The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)
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Bivalvia | Unionida | Unionoidea | Unionidae | Unioninae | Anodontini | Cristariina genus Cristaria Schumacher, 1817: cladomics
7 Recent valid species (10 total species) Cladomics Summarynumber of species = 7 (10 total)
analyzed in 126 cladograms in 85 publications
maximum number of terminals in an analysis =
7 (mean = 1.44, median = 1) Basic Monophylynumber of cladograms that test basic monophyly
(i.e., more than 1 included species) = 21
maximum number of included terminals = 7, (mean =
3.62 median = 3)
proportion of cladograms that support basic monophyly
= 0.95
Pairwise Cladogram ClusteringThe graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.
Dai et al., 2026 [Publ 4757], fig. 3 ** Top Monophyly Cladogram ** iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 7 | 6 | 9 | 27 | 23 | 6 | 0 | +----- Amuranodonta
+-----------------+
| | +-- Anemina
| +--+
| +-- Buldowskia
--+
| +----- Pletholophus
| +--------------+
| | | +-- Beringiana
| | +--+
+--+ +-- Sinanodonta
|
| +----------------- Cristaria_wangyucheni **
| |
+--+ +-------------- Cristaria_bellua **
| |
+--+ +----------- Cristaria_clessini **
| |
+--+ +-- Cristaria_plicata **
| +-----+
| | +-- Cristaria_truncata **
+--+
| +----- Cristaria_radiata__2 **
| |
+--+----- Cristaria_radiata__3 **
|
| +-- Cristaria_liboi **
+--+
+-- Cristaria_radiata__1 **
Chen et al., 2026 [Publ 4805], fig. 4 ** Top Sister-Group Cladogram ** iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 4 | 6 | 28 | 27 | 8 | 5 | +----------- Cristaria_clessini **
|
+-----+ +-------- Cristaria_bellua **
| | |
| +--+ +-- Cristaria_plicata **
| | +--+
| | | +-- Cristaria_truncata **
| +--+
--+ | +-- Cristaria_radiata **
| +--+
| +-- Cristaria_liboi **
|
| +-------------- Beringiana
| |
| | +----- Mimo
+--+ +-----+
| | | +-- Acudonta
| | +--+
+--+ +-- Sinanodonta
|
| +-------- Pletholophus
+--+
| +----- Anemina
+--+
| +-- Amuranodonta
+--+
+-- Buldowskia
Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 4 | 6 | 626 | 27 | 8 | 7 | +-- Cristaria_liboi **
+-----+
| +-- Cristaria_radiata **
|
--+ +-- Cristaria_bellua **
| +--+
| | +-- Cristaria_clessini **
+--+
| +-- Cristaria_plicata **
+--+
+-- Cristaria_truncata **
Wu et al., 2025 [Publ 4442], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 4 | 6 | 29 | 27 | 7 | 5 | +-- Cristaria_clessini **
+-----+
| +-- Cristaria_bellua **
|
+--------+ +-- Cristaria_plicata **
| | +--+
| | | +-- Cristaria_truncata **
| +--+
| | +-- Cristaria_radiata **
| +--+
--+ +-- Cristaria_liboi **
|
| +----- Beringiana
| +--------+
| | | +-- Acudonta
| | +--+
| | +-- Sinanodonta
+--+
| +----- Pletholophus_guangzhouensis
| +-----+
| | | +-- Pletholophus_tenuis
| | +--+
+--+ +-- Pletholophus_reinianus
|
| +-------- Pletholophus_honglinhensis
+--+
| +----- Anemina
+--+
| +-- Amuranodonta
+--+
+-- Buldowskia
Konopleva et al., 2023 [Publ 4269], fig. S1_4 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 5 | 51 | 8 | 5 | 8 | +----- Anemina
+--------+
| | +-- Amuranodonta
| +--+
| +-- Buldowskia
--+
| +-- Beringiana
| +--------+
| | +-- Sinanodonta
+--+
| +-------- Cristaria_radiata **
| |
+--+ +-- Cristaria_truncata **
| +--+
| | +-- Cristaria_plicata **
+--+
| +-- Cristaria_bellua **
+--+
+-- Cristaria_clessini **
