The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)
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Publication ITIS (2012): taxonomic opinions
publication summary Complete CitationITIS. 2012. Integrated Taxonomic Information System. Available electronically.
Taxonomic OpinionsActinonaias carinata carinata Barnes, 1823 = Ortmanniana ligamentina Actinonaias carinata gibba Simpson = Ortmanniana ligamentina Actinonaias ellipsiformis Conrad = Venustaconcha ellipsiformis Actinonaias ligamentina (Lamarck, 1819) = Ortmanniana ligamentina Actinonaias pectorosa (Conrad, 1834) = Ortmanniana pectorosa Alasmidonta arcula (I. Lea, 1838) Alasmidonta atropurpurea (Rafinesque, 1831) Alasmidonta calceolus Lea = Pressodonta viridis Alasmidonta heterodon (I. Lea, 1830) = Prolasmidonta heterodon Alasmidonta marginata Say, 1818 Alasmidonta mccordi Athearn, 1964 Alasmidonta minor = Pressodonta viridis Alasmidonta raveneliana (I. Lea, 1834) Alasmidonta robusta Clarke, 1981 Alasmidonta triangulata (I. Lea, 1858) Alasmidonta undulata (Say, 1817) Alasmidonta varicosa (Lamarck, 1819) Alasmidonta viridis (Rafinesque, 1820) = Pressodonta viridis Alasmidonta wrightiana (Walker, 1901) Amblema costata Rafinesque, 1832 = Amblema plicata Amblema elliottii (I. Lea, 1856) Amblema neislerii (I. Lea, 1858) Amblema plicata (Say, 1817) Anodonta beringiana Middendorff, 1851 = Beringiana beringiana Anodonta californiensis I. Lea, 1852 Anodonta cataracta Say, 1817 = Pyganodon cataracta Anodonta cataracta marginata = Pyganodon cataracta Anodonta couperiana I. Lea, 1840 = Utterbackiana couperiana Anodonta dejecta Lewis, 1875 = Anodonta californiensis Anodonta gibbosa Say, 1824 = Pyganodon gibbosa Anodonta grandis corpulenta Cooper = Pyganodon grandis Anodonta grandis grandis Say, 1829 = Pyganodon grandis Anodonta grandis simpsoniana Lea = Pyganodon grandis Anodonta heardi M.E. Gordon & Hoeh, 1995 = Utterbackiana heardi Anodonta imbecillis Say, 1829 = Utterbackia imbecillis Anodonta implicata Say, 1829 = Utterbackiana implicata Anodonta kennerlyi I. Lea, 1860 Anodonta nuttalliana Lea, 1838 Anodonta oregonensis I. Lea, 1838 Anodonta peggyae Johnson, 1965 = Utterbackia peggyae Anodonta suborbiculata Say, 1831 = Utterbackiana suborbiculata Anodontoides ferussacianus (I. Lea, 1834) Anodontoides radiatus (Conrad, 1834) = Strophitus radiatus Arcidens confragosus (Say, 1829) Arkansia wheeleri Ortmann & Walker, 1912 = Arcidens wheeleri Corbicula fluminalis (O.F. Müller, 1774) Corbicula fluminea (O.F. Müller, 1774) Corbicula manilensis Philippi Cumberlandia monodonta (Say, 1829) Cyclonaias tuberculata (Rafinesque, 1820) Cyprogenia aberti (Conrad, 1850) Cyprogenia irrorata irrorata Lea = Cyprogenia stegaria Cyprogenia irrorata pusilla Simpson = Cyprogenia stegaria Cyprogenia stegaria (Rafinesque, 1820) Cyrtonaias tampicoensis tampicoensis (Lea, 1838) = Cyrtonaias tampicoensis Disconaias salinasensis (Simpson, 1908) = Disconaias fimbriata Dromus dromas caperatus Lea = Dromus dromas Dromus dromas dromas (Lea, 1834) = Dromus dromas Dysnomia nodicella [sic] = Epioblasma othcaloogensis Dysnomia stewardsoni Lea = Epioblasma stewardsonii Dysnomia sulcata sulcata Lea = Epioblasma obliquata Ellipsaria lineolata (Rafinesque, 1820) Elliptio ahenea (I. Lea, 1843) Elliptio angustata (I. Lea, 1831) Elliptio arca (Conrad, 1834) Elliptio