The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)
FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!
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Bivalvia | Unionida | Unionoidea | Margaritiferidae | Margaritiferinae genus Cumberlandia Ortmann, 1912: cladomics 1 Recent valid species (1 total species) Cladomics Summarynumber of species = 1 (1 total) analyzed in 115 cladograms in 60 publications maximum number of terminals in an analysis =
1 (mean = 1, median = 1)
Lyubas et al., 2019 [Publ 3963], fig. 10 ** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram ** | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 15 | 12 | 2 | -- |
+----- Margaritifera
--+
| +-- Cumberlandia_monodonta **
+--+
+-- Pseudunio
Araujo et al., 2016 [Publ 3746], fig. 2a | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 10 | 9 | 2 | -- |
-- Cumberlandia_monodonta **
Wu et al., 2021 [Publ 4366], fig. 3-BI | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 72 | 4 | 2 | -- |
+----- Cumberlandia_monodonta **
--+
| +-- Pseudunio
+--+
+-- Margaritifera
Breton et al., 2011 [Publ 3376], fig. 3 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 37 | 3 | 1 | -- |
+-- Cumberlandia_monodonta **
--+
+-- Margaritifera
Guerra et al., 2017 [Publ 3772], fig. 2a | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 24 | 1 | 1 | -- |
+-- Margaritifera
--+
+-- Cumberlandia_monodonta **
Hoeh et al., 2002 [Publ 2653], fig. 2 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 30 | 1 | 1 | -- |
+-- Margaritifera
--+
+-- Cumberlandia_monodonta **
Rosenberg et al., 1997 [Publ 2720], fig. 2A | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 21 | 2 | 1 | -- |
-- Cumberlandia_monodonta **
Lydeard et al., 2000 [Publ 2654], fig. 2d | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 13 | 0 | 0 | -- |
-- Cumberlandia_monodonta **
Lopes-Lima et al., 2019 [Publ 3901], fig. 2 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 27 | 1 | 1 | -- |
+-- Margaritifera
--+
+-- Cumberlandia_monodonta **
Chen et al., 2025 [Publ 4455], fig. 1 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 25 | 10 | 2 | -- |
+-- Cumberlandia_monodonta **
--+
+-- Margaritifera
Takeuchi et al., 2015 [Publ 3661], fig. 2 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 9 | 7 | 2 | -- |
+-------- Pseudunio_marocanus
--+
| +----- Pseudunio_auricularius
+--+
| +-- Cumberlandia_monodonta **
+--+
+-- Margaritifera
Huff et al., 2004 [Publ 2697], fig. 5-6 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 11 | 6 | 2 | -- |
+----- Margaritifera
--+
| +-- Pseudunio
+--+
+-- Cumberlandia_monodonta **
Walker et al., 2006 [Publ 2666], fig. 2 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 14 | 0 | 0 | -- |
-- Cumberlandia_monodonta **
Zanatta & Murphy, 2006 [Publ 2672], fig. 1 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 55 | 0 | 0 | -- |
-- Cumberlandia_monodonta **
Graf & Cummings, 2006 [Publ 1968], fig. 2 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 46 | 1 | 1 | -- |
+-- Margaritifera
--+
+-- Cumberlandia_monodonta **
Lopes-Lima et al., 2018 [Publ 3829], fig. 3 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 27 | 10 | 2 | -- |
+----- Margaritifera
--+
| +-- Cumberlandia_monodonta **
+--+
+-- Pseudunio
Walker et al., 2006 [Publ 2666], fig. 6-M | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 13 | 2 | 1 | -- |
+-- Cumberlandia_monodonta **
--+
+-- Margaritifera
Araujo et al., 2016 [Publ 3746], fig. 1_2b | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 12 | 9 | 2 | -- |
+----- Margaritifera
--+
| +-- Cumberlandia_monodonta **
+--+
+-- Pseudunio
Hoeh et al., 2009 [Publ 2716], fig. 5 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 45 | 1 | 1 | -- |
+-- Margaritifera
--+
+-- Cumberlandia_monodonta **
Zanatta & Murphy, 2006 [Publ 2672], fig. 2 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 56 | 0 | 0 | -- |
-- Cumberlandia_monodonta **
Lydeard et al., 2000 [Publ 2654], fig. 2b | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 13 | 0 | 0 | -- |
-- Cumberlandia_monodonta **