Dai et al., 2024 [Publ 4385], fig. 3-ML iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 27 | 23 | 6 | 5 | +----- Amuranodonta
+--------+
| | +-- Anemina
| +--+
| +-- Buldowskia
--+
| +----- Pletholophus
| +-----+
| | | +-- Beringiana
| | +--+
+--+ +-- Sinanodonta
|
| +-------- Cristaria_bellua **
+--+
| +----- Cristaria_clessini **
+--+
| +-- Cristaria_plicata **
+--+
+-- Cristaria_truncata **
Xiang et al., 2025 [Publ 4433], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 24 | 23 | 7 | 5 | +----------------- Pletholophus
|
| +----- Beringiana
--+ +--------+
| | | +-- Mimo
| | +--+
| | +-- Sinanodonta
+--+
| +----- Anemina
| +-----+
| | | +-- Amuranodonta
| | +--+
+--+ +-- Buldowskia
|
| +-------- Cristaria_bellua **
+--+
| +----- Cristaria_clessini **
+--+
| +-- Cristaria_plicata **
+--+
+-- Cristaria_truncata **
Dai et al., 2024 [Publ 4385], fig. 3-BI iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 27 | 23 | 6 | 5 | +----- Amuranodonta
+--------+
| | +-- Anemina
| +--+
| +-- Buldowskia
--+
| +----- Pletholophus
| +-----+
| | | +-- Beringiana
| | +--+
+--+ +-- Sinanodonta
|
| +-------- Cristaria_bellua **
+--+
| +----- Cristaria_clessini **
+--+
| +-- Cristaria_plicata **
+--+
+-- Cristaria_truncata **
Bogan et al., 2023 [Publ 4295], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 27 | 23 | 6 | 5 | +-------- Cristaria_bellua **
+--+
| | +----- Cristaria_clessini **
| +--+
| | +-- Cristaria_plicata **
| +--+
--+ +-- Cristaria_truncata **
|
| +----- Amuranodonta
| +--+
| | | +-- Anemina
| | +--+
+--+ +-- Buldowskia
|
| +----- Pletholophus
+--+
| +-- Beringiana
+--+
+-- Sinanodonta
Bogan et al., 2023 [Publ 4295], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 3 | 2 | 2 | 7 | +-- Sinanodonta
+--------+
| +-- Pletholophus
--+
| +-------- Cristaria_radiata **
+--+
| +----- Cristaria_clessini **
+--+
| +-- Cristaria_plicata **
+--+
+-- Cristaria_truncata **
Wu et al., 2026 [Publ 4811], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 93 | 12 | 7 | 6 | +----- Cristaria_liboi **
+-----+
| | +-- Cristaria_plicata **
| +--+
| +-- Cristaria_truncata **
|
--+ +----- Amuranodonta
| +--+
| | | +-- Anemina
| | +--+
| | +-- Buldowskia
+--+
| +-- Beringiana
| +--+
| | +-- Pletholophus
+--+
| +-- Acudonta
+--+
+-- Sinanodonta
Lopes-Lima et al., 2020 [Publ 4080], fig. 8 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 4 | 2 | 1 | 6 | +-------- Sinanodonta
--+
| +----- Cristaria_clessini **
+--+
| +-- Cristaria_plicata **
+--+
+-- Cristaria_radiata **
Wang et al., 2025 [Publ 4488], fig. 3B iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 62 | 12 | 7 | 6 | +----- Cristaria_liboi **
+-----+
| | +-- Cristaria_plicata **
| +--+
| +-- Cristaria_truncata **
|
--+ +----- Amuranodonta
| +--+
| | | +-- Anemina
| | +--+
| | +-- Buldowskia
+--+
| +-- Beringiana
| +--+
| | +-- Pletholophus
+--+
| +-- Acudonta
+--+
+-- Sinanodonta
Wu et al., 2025 [Publ 4442], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 14 | 12 | 7 | 6 | +----- Cristaria_liboi **
+-----+
| | +-- Cristaria_plicata **
| +--+
| +-- Cristaria_truncata **
|
--+ +----- Amuranodonta
| +--+
| | | +-- Anemina
| | +--+
| | +-- Buldowskia
+--+
| +-- Beringiana
| +--+
| | +-- Pletholophus
+--+
| +-- Acudonta
+--+
+-- Sinanodonta
Bolotov et al., 2017 [Publ 3761], fig. S2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 136 | 4 | 2 | -- | +-- Cristaria_plicata **