arctata (Conrad, 1834) Elliptio beadleiana Lea = Pleurobema beadleianum Elliptio buckleyi (I. Lea, 1843) = Elliptio jayensis Elliptio chipolaensis (Walker, 1905) Elliptio cistellaeformis (Lea, 1863) Elliptio complanata (Lightfoot, 1786) Elliptio congaraea (I. Lea, 1831) Elliptio crassidens (Lamarck, 1819) Elliptio dariensis (I. Lea, 1842) Elliptio dilatata (Rafinesque, 1820) = Eurynia dilatata Elliptio downiei (I. Lea, 1858) Elliptio errans (I. Lea, 1834) [sic] = Elliptio icterina Elliptio fisheriana (I. Lea, 1838) Elliptio folliculata (I. Lea, 1838) Elliptio fraterna (I. Lea, 1852) Elliptio hepatica (I. Lea, 1859) = Elliptio icterina Elliptio hopetonensis (I. Lea, 1838) Elliptio icterina (Conrad, 1834) Elliptio jayensis (I. Lea, 1838) Elliptio judithae Clarke, 1986 = Elliptio roanokensis Elliptio lanceolata (I. Lea, 1828) Elliptio lugubris (I. Lea, 1834) = Elliptio icterina Elliptio marsupiobesa Fuller, 1972 Elliptio mcmichaeli Clench & Turner, 1956 Elliptio monichaeli [sic] = Elliptio mcmichaeli Elliptio monroensis (I. Lea, 1843) Elliptio nigella (I. Lea, 1852) Elliptio producta (Conrad, 1836) Elliptio raveneli (Conrad, 1834) = Elliptio icterina Elliptio roanokensis (I. Lea, 1838) Elliptio shepardiana (I. Lea, 1834) Elliptio spinosa (I. Lea, 1836) Elliptio steinstansana R.I. Johnson & Clarke, 1983 = Parvaspina steinstansana Elliptio waccamawensis (I. Lea, 1863) = Elliptio congaraea Elliptio waltoni (Wright, 1888) = Elliptio ahenea Elliptoideus sloatianus (I. Lea, 1840) Epioblasma arcaeformis (I. Lea, 1831) Epioblasma biemarginata (I. Lea, 1857) Epioblasma brevidens (I. Lea, 1831) Epioblasma capsaeformis (I. Lea, 1834) Epioblasma flexuosa (Rafinesque, 1820) Epioblasma florentina curtisii (Frierson & Utterback, 1916) = Epioblasma curtisii Epioblasma florentina florentina (I. Lea, 1857) = Epioblasma florentina Epioblasma florentina walkeri (Wilson & H.W. Clark, 1914) = Epioblasma walkeri Epioblasma haysiana (I. Lea, 1834) Epioblasma lenior (I. Lea, 1842) Epioblasma lewisii (Walker, 1910) Epioblasma metastriata (Conrad, 1838) Epioblasma obliquata obliquata (Rafinesque, 1820) = Epioblasma obliquata Epioblasma obliquata perobliqua (Conrad, 1836) = Epioblasma perobliqua Epioblasma othcaloogensis (I. Lea, 1857) Epioblasma penita (Conrad, 1834) Epioblasma personata (Say, 1829) Epioblasma propinqua (I. Lea, 1857) Epioblasma sampsonii (I. Lea, 1861) Epioblasma stewardsonii (I. Lea, 1852) Epioblasma torulosa gubernaculum (Reeve, 1865) = Epioblasma gubernaculum Epioblasma torulosa rangiana (I. Lea, 1838) = Epioblasma rangiana Epioblasma torulosa torulosa (Rafinesque, 1820) = Epioblasma torulosa Epioblasma triquetra (Rafinesque, 1820) Epioblasma turgidula (I. Lea, 1858) Eupera cubensis (Prime, 1865) Eupera singleyi Pilsbry Fusconaia askewi (Marsh, 1896) Fusconaia barnesiana (I. Lea, 1838) = Pleuronaia barnesiana Fusconaia cerina (Conrad, 1838) Fusconaia cor (Conrad, 1834) Fusconaia cuneolus (I. Lea, 1840) Fusconaia ebena (I. Lea, 1831) = Reginaia ebenus Fusconaia edgariana = Fusconaia cor Fusconaia escambia Clench & Turner, 1956 Fusconaia flava (Rafinesque, 1820) Fusconaia lananensis (Frierson, 1901) Fusconaia masoni (Conrad, 1834) Fusconaia ozarkensis (Call, 1887) Fusconaia subrotunda kirklandiana [sic] = Fusconaia subrotunda Fusconaia subrotunda lesueriana [sic] Lea = Fusconaia subrotunda