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Page last updated
Thu 3 September 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Search the database pages:

Bivalvia | Unionida | Unionoidea | Margaritiferidae | Margaritiferinae

genus Cumberlandia Ortmann, 1912: cladomics
     1 Recent valid species (1 total species)

Cladomics Summary

number of species = 1 (1 total)
analyzed in 112 cladograms in 59 publications
maximum number of terminals in an analysis = 1 (mean = 1, median = 1)

Lyubas et al., 2019 [Publ 3963], fig. 10

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
110115122--
  +----- Margaritifera         
--+                            
  |  +-- Cumberlandia_monodonta **
  +--+                         
     +-- Pseudunio

Wu et al., 2020 [Publ 4121], fig. S1

iTtTbiriSoTroTroSiD
11013392--
  +----- Margaritifera         
--+                            
  |  +-- Cumberlandia_monodonta **
  +--+                         
     +-- Pseudunio

Guerra et al., 2019 [Publ 4168], fig. 1

iTtTbiriSoTroTroSiD
11014052--
  +----- Cumberlandia_monodonta **
--+                            
  |  +-- Pseudunio             
  +--+                         
     +-- Margaritifera

Huang et al., 2002 [Publ 2657], fig. 2

iTtTbiriSoTroTroSiD
11013311--
-- Cumberlandia_monodonta **

Graf & Cummings, 2006 [Publ 1968], fig. 2

iTtTbiriSoTroTroSiD
11014611--
  +-- Margaritifera         
--+                         
  +-- Cumberlandia_monodonta **

Guerra et al., 2017 [Publ 3772], fig. 2a

iTtTbiriSoTroTroSiD
11012411--
  +-- Margaritifera         
--+                         
  +-- Cumberlandia_monodonta **

Campbell et al., 2005 [Publ 2660], fig. 1

iTtTbiriSoTroTroSiD
110110111--
  +-- Margaritifera         
--+                         
  +-- Cumberlandia_monodonta **

Gomes-dos-Santos et al., 2019 [Publ 3943], fig. 1

iTtTbiriSoTroTroSiD
1101942--
  +----- Cumberlandia_monodonta **
--+                            
  |  +-- Pseudunio             
  +--+                         
     +-- Margaritifera

Roe & Hoeh, 2003 [Publ 2655], fig. 4f2

iTtTbiriSoTroTroSiD
11013011--
  +-- Cumberlandia_monodonta **
--+                         
  +-- Margaritifera

Lopes-Lima et al., 2018 [Publ 3829], fig. 3

iTtTbiriSoTroTroSiD
110127102--
  +----- Margaritifera         
--+                            
  |  +-- Cumberlandia_monodonta **
  +--+                         
     +-- Pseudunio

Inoue et al., 2018 [Publ 3858], fig. 1_3a-c

iTtTbiriSoTroTroSiD
110110300--
-- Cumberlandia_monodonta **

Rosenberg et al., 1997 [Publ 2720], fig. 2B

iTtTbiriSoTroTroSiD
11012121--
-- Cumberlandia_monodonta **

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iTtTbiriSoTroTroSiD
11019542--
  +----- Cumberlandia_monodonta **
--+                            
  |  +-- Pseudunio             
  +--+                         
     +-- Margaritifera

Huang et al., 2018 [Publ 3828], fig. 6-M

iTtTbiriSoTroTroSiD
11011600--
-- Cumberlandia_monodonta **

Hoeh et al., 2009 [Publ 2716], fig. 5

iTtTbiriSoTroTroSiD
11014511--
  +-- Margaritifera         
--+                         
  +-- Cumberlandia_monodonta **

Lopes-Lima et al., 2019 [Publ 3901], fig. 3

iTtTbiriSoTroTroSiD
11012811--
  +-- Cumberlandia_monodonta **
--+                         
  +-- Margaritifera

Dai et al., 2025 [Publ 4440], fig. 3

iTtTbiriSoTroTroSiD
11018742--
  +----- Cumberlandia_monodonta **
--+                            
  |  +-- Pseudunio             
  +--+                         
     +-- Margaritifera