+--+
| +-- Cristaria_radiata **
--+
| +-- Pletholophus
+--+
+-- Sinanodonta
Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 120 | 22 | 6 | -- | +-- Cristaria_plicata **
+-----+
| +-- Cristaria_clessini **
|
| +----- Anemina
+--+
--+ | +-- Amuranodonta
| +--+
| +-- Buldowskia
|
| +----- Pletholophus
+--+
| +-- Beringiana
+--+
+-- Sinanodonta
Bolotov et al., 2017 [Publ 3761], fig. S1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 136 | 4 | 2 | -- | +-- Cristaria_plicata **
+--+
| +-- Cristaria_radiata **
--+
| +-- Pletholophus
+--+
+-- Sinanodonta
Klishko et al., 2016 [Publ 3733], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 2 | 2 | 2 | -- | +-- Cristaria_plicata **
+--+
| +-- Cristaria_radiata **
--+
| +-- Sinanodonta
+--+
+-- Beringiana
Bolotov et al., 2017 [Publ 3764], fig. S1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 17 | 0 | 2 | 122 | 4 | 2 | 22 | +-- Alasmidonta
|
+-- Anodonta_anatina
|
+-- Anodonta_cygnea
|
+-- Brachyanodon
|
+-- Lasmigona
|
+-- Pseudanodonta
|
+-- Pyganodon
|
+-- Simpsonaias
|
+-- Strophitus
--+
+-- Sinanodonta_lauta
|
+-- Cristaria_plicata **
|
+-- Cristaria_radiata **
|
+-- Pletholophus
|
+-- Sinanodonta_lucida
|
+-- Sinanodonta_jourdyi
|
+-- Lanceolaria_gladiola
|
+-- Lanceolaria_grayii__1
|
+-- Lanceolaria_grayii__2
Froufe et al., 2017 [Publ 3802], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 10 | 8 | 3 | -- | +-------- Anemina
|
--+ +-- Cristaria_plicata **
| +--+
| | +-- Cristaria_radiata **
+--+
| +-- Beringiana
+--+
+-- Sinanodonta
Klishko et al., 2014 [Publ 3666], fig. 8 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 2 | 2 | 2 | -- | +-- Cristaria_plicata **
+--+
| +-- Cristaria_radiata **
--+
| +-- Beringiana
+--+
+-- Sinanodonta
Chen et al., 2025 [Publ 4487], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 40 | 5 | 5 | -- | Wu et al., 2019 [Publ 3864], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 45 | 3 | 2 | -- | +----- Cristaria_plicata **
--+
| +-- Anemina
+--+
+-- Sinanodonta
Bolotov et al., 2018 [Publ 3826], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 81 | 3 | 2 | -- | +----- Cristaria_plicata **
--+
| +-- Pletholophus
+--+
+-- Sinanodonta
Lopes-Lima et al., 2020 [Publ 4080], fig. 9 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 5 | 3 | 2 | -- | +----- Cristaria_plicata **
--+
| +-- Sinanodonta
+--+
+-- Pletholophus
Huang et al., 2018 [Publ 3828], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 41 | 3 | 2 | -- | +----- Cristaria_plicata **
--+
| +-- Anemina
+--+
+-- Sinanodonta
Huang et al., 2019 [Publ 3843], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 44 | 7 | 2 | -- | Wu et al., 2022 [Publ 4238], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 73 | 3 | 2 | -- | +----- Cristaria_plicata **
--+
| +-- Anemina
+--+
+-- Sinanodonta
Huang et al., 2019 [Publ 3843], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 199 | 8 | 3 | -- | +-------- Anemina
--+
| +----- Cristaria_plicata **
+--+
| +-- Pletholophus
+--+
+-- Sinanodonta
Dai et al., 2023 [Publ 4285], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 52 | 2 | 2 | -- | +----- Cristaria_plicata **
--+
| +-- Anemina
+--+
+-- Sinanodonta