Fusconaia subrotunda pilaris Lea = Fusconaia subrotunda Fusconaia subrotunda subrotunda (Lee, 1831) [sic] = Fusconaia subrotunda Fusconaia succissa (I. Lea, 1852) = Pustulosa succissa Fusconaia undata Barnes = Fusconaia flava Glebula rotundata (Lamarck, 1819) Gonidea angulata (I. Lea, 1838) Hemistena lata (Rafinesque, 1820) Lampsilis abrupta (Say, 1831) Lampsilis altilis (Conrad, 1834) = Hamiota altilis Lampsilis australis Simpson, 1900 = Hamiota australis Lampsilis binominata Simpson, 1900 Lampsilis bracteata (Gould, 1855) Lampsilis cardium Rafinesque, 1820 Lampsilis cariosa (Say, 1817) Lampsilis dolabraeformis (I. Lea, 1838) Lampsilis excavatus Lea = Lampsilis ornata Lampsilis fasciola Rafinesque, 1820 Lampsilis fullerkati R.I. Johnson, 1984 = Lampsilis radiata Lampsilis haddletoni Athearn, 1964 = Obovaria haddletoni Lampsilis higginsii (I. Lea, 1857) Lampsilis hydiana (I. Lea, 1838) Lampsilis ochracea Say = Atlanticoncha ochracea Lampsilis orbiculata Hildreth = Obovaria subrotunda Lampsilis ornata (Conrad, 1835) Lampsilis ovata (Say, 1817) Lampsilis perovalis (Conrad, 1834) = Hamiota perovalis Lampsilis perpasta Lea = Lampsilis straminea Lampsilis powellii (I. Lea, 1852) Lampsilis radiata conspicua (I. Lea, 1872) = Lampsilis radiata Lampsilis radiata radiata (Gmelin, 1791) = Lampsilis radiata Lampsilis rafinesqueana Frierson, 1927 Lampsilis reeveiana brevicula (Call, 1887) = Lampsilis reeveiana Lampsilis reeveiana brittsi Simpson, 1900 = Lampsilis brittsi Lampsilis reeveiana reeveiana (I. Lea, 1852) = Lampsilis reeveiana Lampsilis satura (I. Lea, 1852) Lampsilis siliquoidea (Barnes, 1823) Lampsilis splendida (I. Lea, 1838) Lampsilis straminea claibornensis (I. Lea, 1838) = Lampsilis straminea Lampsilis straminea straminea (Conrad, 1834) = Lampsilis straminea Lampsilis streckeri Frierson, 1927 Lampsilis subangulata (I. Lea, 1840) = Hamiota subangulata Lampsilis teres (Rafinesque, 1820) Lampsilis virescens (I. Lea, 1858) Lasmigona complanata alabamensis Clarke, 1985 = Lasmigona alabamensis Lasmigona complanata complanata (Barnes, 1823) = Lasmigona complanata Lasmigona compressa (I. Lea, 1829) = Platynaias compressa Lasmigona costata (Rafinesque, 1820) Lasmigona decorata (I. Lea, 1852) = Platynaias decorata Lasmigona holstonia (I. Lea, 1838) = Alasminota holstonia Lasmigona subviridis (Conrad, 1835) = Platynaias subviridis Lemiox rimosus (Rafinesque, 1831) Leptodea amphichaena Frierson = Potamilus amphichaenus Leptodea fragilis (Rafinesque, 1820) = Potamilus fragilis Leptodea laevissima Lea = Potamilus ohiensis Leptodea leptodon (Rafinesque, 1820) = Potamilus leptodon Leptodea ochracea (Say, 1817) = Atlanticoncha ochracea Lexingtonia dolabelloides conradi (Vanatta) = Pleuronaia dolabelloides Lexingtonia dolabelloides dolabelloides (Lea, 1840) = Pleuronaia dolabelloides Lexingtonia subplana (Conrad, 1837) = Fusconaia masoni Ligumia latissima = Ligumia recta Ligumia nasuta (Say, 1817) = Sagittunio nasutus Ligumia recta (Lamarck, 1819) Ligumia subrostrata (Say, 1831) = Sagittunio subrostratus Margaritifera falcata (Gould, 1850) Margaritifera hembeli (Conrad, 1838) Margaritifera margaritifera (Linnaeus, 1758) Margaritifera marrianae R.I. Johnson, 1983 Medionidus acutissimus (I. Lea, 1831) Medionidus conradicus (I. Lea, 1834) Medionidus mcglameriae van der Schalie, 1939 = Potamilus fragilis