Guerra et al., 2019 [Publ 4168], fig. 1 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 40 | 5 | 2 | -- |
+----- Cumberlandia_monodonta **
--+
| +-- Pseudunio
+--+
+-- Margaritifera
Inoue et al., 2018 [Publ 3858], fig. 1_3a-c | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 104 | 0 | 0 | -- |
-- Cumberlandia_monodonta **
Walker et al., 2006 [Publ 2666], fig. 3 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 14 | 0 | 0 | -- |
-- Cumberlandia_monodonta **
Chase et al., 2018 [Publ 3861], fig. 1-M | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 10 | 0 | 0 | -- |
-- Cumberlandia_monodonta **
Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 75 | 8 | 2 | -- |
-- Cumberlandia_monodonta **
Gomes-dos-Santos et al., 2019 [Publ 3943], fig. 1 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 9 | 4 | 2 | -- |
+----- Cumberlandia_monodonta **
--+
| +-- Pseudunio
+--+
+-- Margaritifera
Lopes-Lima et al., 2019 [Publ 3901], fig. 4 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 25 | 1 | 1 | -- |
+-- Margaritifera
--+
+-- Cumberlandia_monodonta **
Walker et al., 2006 [Publ 2666], fig. 4 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 14 | 0 | 0 | -- |
-- Cumberlandia_monodonta **
Rosenberg et al., 1997 [Publ 2720], fig. 2B | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 21 | 2 | 1 | -- |
-- Cumberlandia_monodonta **
Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 75 | 8 | 2 | -- |
-- Cumberlandia_monodonta **
Vikhrev et al., 2018 [Publ 3812], fig. 3 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 12 | 9 | 2 | -- |
+----- Margaritifera
--+
| +-- Cumberlandia_monodonta **
+--+
+-- Pseudunio
Graf, 2002 [Publ 2651], fig. 2 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 17 | 0 | 0 | -- |
-- Cumberlandia_monodonta **
Roe & Hoeh, 2003 [Publ 2655], fig. 4f3 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 30 | 1 | 1 | -- |
+-- Cumberlandia_monodonta **
--+
+-- Margaritifera
Walker et al., 2006 [Publ 2666], fig. 6 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 35 | 5 | 2 | -- |
+----- Cumberlandia_monodonta **
--+
| +-- Pseudunio
+--+
+-- Margaritifera
Chen et al., 2025 [Publ 4448], fig. 3 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 84 | 4 | 2 | -- |
+----- Cumberlandia_monodonta **
--+
| +-- Pseudunio
+--+
+-- Margaritifera
Wu et al., 2020 [Publ 4121], fig. 2 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 33 | 9 | 2 | -- |
-- Cumberlandia_monodonta **
Whelan et al., 2011 [Publ 3380], fig. 1-ML | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 35 | 1 | 1 | -- |
+-- Margaritifera
--+
+-- Cumberlandia_monodonta **
Wu et al., 2021 [Publ 4366], fig. 3-ML | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 72 | 4 | 2 | -- |
+----- Cumberlandia_monodonta **
--+
| +-- Pseudunio
+--+
+-- Margaritifera
Wu et al., 2019 [Publ 3864], fig. 3 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 45 | 2 | 1 | -- |
+-- Cumberlandia_monodonta **
--+
+-- Margaritifera
Teiga-Teixeira et al., 2020 [Publ 4173], fig. 1 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 18 | 1 | 1 | -- |
+-- Margaritifera
--+
+-- Cumberlandia_monodonta **
Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 80 | 1 | 1 | -- |
+-- Cumberlandia_monodonta **
--+
+-- Margaritifera
Wu et al., 2022 [Publ 4238], fig. 3 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 72 | 4 | 2 | -- |
+----- Cumberlandia_monodonta **
--+
| +-- Pseudunio
+--+
+-- Margaritifera
Sano et al., 2020 [Publ 4138], fig. 2 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 68 | 9 | 2 | -- |
-- Cumberlandia_monodonta **
Inoue et al., 2014 [Publ 3635], fig. 3A | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 19 | 0 | 0 | -- |
-- Cumberlandia_monodonta **
Guerra et al., 2017 [Publ 3772], fig. 2b-M | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 7 | 0 | 0 | -- |
-- Cumberlandia_monodonta **
Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 139 | 10 | 2 | -- |
+----- Cumberlandia_monodonta **
--+
| +-- Pseudunio
+--+
+-- Margaritifera