Wu et al., 2016 [Publ 3711], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 20 | 2 | 2 | -- | Smith et al., 2018 [Publ 3790], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 17 | 0 | 0 | -- | Huang et al., 2019 [Publ 3843], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 55 | 3 | 2 | -- | +----- Cristaria_plicata **
--+
| +-- Anemina
+--+
+-- Sinanodonta
Pfeiffer et al., 2019 [Publ 3937], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 56 | 1 | 1 | -- | +-- Cristaria_plicata **
--+
+-- Sinanodonta
Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 79 | 4 | 3 | -- | +-------- Cristaria_plicata **
--+
| +----- Anemina
+--+
| +-- Pletholophus
+--+
+-- Sinanodonta
Dai et al., 2025 [Publ 4441], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 46 | 2 | 2 | -- | +----- Sinanodonta
--+
| +-- Anemina
+--+
+-- Cristaria_plicata **
Wu et al., 2020 [Publ 4147], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 29 | 2 | 2 | -- | +----- Cristaria_plicata **
--+
| +-- Sinanodonta
+--+
+-- Anemina
Huang et al., 2019 [Publ 3843], fig. S3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 197 | 8 | 3 | -- | +-------- Anemina
--+
| +----- Cristaria_plicata **
+--+
| +-- Pletholophus
+--+
+-- Sinanodonta
Lopes-Lima et al., 2017 [Publ 3792], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 31 | 3 | 2 | -- | +----- Cristaria_plicata **
--+
| +-- Anemina
+--+
+-- Sinanodonta
Hou et al., 2024 [Publ 4400], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 75 | 5 | 3 | -- | +-------- Cristaria_plicata **
--+
| +----- Anemina
+--+
| +-- Beringiana
+--+
+-- Sinanodonta
Sano et al., 2020 [Publ 4138], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 67 | 10 | 5 | -- | Wu et al., 2024 [Publ 4388], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 52 | 3 | 3 | -- | +-------- Cristaria_plicata **
--+
| +----- Anemina
+--+
| +-- Beringiana
+--+
+-- Sinanodonta
Huang et al., 2019 [Publ 3843], fig. S4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 199 | 8 | 3 | -- | Wu et al., 2023 [Publ 4339], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 18 | 3 | 2 | -- | +----- Anemina
--+
| +-- Cristaria_plicata **
+--+
+-- Sinanodonta
Zhang et al., 2004 [Publ 2735], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 13 | 1 | 1 | -- | +-- Sinanodonta
--+
+-- Cristaria_plicata **
Pavan-Kumar et al., 2022 [Publ 4263], fig. S5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 38 | 3 | 2 | -- | +----- Cristaria_plicata **
--+
| +-- Anemina
+--+
+-- Sinanodonta
Pfarrer et al., 2022 [Publ 4305], fig. 2-5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 11 | 1 | 1 | -- | +-- Cristaria_plicata **
--+
+-- Sinanodonta
Sano et al., 2022 [Publ 4311], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 17 | 17 | 4 | -- | +-- Beringiana
+-----+
| +-- Sinanodonta
--+
| +----- Pletholophus
+--+
| +-- Cristaria_plicata **
+--+
+-- Buldowskia
Wu et al., 2021 [Publ 4366], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 68 | 3 | 2 | -- | +----- Cristaria_plicata **
--+
| +-- Anemina
+--+
+-- Sinanodonta
Soroka & Burzynski, 2017 [Publ 3837], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 22 | 3 | 2 | -- | +----- Cristaria_plicata **
--+
| +-- Anemina
+--+
+-- Sinanodonta
Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 140 | 3 | 2 | -- | +----- Cristaria_plicata **
--+
| +-- Anemina
+--+
+-- Sinanodonta