Walker et al., 2006 [Publ 2666], fig. 3

iTtTbiriSoTroTroSiD
11011400--
-- Cumberlandia_monodonta **

Chase et al., 2018 [Publ 3861], fig. 1-M

iTtTbiriSoTroTroSiD
11011000--
-- Cumberlandia_monodonta **

Guerra et al., 2017 [Publ 3772], fig. 2b-M

iTtTbiriSoTroTroSiD
1101700--
-- Cumberlandia_monodonta **

Teiga-Teixeira et al., 2020 [Publ 4173], fig. 1

iTtTbiriSoTroTroSiD
11011811--
  +-- Margaritifera         
--+                         
  +-- Cumberlandia_monodonta **

Bogan & Hoeh, 2000 [Publ 2650], fig. 1

iTtTbiriSoTroTroSiD
11013311--
  +-- Cumberlandia_monodonta **
--+                         
  +-- Margaritifera

Graf & Ó Foighil, 2000 [Publ 2649], fig. 1_2

iTtTbiriSoTroTroSiD
11011000--
-- Cumberlandia_monodonta **

Guerra et al., 2017 [Publ 3772], fig. 2b

iTtTbiriSoTroTroSiD
11012411--
  +-- Margaritifera         
--+                         
  +-- Cumberlandia_monodonta **

Wu et al., 2021 [Publ 4366], fig. 6

iTtTbiriSoTroTroSiD
11019052--
  +----- Margaritifera         
--+                            
  |  +-- Cumberlandia_monodonta **
  +--+                         
     +-- Pseudunio

Sano et al., 2020 [Publ 4138], fig. 4

iTtTbiriSoTroTroSiD
11016792--
-- Cumberlandia_monodonta **

Vikhrev et al., 2018 [Publ 3812], fig. 3

iTtTbiriSoTroTroSiD
11011292--
  +----- Margaritifera         
--+                            
  |  +-- Cumberlandia_monodonta **
  +--+                         
     +-- Pseudunio

Hoeh et al., 2002 [Publ 2653], fig. 2

iTtTbiriSoTroTroSiD
11013011--
  +-- Margaritifera         
--+                         
  +-- Cumberlandia_monodonta **

Chase et al., 2018 [Publ 3847], fig. 4

iTtTbiriSoTroTroSiD
11013731--
  +-- Cumberlandia_monodonta **
--+                         
  +-- Margaritifera

Whelan et al., 2011 [Publ 3380], fig. 1-ML

iTtTbiriSoTroTroSiD
11013511--
  +-- Margaritifera         
--+                         
  +-- Cumberlandia_monodonta **

Chen et al., 2025 [Publ 4448], fig. 3

iTtTbiriSoTroTroSiD
11018542--
  +----- Cumberlandia_monodonta **
--+                            
  |  +-- Pseudunio             
  +--+                         
     +-- Margaritifera

Lopes-Lima et al., 2019 [Publ 3901], fig. 2

iTtTbiriSoTroTroSiD
11012711--
  +-- Margaritifera         
--+                         
  +-- Cumberlandia_monodonta **

Hoeh et al., 2001 [Publ 2652], fig. 14f4

iTtTbiriSoTroTroSiD
11013011--
  +-- Cumberlandia_monodonta **
--+                         
  +-- Margaritifera

Walker et al., 2006 [Publ 2666], fig. 2

iTtTbiriSoTroTroSiD
11011400--
-- Cumberlandia_monodonta **

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

iTtTbiriSoTroTroSiD
11017582--
-- Cumberlandia_monodonta **

Wu et al., 2019 [Publ 3864], fig. 3

iTtTbiriSoTroTroSiD
11014521--
  +-- Cumberlandia_monodonta **
--+                         
  +-- Margaritifera

Wu et al., 2021 [Publ 4366], fig. 4

iTtTbiriSoTroTroSiD
11016842--
  +----- Cumberlandia_monodonta **
--+                            
  |  +-- Pseudunio             
  +--+                         
     +-- Margaritifera

