The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)
FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!
Publication Campbell & Lydeard (2012): cladograms
publication summary | cladograms Complete CitationCampbell, D.C. & C. Lydeard. 2012. The genera of the Pleurobemini (Bivalvia: Unionidae: Ambleminae). American Malacological Bulletin 30(1): 19-38. Available electronically.
Cladogram Figures(click the figure number to see the cladogram) Cladogram Figure 1-2
+-- Uniomerus_declivis
+-----------------+
| +-- Uniomerus_tetralasmus
+--------------------+
| | +----------------- Theliderma_metanevra
| | |
| +--+ +-------------- Quadrula_verrucosa
| | |
| | | +-- Quadrula_quadrula
| +--+ +--------+
| | | +-- Quadrula_nobilis
| | |
| +--+ +-- Pustulosa_infucata
| | +-----+
| | | +-- Pustulosa_kleiniana
| +--+
| | +----- Cyclonaias_tuberculata
| +--+
| | +-- Pustulosa_pustulosa
| +--+
| +-- Pustulosa_succissa
|
| +----- Plectomerus_dombeyanus
| +-----------------+
| | | +-- Reginaia_ebenus
| | +--+
--+ | +-- Reginaia_rotulata
| |
| | +----- Popenaias_popeii
| +--------------+ +--------------+
| | | | | +-- Amblema_elliottii
| | | | +--+
| | | | +-- Amblema_plicata
| | | |
| | | | +----------------- Potamilus_alatus
| | +--+ |
| | | +----------------- Potamilus_capax
| | | |
| | | +----------------- Glebula_rotundata
| | | |
| | | | +-- Toxolasma_lividum
| | | +--------------+
| | +--+ +-- Paetulunio_fabalis
| | |
| | | +-- Dromus_dromas
| | +--------------+
| | | +-- Lemiox_rimosus
| | |
| | | +-- Lampsilis_ovata
+--+ | +-----------+
| | | +-- Obliquaria_reflexa
| +--+
| | +----------- Lampsilis_virescens
| | |
| +--+ +-- Epioblasma_rangiana
| | +-----+
| | | +-- Obovaria_arkansasensis
| +--+
| | +----- Obovaria_olivaria
| +--+
| | +-- Venustaconcha_ellipsiformis
| +--+
| +-- Venustaconcha_pleasii
|
| +----------------------------------- Elliptoideus_sloatianus
| |
| | +-------------------------------- Pleurobema_stabile
| | |
| | | +-- Pleurobema_clava
| | | +-----------+
+--+ | | +-- Pleurobema_oviforme
| | |
| | +--------------+ +----------- Pleurobema_pyriforme
| | | | |
| | | | | +-- Pleurobema_athearni
| | | | | +--+
+--+ | +--+ | +-- Pleurobema_georgianum
| | | +--+
| | | | | +-- Pleurobema_decisum
| | | | +--+
| | | | +-- Pleurobema_hanleyianum
| | +--+
| | | +----- Pleurobema_beadleianum
| | | |
| | | +----- Pleurobema_strodeanum
| | +--+
+--+ +----- Pleurobema_rubellum
| |
| | +-- Pleurobema_perovatum
| +--+
| +-- Pleurobema_taitianum
|
| +----- Pleurobema_riddellii
| +--------------------+
| | | +-- Pleurobema_sintoxia
| | +--+
| | +-- Pleurobema_cordatum
| |
| | +----------------- Fusconaia_subrotunda
| | |
| | +-----+ +-- Fusconaia_cor
+--+ | | +-----------+
| | | | +-- Fusconaia_cuneolus
| | +--+
| | | +----------- Fusconaia_ozarkensis
| | | |
| | +--+ +-- Fusconaia_escambia
| | | +-----+
| | | | +-- Fusconaia_burkei
| | +--+
+--+ | +----- Fusconaia_masoni
| +--+
| | +-- Fusconaia_flava
| +--+
| +-- Fusconaia_cerina
|
| +----- Pleuronaia_gibber
| +--------------+
| | | +-- Pleuronaia_barnesiana
| | +--+
| | +-- Pleuronaia_dolabelloides
+--+
| +-- Eurynia_dilatata
| +--------------+
| | +-- Parvaspina_collina
| |
+--+ +-- Hemistena_lata
| +-----------+
| | +-- Plethobasus_cyphyus
| |
+--+ +----------- Elliptio_shepardiana