Wu et al., 2024 [Publ 4350], fig. 8b_S2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 91 | 5 | 3 | -- | +-------- Cristaria_plicata **
--+
| +----- Anemina
+--+
| +-- Beringiana
+--+
+-- Sinanodonta
Huang et al., 2002 [Publ 2657], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 33 | 1 | 1 | -- | +-- Sinanodonta
--+
+-- Cristaria_plicata **
Wu et al., 2018 [Publ 4143], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 21 | 2 | 2 | -- | Chen et al., 2012 [Publ 3624], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 6 | 0 | 0 | -- | Pfeiffer & Graf, 2013 [Publ 3531], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 30 | 1 | 1 | -- | +-- Cristaria_plicata **
--+
+-- Sinanodonta
Xiang et al., 2025 [Publ 4433], fig. 7 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 12 | 6 | 4 | -- | +----------- Cristaria_plicata **
--+
| +-------- Anemina
+--+
| +----- Beringiana
+--+
| +-- Mimo
+--+
+-- Sinanodonta
Huang et al., 2002 [Publ 2657], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 33 | 1 | 1 | -- | +-- Sinanodonta
--+
+-- Cristaria_plicata **
Wu et al., 2023 [Publ 4339], fig. 4_S5-nt iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 95 | 5 | 3 | -- | +-------- Cristaria_plicata **
--+
| +----- Anemina
+--+
| +-- Beringiana
+--+
+-- Sinanodonta
Sano et al., 2017 [Publ 3799], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 20 | 7 | 4 | -- | +-- Cristaria_plicata **
+-----+
| +-- Buldowskia
--+
| +----- Pletholophus
+--+
| +-- Sinanodonta
+--+
+-- Beringiana
Fonseca et al., 2016 [Publ 3775], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 30 | 3 | 2 | -- | +----- Cristaria_plicata **
--+
| +-- Anemina
+--+
+-- Sinanodonta
Mabuchi & Nishida, 2020 [Publ 4120], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 11 | 4 | 2 | -- | +-------------- Cristaria_plicata **
|
| +----------- Sinanodonta_tumens
| |
| | +-- Sinanodonta_lauta__5
| | |
--+ | +-- Sinanodonta_lauta__6
| | |
| | +-- Beringiana_japonica__15
| | |
| | +-- Beringiana_japonica__16
| | +-----+
| | | +-- Beringiana_japonica__17
+--+ | |
| | +-- Beringiana_japonica__18
| | |
| | +-- Beringiana_japonica__19
| | |
| | +-- Beringiana_japonica__20
| |
| | +----- Beringiana_japonica__1
| | |
| | +----- Beringiana_japonica__2
| | |
+--+ +----- Beringiana_japonica__3
| |
| +----- Beringiana_japonica__4
| |
| +----- Beringiana_japonica__5
| |
| +----- Sinanodonta_lauta__1
| |
| +----- Beringiana_japonica__6
| |
| +----- Sinanodonta_lauta__2
| |
| +----- Sinanodonta_calipygos__1
| |
+--+----- Beringiana_japonica__7
|
+----- Sinanodonta_lauta__4
|
+----- Beringiana_japonica__8
|
+----- Beringiana_japonica__9
|
+----- Beringiana_japonica__10
|
+----- Beringiana_japonica__11
|
+----- Beringiana_japonica__12
|
+----- Beringiana_japonica__13
|
+----- Beringiana_japonica__14
|
| +-- Sinanodonta_lauta__3
+--+
+-- Sinanodonta_calipygos__2
Sano et al., 2017 [Publ 3799], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 32 | 8 | 4 | -- | +-- Cristaria_plicata **
+-----+
| +-- Buldowskia
--+
| +----- Beringiana
+--+
| +-- Pletholophus
+--+
+-- Sinanodonta
Wang et al., 2018 [Publ 3865], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 30 | 3 | 2 | -- | +----- Cristaria_plicata **
--+
| +-- Anemina
+--+
+-- Sinanodonta
Bogan et al., 2023 [Publ 4295], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 5 | 4 | 2 | -- | +----- Cristaria_truncata **
--+