Araujo et al., 2016 [Publ 3746], fig. 2a

iTtTbiriSoTroTroSiD
11011092--
-- Cumberlandia_monodonta **

Bolotov et al., 2016 [Publ 3743], fig. 4

iTtTbiriSoTroTroSiD
11012982--
-- Cumberlandia_monodonta **

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B

iTtTbiriSoTroTroSiD
11012911--
  +-- Margaritifera         
--+                         
  +-- Cumberlandia_monodonta **

Wu et al., 2021 [Publ 4366], fig. 2_I

iTtTbiriSoTroTroSiD
11019052--
  +----- Margaritifera         
--+                            
  |  +-- Pseudunio             
  +--+                         
     +-- Cumberlandia_monodonta **

Wu et al., 2022 [Publ 4238], fig. 3

iTtTbiriSoTroTroSiD
11017342--
  +----- Cumberlandia_monodonta **
--+                            
  |  +-- Pseudunio             
  +--+                         
     +-- Margaritifera

Huang et al., 2018 [Publ 3828], fig. 6

iTtTbiriSoTroTroSiD
11014142--
  +----- Cumberlandia_monodonta **
--+                            
  |  +-- Pseudunio             
  +--+                         
     +-- Margaritifera

Zanatta & Murphy, 2006 [Publ 2672], fig. 2

iTtTbiriSoTroTroSiD
11015600--
-- Cumberlandia_monodonta **

Huang et al., 2002 [Publ 2657], fig. 1

iTtTbiriSoTroTroSiD
11013311--
-- Cumberlandia_monodonta **

Wu et al., 2020 [Publ 4121], fig. 3

iTtTbiriSoTroTroSiD
11014821--
  +-- Cumberlandia_monodonta **
--+                         
  +-- Margaritifera

Rosenberg et al., 1994 [Publ 2719], fig. 3

iTtTbiriSoTroTroSiD
11011821--
-- Cumberlandia_monodonta **

Huang et al., 2019 [Publ 3843], fig. 3

iTtTbiriSoTroTroSiD
11015542--
  +----- Cumberlandia_monodonta **
--+                            
  |  +-- Pseudunio             
  +--+                         
     +-- Margaritifera

Burzynski & Soroka, 2018 [Publ 3862], fig. 5a

iTtTbiriSoTroTroSiD
11011911--
  +-- Pseudunio             
--+                         
  +-- Cumberlandia_monodonta **

Sano et al., 2020 [Publ 4138], fig. 3

iTtTbiriSoTroTroSiD
11016792--
-- Cumberlandia_monodonta **

Graf, 2002 [Publ 2651], fig. 2

iTtTbiriSoTroTroSiD
11011700--
-- Cumberlandia_monodonta **

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
1101198102--
  +----- Margaritifera         
--+                            
  |  +-- Cumberlandia_monodonta **
  +--+                         
     +-- Pseudunio

Hoeh et al., 2009 [Publ 2716], fig. 4

iTtTbiriSoTroTroSiD
11014511--
  +-- Margaritifera         
--+                         
  +-- Cumberlandia_monodonta **

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A

iTtTbiriSoTroTroSiD
11012911--
  +-- Margaritifera         
--+                         
  +-- Cumberlandia_monodonta **

Wu et al., 2020 [Publ 4121], fig. 2

iTtTbiriSoTroTroSiD
11013392--
-- Cumberlandia_monodonta **

Roe & Hoeh, 2003 [Publ 2655], fig. 4f3

iTtTbiriSoTroTroSiD
11013011--
  +-- Cumberlandia_monodonta **
--+                         
  +-- Margaritifera

Lydeard et al., 2000 [Publ 2654], fig. 2c

iTtTbiriSoTroTroSiD
11011300--
-- Cumberlandia_monodonta **

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
1101200102--
  +-- Cumberlandia_monodonta **
  |                         
--+-- Pseudunio             
  |                         
  +-- Margaritifera

Lopes-Lima et al., 2018 [Publ 3829], fig. 1

iTtTbiriSoTroTroSiD
110127102--
  +----- Margaritifera         
--+                            
  |  +-- Cumberlandia_monodonta **
  +--+                         
     +-- Pseudunio

Huff et al., 2004 [Publ 2697], fig. 5-6

iTtTbiriSoTroTroSiD
11011162--
  +----- Margaritifera         
--+                            
  |  +-- Pseudunio             
  +--+                         
     +-- Cumberlandia_monodonta **