Zhou et al., 2007 [Publ 2658], fig. 1 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 20 | 1 | 1 | -- |
+-- Margaritifera
--+
+-- Cumberlandia_monodonta **
Wu et al., 2020 [Publ 4121], fig. 3 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 47 | 2 | 1 | -- |
+-- Cumberlandia_monodonta **
--+
+-- Margaritifera
Rosenberg et al., 1994 [Publ 2719], fig. 3 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 18 | 2 | 1 | -- |
-- Cumberlandia_monodonta **
Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 29 | 1 | 1 | -- |
+-- Margaritifera
--+
+-- Cumberlandia_monodonta **
Huang et al., 2019 [Publ 3843], fig. 3 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 55 | 4 | 2 | -- |
+----- Cumberlandia_monodonta **
--+
| +-- Pseudunio
+--+
+-- Margaritifera
Bolotov et al., 2016 [Publ 3743], fig. 2 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 29 | 8 | 2 | -- |
-- Cumberlandia_monodonta **
Campbell & Lydeard, 2026 [Publ 4810], fig. 4 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 46 | 2 | 2 | -- |
+----- Pseudunio
--+
| +-- Cumberlandia_monodonta **
+--+
+-- Margaritifera
Campbell et al., 2005 [Publ 2660], fig. 1 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 101 | 1 | 1 | -- |
+-- Margaritifera
--+
+-- Cumberlandia_monodonta **
Wu et al., 2023 [Publ 4339], fig. 4_S5-aa | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 94 | 4 | 2 | -- |
+----- Cumberlandia_monodonta **
--+
| +-- Pseudunio
+--+
+-- Margaritifera
Guerra et al., 2019 [Publ 4168], fig. 2 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 40 | 5 | 2 | -- |
+----- Cumberlandia_monodonta **
--+
| +-- Pseudunio
+--+
+-- Margaritifera
Sano et al., 2020 [Publ 4138], fig. 3 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 68 | 9 | 2 | -- |
-- Cumberlandia_monodonta **
Bolotov et al., 2017 [Publ 3761], fig. S1 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 138 | 8 | 2 | -- |
-- Cumberlandia_monodonta **
Campbell et al., 2005 [Publ 2660], fig. 2 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 101 | 1 | 1 | -- |
+-- Cumberlandia_monodonta **
--+
+-- Margaritifera
Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 29 | 1 | 1 | -- |
+-- Margaritifera
--+
+-- Cumberlandia_monodonta **
Sano et al., 2020 [Publ 4138], fig. 4 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 68 | 9 | 2 | -- |
-- Cumberlandia_monodonta **
Chase et al., 2018 [Publ 3847], fig. 4 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 38 | 3 | 1 | -- |
+-- Cumberlandia_monodonta **
--+
+-- Margaritifera
Guerra et al., 2019 [Publ 4168], fig. 2-M | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 21 | 2 | 2 | -- |
+----- Pseudunio
--+
| +-- Margaritifera
+--+
+-- Cumberlandia_monodonta **
Wu et al., 2023 [Publ 4339], fig. 4_S5-nt | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 94 | 4 | 2 | -- |
+----- Cumberlandia_monodonta **
--+
| +-- Pseudunio
+--+
+-- Margaritifera
Huang et al., 2019 [Publ 3843], fig. S3 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 198 | 10 | 2 | -- |
+----- Margaritifera
--+
| +-- Cumberlandia_monodonta **
+--+
+-- Pseudunio
Bogan & Hoeh, 2000 [Publ 2650], fig. 1 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 33 | 1 | 1 | -- |
+-- Cumberlandia_monodonta **
--+
+-- Margaritifera
Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 631 | 10 | 2 | -- |
+-- Cumberlandia_monodonta **
--+
+-- Margaritifera
Huang et al., 2019 [Publ 3843], fig. 2 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 200 | 10 | 2 | -- |
+-- Cumberlandia_monodonta **
|
--+-- Pseudunio
|
+-- Margaritifera
Graf & Ó Foighil, 2000 [Publ 2649], fig. 1_2 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 10 | 0 | 0 | -- |
-- Cumberlandia_monodonta **
Wu et al., 2020 [Publ 4121], fig. S1 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 33 | 9 | 2 | -- |
+----- Margaritifera
--+
| +-- Cumberlandia_monodonta **
+--+
+-- Pseudunio
Guerra et al., 2019 [Publ 4168], fig. 1-M | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 21 | 2 | 2 | -- |
+----- Cumberlandia_monodonta **
--+