| +-- Sinanodonta
+--+
+-- Pletholophus
Dai et al., 2024 [Publ 4358], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 52 | 2 | 2 | -- | +----- Cristaria_plicata **
--+
| +-- Anemina
+--+
+-- Sinanodonta
Chen et al., 2025 [Publ 4663], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 47 | 2 | 2 | -- | +----- Sinanodonta
--+
| +-- Anemina
+--+
+-- Cristaria_plicata **
Bolotov et al., 2020 [Publ 4084], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 35 | 8 | 5 | -- | +----- Amuranodonta
+--+
| | +-- Anemina
| +--+
--+ +-- Buldowskia
|
| +----- Cristaria_plicata **
+--+
| +-- Beringiana
+--+
+-- Sinanodonta
Fukata & Iigo, 2020 [Publ 4106], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 53 | 3 | 2 | -- | +----- Cristaria_plicata **
--+
| +-- Anemina
+--+
+-- Sinanodonta
Bogan et al., 2023 [Publ 4295], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 12 | 11 | 2 | -- | +----- Pletholophus
--+
| +-- Cristaria_truncata **
+--+
+-- Sinanodonta
Xue et al., 2016 [Publ 3659], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 14 | 3 | 2 | -- | Plazzi et al., 2011 [Publ 3676], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 17 | 0 | 0 | -- | Jiang et al., 2010 [Publ 3438], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 3 | 0 | 0 | -- | Wu et al., 2021 [Publ 4366], fig. 3-BI iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 73 | 3 | 2 | -- | +----- Cristaria_plicata **
--+
| +-- Anemina
+--+
+-- Sinanodonta
Dai et al., 2025 [Publ 4440], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 44 | 5 | 5 | -- | Cha et al., 2018 [Publ 3916], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 37 | 3 | 2 | -- | +----- Cristaria_plicata **
--+
| +-- Anemina
+--+
+-- Sinanodonta
Wang et al., 2018 [Publ 3846], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 30 | 3 | 2 | -- | +----- Cristaria_plicata **
--+
| +-- Anemina
+--+
+-- Sinanodonta
Dai et al., 2025 [Publ 4440], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 44 | 5 | 5 | -- | Wu et al., 2024 [Publ 4379], fig. 4_6 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 22 | 2 | 1 | -- | +-- Cristaria_plicata **
--+
+-- Sinanodonta
Aunins et al., 2018 [Publ 3863], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 39 | 3 | 2 | -- | +----- Cristaria_plicata **
--+
| +-- Anemina
+--+
+-- Sinanodonta
Wu et al., 2024 [Publ 4350], fig. 8a_S1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 91 | 5 | 3 | -- | +-------- Cristaria_plicata **
--+
| +----- Anemina
+--+
| +-- Beringiana
+--+
+-- Sinanodonta
Dai et al., 2025 [Publ 4440], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 87 | 3 | 2 | -- | +----- Cristaria_plicata **
--+
| +-- Anemina
+--+
+-- Sinanodonta
Wen et al., 2017 [Publ 3795], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 15 | 1 | 1 | -- | +-- Sinanodonta
--+
+-- Cristaria_plicata **
Riccardi et al., 2020 [Publ 4089], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 18 | 3 | 2 | -- | +----- Cristaria_plicata **
--+
| +-- Anemina
+--+
+-- Sinanodonta
Zhang et al., 2013 [Publ 4214], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 3 | 1 | 1 | -- | Wu et al., 2022 [Publ 4238], fig. 2d iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 38 | 3 | 2 | -- | +----- Cristaria_plicata **
--+
| +-- Anemina
+--+
+-- Sinanodonta