Lydeard et al., 1996 [Publ 2656], fig. 4

iTtTbiriSoTroTroSiD
11012721--
  +-- Cumberlandia_monodonta **
--+                         
  +-- Margaritifera

Whelan et al., 2011 [Publ 3380], fig. 1-MP

iTtTbiriSoTroTroSiD
11013511--
  +-- Margaritifera         
--+                         
  +-- Cumberlandia_monodonta **

Fukata & Iigo, 2020 [Publ 4106], fig. 1

iTtTbiriSoTroTroSiD
11015311--
  +-- Pseudunio             
--+                         
  +-- Cumberlandia_monodonta **

Marshall et al., 2014 [Publ 3593], fig. 1

iTtTbiriSoTroTroSiD
11011300--
-- Cumberlandia_monodonta **

Guerra et al., 2019 [Publ 4168], fig. 1-M

iTtTbiriSoTroTroSiD
11012122--
  +----- Cumberlandia_monodonta **
--+                            
  |  +-- Margaritifera         
  +--+                         
     +-- Pseudunio

Sano et al., 2020 [Publ 4138], fig. 2

iTtTbiriSoTroTroSiD
11016792--
-- Cumberlandia_monodonta **

Gomes-dos-Santos et al., 2019 [Publ 3943], fig. 1-M

iTtTbiriSoTroTroSiD
1101522--
  +----- Margaritifera         
--+                            
  |  +-- Pseudunio             
  +--+                         
     +-- Cumberlandia_monodonta **

Walker et al., 2006 [Publ 2666], fig. 6-M

iTtTbiriSoTroTroSiD
11011321--
  +-- Cumberlandia_monodonta **
--+                         
  +-- Margaritifera

Bolotov et al., 2017 [Publ 3761], fig. S2

iTtTbiriSoTroTroSiD
110113882--
-- Cumberlandia_monodonta **

Wu et al., 2021 [Publ 4366], fig. 3-ML

iTtTbiriSoTroTroSiD
11017342--
  +----- Cumberlandia_monodonta **
--+                            
  |  +-- Pseudunio             
  +--+                         
     +-- Margaritifera

Bolotov et al., 2015 [Publ 3646], fig. 2A

iTtTbiriSoTroTroSiD
1101872--
  +----------- Margaritifera_margaritifera
  |                                       
  |     +----- Margaritifera_dahurica__1  
  |     |                                 
  +-----+----- Margaritifera_dahurica__2  
--+     |                                 
  |     |  +-- Pseudunio                  
  |     +--+                              
  |        +-- Cumberlandia_monodonta **     
  |                                       
  |  +-------- Margaritifera_falcata      
  +--+                                    
     |  +----- Margaritifera_laevis       
     +--+                                 
        |  +-- Margaritifera_middendorffi 
        +--+                              
           +-- Margaritifera_marrianae

Chase et al., 2018 [Publ 3861], fig. 1

iTtTbiriSoTroTroSiD
11011000--
-- Cumberlandia_monodonta **

Lydeard et al., 2000 [Publ 2654], fig. 2b

iTtTbiriSoTroTroSiD
11011300--
-- Cumberlandia_monodonta **

Huff et al., 2004 [Publ 2697], fig. 4

iTtTbiriSoTroTroSiD
11011062--
  +-- Pseudunio             
  |                         
--+-- Cumberlandia_monodonta **
  |                         
  +-- Margaritifera

Machordom et al., 2003 [Publ 2688], fig. 3

iTtTbiriSoTroTroSiD
1101322--
  +----- Cumberlandia_monodonta **
--+                            
  |  +-- Margaritifera         
  +--+                         
     +-- Pseudunio

Bolotov et al., 2016 [Publ 3743], fig. 2

iTtTbiriSoTroTroSiD
11012982--
-- Cumberlandia_monodonta **

Aunins et al., 2018 [Publ 3863], fig. 1

iTtTbiriSoTroTroSiD
11013921--
  +-- Cumberlandia_monodonta **
--+                         
  +-- Margaritifera