| |
| | +-------- Elliptio_complanata
+--+ |
| | +----- Elliptio_purpurella
| +--+
| | | +-- Elliptio_arca
+--+ +--+
| +-- Elliptio_arctata
|
| +-- Elliptio_mcmichaeli
| |
+-----+-- Elliptio_pullata
|
+-- Elliptio_crassidens
Cladogram Figure 3
+-- Uniomerus_declivis
+--------------+
| +-- Uniomerus_tetralasmus
|
+--------------------+ +-- Quadrula_verrucosa
| | +-----------+
| | | +-- Quadrula_nobilis
| | |
| +--+ +----------- Quadrula_quadrula
| | |
| | +----------- Theliderma_metanevra
| +--+
| | +-- Pustulosa_infucata
| | +-----+
| | | +-- Pustulosa_kleiniana
| +--+
| | +----- Cyclonaias_tuberculata
| +--+
| | +-- Pustulosa_pustulosa
| +--+
| +-- Pustulosa_succissa
|
| +-- Amblema_elliottii
| +--+
| | +-- Amblema_plicata
| +--------------------------+
| | | +-- Reginaia_ebenus
--+ | +--+
| | +-- Reginaia_rotulata
| +--+
| | | +----------------------------- Popenaias_popeii
| | | |
| | | | +-- Glebula_rotundata
| | +--+ +-----------------------+
| | | | +-- Paetulunio_fabalis
| | | |
| | +--+ +-- Toxolasma_lividum
| | | +--------------------+
| | | | +-- Plectomerus_dombeyanus
| | | |
| | +--+ +-------------------- Obliquaria_reflexa
| | | |
| | | | +-- Dromus_dromas
| | | | +--+
| | +--+ | +-- Lemiox_rimosus
| | | +-----------+
| | | | | +-- Potamilus_alatus
| | | | +--+
| | +--+ +-- Potamilus_capax
| | |
| | | +-------------- Lampsilis_ovata
+--+ | |
| +--+ +----------- Lampsilis_virescens
| | |
| +--+ +-- Epioblasma_rangiana
| | +-----+
| | | +-- Obovaria_arkansasensis
| +--+
| | +----- Obovaria_olivaria
| +--+
| | +-- Venustaconcha_ellipsiformis
| +--+
| +-- Venustaconcha_pleasii
|
| +-------------------------------- Elliptoideus_sloatianus
| |
| | +-- Eurynia_dilatata
| | +--------------------------+
| | | +-- Parvaspina_collina
| | |
| | | +-- Hemistena_lata
| | +--------------------------+
| | | +-- Plethobasus_cyphyus
+--+ |
| | +----------- Elliptio_shepardiana
| | |
| | | +----- Elliptio_crassidens
| +-----------------+ +--+
| | | | | +-- Elliptio_mcmichaeli
| | | | +--+
| | +--+ +-- Elliptio_pullata
| | |
| | | +----- Elliptio_complanata
+--+ | |
| +--+ +-- Elliptio_purpurella
| | |
| +--+-- Elliptio_arca
| |
| +-- Elliptio_arctata
|
| +----- Pleuronaia_gibber
| +--------------------+
| | | +-- Pleuronaia_barnesiana
| | +--+
| | +-- Pleuronaia_dolabelloides
| |
| | +-------------- Fusconaia_ozarkensis
| | |
| | | +----- Fusconaia_subrotunda
| | +--------+ +-----+
+--+ | | | | +-- Fusconaia_cor
| | | | +--+
| | | | +-- Fusconaia_cuneolus
| | +--+
| | | +-- Fusconaia_escambia
| | | +-----+
| | | | +-- Fusconaia_burkei
| | +--+
| | | +----- Fusconaia_masoni
+--+ +--+
| | +-- Fusconaia_cerina
| +--+
| +-- Fusconaia_flava
|
| +----- Pleurobema_riddellii
| +--------------+
| | | +-- Pleurobema_sintoxia
| | +--+
| | +-- Pleurobema_cordatum
+--+
| +----------------- Pleurobema_stabile
| |
| | +----------- Pleurobema_strodeanum
| | +--+
+--+ | | +-------- Pleurobema_rubellum
| | +--+
| | | +----- Pleurobema_beadleianum
| | +--+
| | | +-- Pleurobema_perovatum
+--+ +--+
| +-- Pleurobema_taitianum
|
| +-- Pleurobema_clava
| +--------+
| | +-- Pleurobema_oviforme
+--+
| +-------- Pleurobema_pyriforme
| |
+--+ +-- Pleurobema_athearni
| +--+
| | +-- Pleurobema_georgianum
+--+
| +-- Pleurobema_decisum
+--+
+-- Pleurobema_hanleyianum
|