Wei et al., 2016 [Publ 3665], fig. 0 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 5 | 0 | 0 | -- | Chen et al., 2023 [Publ 4302], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 40 | 5 | 5 | -- | Sheng et al., 2014 [Publ 3704], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 7 | 0 | 0 | -- | Gomes-dos-Santos et al., 2019 [Publ 4042], fig. 8 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 5 | 2 | 2 | -- | +----- Cristaria_radiata **
--+
| +-- Anemina
+--+
+-- Sinanodonta
Wu et al., 2018 [Publ 3836], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 52 | 5 | 2 | -- | +-- Anemina
|
--+-- Cristaria_plicata **
|
+-- Sinanodonta
Zhou et al., 2016 [Publ 3710], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 19 | 1 | 1 | -- | Chen et al., 2025 [Publ 4487], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 40 | 5 | 5 | -- | +----------- Cristaria_plicata **
|
--+ +-- Anemina
| +-----+
| | +-- Amuranodonta
+--+
| +----- Pletholophus
+--+
| +-- Sinanodonta
+--+
+-- Beringiana
Smith, 2021 [Publ 4331], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 19 | 0 | 0 | -- | Chen et al., 2023 [Publ 4302], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 40 | 5 | 5 | -- | +----------- Cristaria_plicata **
|
--+ +-- Anemina
| +-----+
| | +-- Amuranodonta
+--+
| +----- Pletholophus
+--+
| +-- Sinanodonta
+--+
+-- Beringiana
Chen et al., 2020 [Publ 4170], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 33 | 2 | 2 | -- | Breton et al., 2011 [Publ 3376], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 37 | 0 | 0 | -- | Wu et al., 2021 [Publ 4366], fig. 2_I iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 90 | 3 | 2 | -- | +----- Anemina
--+
| +-- Cristaria_plicata **
+--+
+-- Sinanodonta
Chen et al., 2025 [Publ 4448], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 85 | 4 | 2 | -- | +----- Cristaria_plicata **
--+
| +-- Anemina
+--+
+-- Sinanodonta
Pavan-Kumar et al., 2022 [Publ 4263], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 36 | 2 | 2 | -- | +----- Cristaria_plicata **
--+
| +-- Sinanodonta
+--+
+-- Anemina
Huang et al., 2013 [Publ 3629], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 15 | 1 | 1 | -- | +-- Sinanodonta
--+
+-- Cristaria_plicata **
Froufe et al., 2017 [Publ 3802], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 3 | 0 | 0 | -- | Wu et al., 2023 [Publ 4339], fig. 4_S5-aa iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 95 | 5 | 3 | -- | +-- Anemina
+--+
| +-- Cristaria_plicata **
--+
| +-- Beringiana
+--+
+-- Sinanodonta
Froufe et al., 2019 [Publ 3936], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 43 | 3 | 2 | -- | +----- Cristaria_plicata **
--+
| +-- Anemina
+--+
+-- Sinanodonta
Xia et al., 2016 [Publ 3712], fig. 7B iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 2 | 1 | 1 | -- | Peng et al., 2012 [Publ 3689], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 2 | 0 | 0 | -- | Pfeiffer et al., 2019 [Publ 3937], fig. 4-5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 56 | 1 | 1 | -- | +-- Cristaria_plicata **
--+
+-- Sinanodonta
Guerra et al., 2017 [Publ 3772], fig. 2a iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 24 | 1 | 1 | -- | Sano et al., 2020 [Publ 4138], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 22 | 0 | 3 | 33 | 9 | 5 | 36 | +----- Lanceolaria_grayii__1
|
+----- Lanceolaria_grayii__2
|