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iTtTbiriSoTroTroSiD
11019542--
  +----- Cumberlandia_monodonta **
--+                            
  |  +-- Pseudunio             
  +--+                         
     +-- Margaritifera

Bolotov et al., 2015 [Publ 3646], fig. 2B

iTtTbiriSoTroTroSiD
1101872--
  +----- Cumberlandia_monodonta **
--+                            
  |  +-- Pseudunio             
  +--+                         
     +-- Margaritifera

Guerra et al., 2019 [Publ 4168], fig. 2

iTtTbiriSoTroTroSiD
11014052--
  +----- Cumberlandia_monodonta **
--+                            
  |  +-- Pseudunio             
  +--+                         
     +-- Margaritifera

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iTtTbiriSoTroTroSiD
11017911--
  +-- Cumberlandia_monodonta **
--+                         
  +-- Margaritifera

Zanatta & Murphy, 2006 [Publ 2672], fig. 1

iTtTbiriSoTroTroSiD
11015500--
-- Cumberlandia_monodonta **

Hoeh et al., 2001 [Publ 2652], fig. 14f2

iTtTbiriSoTroTroSiD
11013011--
  +-- Cumberlandia_monodonta **
--+                         
  +-- Margaritifera

Huang et al., 2018 [Publ 3828], fig. 3_5

iTtTbiriSoTroTroSiD
11012892--
  +----- Margaritifera         
--+                            
  |  +-- Cumberlandia_monodonta **
  +--+                         
     +-- Pseudunio

Inoue et al., 2013 [Publ 3579], fig. 3

iTtTbiriSoTroTroSiD
1101872--
  +----- Cumberlandia_monodonta **
--+                            
  |  +-- Pseudunio             
  +--+                         
     +-- Margaritifera

Campbell et al., 2005 [Publ 2660], fig. 2

iTtTbiriSoTroTroSiD
110110111--
  +-- Cumberlandia_monodonta **
--+                         
  +-- Margaritifera

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

iTtTbiriSoTroTroSiD
11017582--
-- Cumberlandia_monodonta **

Hoeh et al., 2009 [Publ 2716], fig. 3

iTtTbiriSoTroTroSiD
11014611--
  +-- Margaritifera         
--+                         
  +-- Cumberlandia_monodonta **

Wu et al., 2021 [Publ 4366], fig. 3-BI

iTtTbiriSoTroTroSiD
11017342--
  +----- Cumberlandia_monodonta **
--+                            
  |  +-- Pseudunio             
  +--+                         
     +-- Margaritifera

Lydeard et al., 2000 [Publ 2654], fig. 2a

iTtTbiriSoTroTroSiD
11011300--
-- Cumberlandia_monodonta **

Lydeard et al., 1996 [Publ 2656], fig. 3

iTtTbiriSoTroTroSiD
11012621--
  +-- Cumberlandia_monodonta **
--+                         
  +-- Margaritifera

Guerra et al., 2019 [Publ 4168], fig. 2-M

iTtTbiriSoTroTroSiD
11012122--
  +----- Pseudunio             
--+                            
  |  +-- Margaritifera         
  +--+                         
     +-- Cumberlandia_monodonta **

Walker et al., 2006 [Publ 2666], fig. 6

iTtTbiriSoTroTroSiD
11013552--
  +----- Cumberlandia_monodonta **
--+                            
  |  +-- Pseudunio             
  +--+                         
     +-- Margaritifera

Inoue et al., 2014 [Publ 3635], fig. 3A

iTtTbiriSoTroTroSiD
11011900--
-- Cumberlandia_monodonta **

Guerra et al., 2017 [Publ 3772], fig. 2a-M

iTtTbiriSoTroTroSiD
1101700--
-- Cumberlandia_monodonta **

Lydeard et al., 2000 [Publ 2654], fig. 2d

iTtTbiriSoTroTroSiD
11011300--
-- Cumberlandia_monodonta **

Chase et al., 2018 [Publ 3847], fig. 5

iTtTbiriSoTroTroSiD
1101900--
-- Cumberlandia_monodonta **

Takeuchi et al., 2015 [Publ 3661], fig. 3

iTtTbiriSoTroTroSiD
1101972--
  +----- Pseudunio_auricularius__1
  |                               
--+----- Pseudunio_auricularius__2
  |                               
  |  +-- Cumberlandia_monodonta **   
  +--+                            
     +-- Margaritifera