| +-- Margaritifera
+--+
+-- Pseudunio
Wu et al., 2021 [Publ 4366], fig. 4 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 67 | 4 | 2 | -- |
+----- Cumberlandia_monodonta **
--+
| +-- Pseudunio
+--+
+-- Margaritifera
Bolotov et al., 2016 [Publ 3743], fig. 4 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 29 | 8 | 2 | -- |
-- Cumberlandia_monodonta **
Hoeh et al., 2001 [Publ 2652], fig. 14f4 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 30 | 1 | 1 | -- |
+-- Cumberlandia_monodonta **
--+
+-- Margaritifera
Huang et al., 2018 [Publ 3828], fig. 6-M | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 16 | 0 | 0 | -- |
-- Cumberlandia_monodonta **
Gomes-dos-Santos et al., 2019 [Publ 3943], fig. 1-M | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 5 | 2 | 2 | -- |
+----- Margaritifera
--+
| +-- Pseudunio
+--+
+-- Cumberlandia_monodonta **
Bolotov et al., 2015 [Publ 3646], fig. 2A | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 8 | 7 | 2 | -- |
+----------- Margaritifera_margaritifera
|
| +----- Margaritifera_dahurica__1
| |
+-----+----- Margaritifera_dahurica__2
--+ |
| | +-- Pseudunio
| +--+
| +-- Cumberlandia_monodonta **
|
| +-------- Margaritifera_falcata
+--+
| +----- Margaritifera_laevis
+--+
| +-- Margaritifera_middendorffi
+--+
+-- Margaritifera_marrianae
Wu et al., 2021 [Publ 4366], fig. 6 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 90 | 5 | 2 | -- |
+----- Margaritifera
--+
| +-- Cumberlandia_monodonta **
+--+
+-- Pseudunio
Wu et al., 2021 [Publ 4366], fig. 2_I | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 90 | 5 | 2 | -- |
+----- Margaritifera
--+
| +-- Pseudunio
+--+
+-- Cumberlandia_monodonta **
Lydeard et al., 2000 [Publ 2654], fig. 2c | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 13 | 0 | 0 | -- |
-- Cumberlandia_monodonta **
Guerra et al., 2017 [Publ 3772], fig. 2b | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 24 | 1 | 1 | -- |
+-- Margaritifera
--+
+-- Cumberlandia_monodonta **
Roe & Hoeh, 2003 [Publ 2655], fig. 4f2 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 30 | 1 | 1 | -- |
+-- Cumberlandia_monodonta **
--+
+-- Margaritifera
Machordom et al., 2003 [Publ 2688], fig. 3 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 3 | 2 | 2 | -- |
+----- Cumberlandia_monodonta **
--+
| +-- Margaritifera
+--+
+-- Pseudunio
Marshall et al., 2014 [Publ 3593], fig. 1 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 13 | 0 | 0 | -- |
-- Cumberlandia_monodonta **
Hoeh et al., 2001 [Publ 2652], fig. 14f2 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 30 | 1 | 1 | -- |
+-- Cumberlandia_monodonta **
--+
+-- Margaritifera
Lydeard et al., 2000 [Publ 2654], fig. 2a | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 13 | 0 | 0 | -- |
-- Cumberlandia_monodonta **
Inoue et al., 2013 [Publ 3579], fig. 3 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 8 | 7 | 2 | -- |
+----- Cumberlandia_monodonta **
--+
| +-- Pseudunio
+--+
+-- Margaritifera
Hoeh et al., 2009 [Publ 2716], fig. 3 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 46 | 1 | 1 | -- |
+-- Margaritifera
--+
+-- Cumberlandia_monodonta **
Chase et al., 2018 [Publ 3861], fig. 1 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 10 | 0 | 0 | -- |
-- Cumberlandia_monodonta **
Huang et al., 2018 [Publ 3828], fig. 6 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 41 | 4 | 2 | -- |
+----- Cumberlandia_monodonta **
--+
| +-- Pseudunio
+--+
+-- Margaritifera
Huang et al., 2002 [Publ 2657], fig. 1 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 33 | 1 | 1 | -- |
-- Cumberlandia_monodonta **
Hoeh et al., 2009 [Publ 2716], fig. 4 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 45 | 1 | 1 | -- |
+-- Margaritifera
--+
+-- Cumberlandia_monodonta **
Aunins et al., 2018 [Publ 3863], fig. 1 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 39 | 2 | 1 | -- |
+-- Cumberlandia_monodonta **
--+
+-- Margaritifera
Huff et al., 2004 [Publ 2697], fig. 2 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 11 | 6 | 2 | -- |