+----- Cristaria_plicata__1 **
|
+----- Cristaria_plicata__2 **
|
+----- Sinanodonta_lauta
|
+----- Beringiana_japonica
|
+----- Beringiana_fukuharai__1
|
+----- Beringiana_beringiana
|
+----- Beringiana_fukuharai__2
|
+----- Beringiana_fukuharai__3
|
+----- Buldowskia_iwakawai__1
|
+----- Buldowskia_iwakawai__2
|
+----- Sinanodonta_tumens__1
--+
+----- Sinanodonta_tumens__2
|
+----- Pletholophus
|
+----- Anemina
|
+----- Lanceolaria_lanceolata
|
+----- Cristaria_plicata__3 **
|
+----- Lanceolaria_gladiola
|
+----- Sinanodonta_angula
|
+----- Lepidodesma
|
| +-- Scriptolamprotula
+--+
| +-- Acuticosta
|
| +-- Cuneopsis
+--+
| +-- Nodularia
|
| +-- Schistodesmus
+--+
+-- Inversiunio
Guerra et al., 2017 [Publ 3772], fig. 2b iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 24 | 1 | 1 | -- | +-- Cristaria_plicata **
--+
+-- Sinanodonta
Wu et al., 2020 [Publ 4121], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 48 | 3 | 2 | -- | +----- Cristaria_plicata **
--+
| +-- Anemina
+--+
+-- Sinanodonta
Froufe et al., 2019 [Publ 3936], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 43 | 3 | 2 | -- | +----- Cristaria_plicata **
--+
| +-- Anemina
+--+
+-- Sinanodonta
Sano et al., 2020 [Publ 4138], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 67 | 10 | 5 | -- | Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 75 | 4 | 2 | -- | +----- Cristaria_plicata **
--+
| +-- Pletholophus
+--+
+-- Sinanodonta
Wu et al., 2021 [Publ 4366], fig. 3-ML iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 73 | 3 | 2 | -- | +----- Cristaria_plicata **
--+
| +-- Anemina
+--+
+-- Sinanodonta
Wu et al., 2024 [Publ 4379], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 53 | 5 | 3 | -- | +-------- Cristaria_plicata **
--+
| +----- Anemina
+--+
| +-- Beringiana
+--+
+-- Sinanodonta
Hou et al., 2025 [Publ 4445], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 52 | 5 | 3 | -- | +-------- Cristaria_plicata **
--+
| +----- Anemina
+--+
| +-- Beringiana
+--+
+-- Sinanodonta
Wu et al., 2021 [Publ 4366], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 90 | 3 | 2 | -- | +----- Anemina
--+
| +-- Cristaria_plicata **
+--+
+-- Sinanodonta
Zieritz et al., 2021 [Publ 4231], fig. 3-F_5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 47 | 3 | 2 | -- | +----- Cristaria_plicata **
--+
| +-- Anemina
+--+
+-- Sinanodonta
Dai et al., 2024 [Publ 4389], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 45 | 2 | 2 | -- | +----- Sinanodonta
--+
| +-- Anemina
+--+
+-- Cristaria_plicata **
Sano et al., 2020 [Publ 4138], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 67 | 10 | 5 | -- | Zhou et al., 2007 [Publ 2658], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 20 | 1 | 1 | -- | +-- Sinanodonta
--+
+-- Cristaria_plicata **
Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 75 | 4 | 2 | -- | +----- Cristaria_plicata **
--+
| +-- Pletholophus
+--+
+-- Sinanodonta
Sano et al., 2022 [Publ 4311], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 11 | 11 | 4 | -- | +-- Beringiana
+-----+
| +-- Sinanodonta
--+
| +----- Pletholophus
+--+
| +-- Cristaria_plicata **
+--+
+-- Buldowskia
Huang et al., 2019 [Publ 3843], fig. S1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 46 | 7 | 2 | -- | Zhou et al., 2007 [Publ 2658], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 17 | 1 | 1 | -- |
|