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

iTtTbiriSoTroTroSiD
11013911--
  +-- Margaritifera         
--+                         
  +-- Cumberlandia_monodonta **

Lopes-Lima et al., 2019 [Publ 3901], fig. 4

iTtTbiriSoTroTroSiD
11012511--
  +-- Margaritifera         
--+                         
  +-- Cumberlandia_monodonta **

Huff et al., 2004 [Publ 2697], fig. 3

iTtTbiriSoTroTroSiD
1101752--
  +-------- Margaritifera_margaritifera  
  |                                      
  |  +----- Cumberlandia_monodonta **       
--+  |                                   
  |  |  +-- Pseudunio                    
  |  |  |                                
  +--+  +-- Margaritifera_dahurica       
     |  |                                
     |  +-- Margaritifera_middendorffi__1
     |  |                                
     |  +-- Margaritifera_middendorffi__2
     +--+                                
        +-- Margaritifera_falcata__1     
        |                                
        +-- Margaritifera_falcata__2     
        |                                
        +-- Margaritifera_falcata__3     
        |                                
        +-- Margaritifera_falcata__4

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
1101200102--
  +----- Cumberlandia_monodonta **
--+                            
  |  +-- Pseudunio             
  +--+                         
     +-- Margaritifera

Walker et al., 2006 [Publ 2666], fig. 4

iTtTbiriSoTroTroSiD
11011400--
-- Cumberlandia_monodonta **

Chen et al., 2025 [Publ 4455], fig. 1

iTtTbiriSoTroTroSiD
110125102--
  +-- Cumberlandia_monodonta **
--+                         
  +-- Margaritifera

Araujo et al., 2009 [Publ 2395], fig. 3

iTtTbiriSoTroTroSiD
1101762--
  +----- Cumberlandia_monodonta **
--+                            
  |  +-- Pseudunio             
  +--+                         
     +-- Margaritifera

Bolotov et al., 2017 [Publ 3761], fig. S1

iTtTbiriSoTroTroSiD
110113882--
-- Cumberlandia_monodonta **

Araujo et al., 2016 [Publ 3746], fig. 1_2b

iTtTbiriSoTroTroSiD
11011292--
  +----- Margaritifera         
--+                            
  |  +-- Cumberlandia_monodonta **
  +--+                         
     +-- Pseudunio

Breton et al., 2011 [Publ 3376], fig. 3

iTtTbiriSoTroTroSiD
11013631--
  +-- Cumberlandia_monodonta **
--+                         
  +-- Margaritifera

Takeuchi et al., 2015 [Publ 3661], fig. 2

iTtTbiriSoTroTroSiD
1101972--
  +-------- Pseudunio_marocanus   
--+                               
  |  +----- Pseudunio_auricularius
  +--+                            
     |  +-- Cumberlandia_monodonta **
     +--+                         
        +-- Margaritifera

Zhou et al., 2007 [Publ 2658], fig. 1

iTtTbiriSoTroTroSiD
11012011--
  +-- Margaritifera         
--+                         
  +-- Cumberlandia_monodonta **

Huff et al., 2004 [Publ 2697], fig. 2

iTtTbiriSoTroTroSiD
11011162--
  +-------- Margaritifera_dahurica        
  |                                       
  +-------- Margaritifera_margaritifera__1
  |                                       
  +-------- Margaritifera_margaritifera__2
  |                                       
--+-------- Margaritifera_margaritifera__3
  |                                       
  |     +-- Pseudunio                     
  +-----+                                 
  |     +-- Cumberlandia_monodonta **        
  |                                       
  |  +----- Margaritifera_falcata         
  +--+                                    
     |  +-- Margaritifera_middendorffi    
     +--+                                 
        +-- Margaritifera_marrianae

Rosenberg et al., 1997 [Publ 2720], fig. 2A

iTtTbiriSoTroTroSiD
11012121--
-- Cumberlandia_monodonta **
 
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