+-------- Margaritifera_dahurica
|
+-------- Margaritifera_margaritifera__1
|
+-------- Margaritifera_margaritifera__2
|
--+-------- Margaritifera_margaritifera__3
|
| +-- Pseudunio
+-----+
| +-- Cumberlandia_monodonta **
|
| +----- Margaritifera_falcata
+--+
| +-- Margaritifera_middendorffi
+--+
+-- Margaritifera_marrianae
Lydeard et al., 1996 [Publ 2656], fig. 3 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 26 | 2 | 1 | -- |
+-- Cumberlandia_monodonta **
--+
+-- Margaritifera
Bolotov et al., 2015 [Publ 3646], fig. 2B | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 8 | 7 | 2 | -- |
+----- Cumberlandia_monodonta **
--+
| +-- Pseudunio
+--+
+-- Margaritifera
Huang et al., 2002 [Publ 2657], fig. 2 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 33 | 1 | 1 | -- |
-- Cumberlandia_monodonta **
Whelan et al., 2011 [Publ 3380], fig. 1-MP | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 35 | 1 | 1 | -- |
+-- Margaritifera
--+
+-- Cumberlandia_monodonta **
Lydeard et al., 1996 [Publ 2656], fig. 4 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 27 | 2 | 1 | -- |
+-- Cumberlandia_monodonta **
--+
+-- Margaritifera
Chase et al., 2018 [Publ 3847], fig. 5 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 9 | 0 | 0 | -- |
-- Cumberlandia_monodonta **
Guerra et al., 2017 [Publ 3772], fig. 2a-M | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 7 | 0 | 0 | -- |
-- Cumberlandia_monodonta **
Lopes-Lima et al., 2019 [Publ 3901], fig. 3 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 28 | 1 | 1 | -- |
+-- Cumberlandia_monodonta **
--+
+-- Margaritifera
Dai et al., 2025 [Publ 4440], fig. 3 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 86 | 4 | 2 | -- |
+----- Cumberlandia_monodonta **
--+
| +-- Pseudunio
+--+
+-- Margaritifera
Takeuchi et al., 2015 [Publ 3661], fig. 3 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 9 | 7 | 2 | -- |
+----- Pseudunio_auricularius__1
|
--+----- Pseudunio_auricularius__2
|
| +-- Cumberlandia_monodonta **
+--+
+-- Margaritifera
Burzynski & Soroka, 2018 [Publ 3862], fig. 5a | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 19 | 1 | 1 | -- |
+-- Pseudunio
--+
+-- Cumberlandia_monodonta **
Huang et al., 2018 [Publ 3828], fig. 3_5 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 28 | 9 | 2 | -- |
+----- Margaritifera
--+
| +-- Cumberlandia_monodonta **
+--+
+-- Pseudunio
Huff et al., 2004 [Publ 2697], fig. 3 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 7 | 5 | 2 | -- |
+-------- Margaritifera_margaritifera
|
| +----- Cumberlandia_monodonta **
--+ |
| | +-- Pseudunio
| | |
+--+ +-- Margaritifera_dahurica
| |
| +-- Margaritifera_middendorffi__1
| |
| +-- Margaritifera_middendorffi__2
+--+
+-- Margaritifera_falcata__1
|
+-- Margaritifera_falcata__2
|
+-- Margaritifera_falcata__3
|
+-- Margaritifera_falcata__4
Huang et al., 2019 [Publ 3843], fig. S4 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 200 | 10 | 2 | -- |
+----- Cumberlandia_monodonta **
--+
| +-- Pseudunio
+--+
+-- Margaritifera
Lopes-Lima et al., 2018 [Publ 3829], fig. 1 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 27 | 10 | 2 | -- |
+----- Margaritifera
--+
| +-- Cumberlandia_monodonta **
+--+
+-- Pseudunio
Bolotov et al., 2017 [Publ 3761], fig. S2 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 138 | 8 | 2 | -- |
-- Cumberlandia_monodonta **
Fukata & Iigo, 2020 [Publ 4106], fig. 1 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 53 | 1 | 1 | -- |
+-- Pseudunio
--+
+-- Cumberlandia_monodonta **
Pfeiffer & Graf, 2015 [Publ 3620], fig. 2 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 39 | 1 | 1 | -- |
+-- Margaritifera
--+
+-- Cumberlandia_monodonta **
Araujo et al., 2009 [Publ 2395], fig. 3 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 7 | 6 | 2 | -- |
+----- Cumberlandia_monodonta **
--+
| +-- Pseudunio
+--+
+-- Margaritifera
Huff et al., 2004 [Publ 2697], fig. 4 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 10 | 6 | 2 | -- |
+-- Pseudunio
|
--+-- Cumberlandia_monodonta **
|
+-- Margaritifera
|