Medionidus parvulus (I. Lea, 1860) Medionidus penicillatus (I. Lea, 1857) Medionidus simpsonianus Walker, 1905 Medionidus walkeri (Wright, 1897) Megalonaias boykiniana (Lea, 1840) = Megalonaias nervosa Megalonaias gigantea = Megalonaias nervosa Megalonaias nervosa (Rafinesque, 1820) Megalonaias nicklineana [sic] = Megalonaias nickliniana Musculium lacustre (Müller, 1774) = Sphaerium lacustre Musculium partumeium (Say, 1822) = Sphaerium partumeium Musculium securis (Prime, 1852) = Sphaerium securis Musculium transversum (Say, 1829) = Sphaerium transversum Obliquaria reflexa Rafinesque, 1820 Obovaria jacksoniana (Frierson, 1912) = Obovaria arkansasensis Obovaria olivaria (Rafinesque, 1820) Obovaria retusa (Lamarck, 1819) Obovaria rotulata (Wright, 1899) = Reginaia rotulata Obovaria subrotunda (Rafinesque, 1820) Obovaria unicolor (I. Lea, 1845) Pegias fabula (I. Lea, 1838) Pisidium adamsi Stimpson, 1851 = Euglesa adamsii Pisidium amnicum (Muller, 1774) Pisidium casertanum (Poli, 1791) = Euglesa casertana Pisidium compressum Prime, 1852 = Euglesa compressa Pisidium conventus Clessin, 1877 = Conventus conventus Pisidium cruciatum Sterki, 1895 Pisidium dubium (Say, 1817) Pisidium equilaterale Prime, 1852 = Euglesa equilateralis Pisidium fallax Sterki, 1896 = Euglesa fallax Pisidium ferrugineum Prime, 1852 = Euglesa ferruginea Pisidium henslowanum (Sheppard, 1825) = Euglesa henslowana Pisidium idahoense E.W. Roper, 1890 = Pisidium idahoensis Pisidium insigne Gabb, 1868 = Conventus insigne Pisidium lilljeborgi Clessin, 1886 = Euglesa lilljeborgii Pisidium milium Held, 1836 = Euglesa milium Pisidium nitidum Jenyns, 1832 = Euglesa nitida Pisidium punctatum Sterki, 1895 Pisidium punctiferum (Guppy, 1867) Pisidium rotundatum Prime, 1852 = Euglesa rotundata Pisidium sanguinichristi Taylor, 1987 Pisidium subtruncatum Malm, 1855 = Euglesa subtruncata Pisidium supinum Schmidt, 1850 = Euglesa supina Pisidium ultramontanum Prime, 1865 = Euglesa ultramontana Pisidium variabile Prime, 1852 = Euglesa variabilis Pisidium ventricosum Prime, 1851 = Euglesa ventricosa Pisidium walkeri Sterki, 1895 = Euglesa walkeri Plectomerus dombeyanus (Valenciennes, 1827) Plethobasus cicatricosus (Say, 1829) Plethobasus cooperianus (I. Lea, 1834) Plethobasus cyphyus (Rafinesque, 1820) Pleurobema aldrichianum Goodrich = Pleurobema hanleyianum Pleurobema altum (Conrad, 1854) Pleurobema avellanum Simpson, 1900 = Pleurobema rubellum Pleurobema beadleianum (I. Lea, 1861) Pleurobema bournianum (I. Lea, 1840) = Pleurobema clava Pleurobema chattanoogaense (I. Lea, 1858) = Pleurobema decisum Pleurobema clava (Lamarck, 1819) Pleurobema coccineum (Conrad, 1834) = Pleurobema sintoxia Pleurobema collina (Conrad, 1837) = Parvaspina collina Pleurobema cordatum cordatum (Rafinesque, 1820) = Pleurobema cordatum Pleurobema curtum (I. Lea, 1859) Pleurobema decisum (I. Lea, 1831) Pleurobema flavidulum (I. Lea, 1861) = Pleurobema perovatum Pleurobema furvum (Conrad, 1834) = Pleurobema rubellum Pleurobema georgianum (I. Lea, 1841) Pleurobema gibberum (I. Lea, 1838) = Pleuronaia gibber Pleurobema hagleri (Frierson, 1900) = Pleurobema rubellum Pleurobema hanleyianum (I. Lea, 1852) Pleurobema irrasum Lea = Pleurobema georgianum Pleurobema johannis (I. Lea, 1859) = Pleurobema perovatum Pleurobema marshalli Frierson, 1927 Pleurobema meredithi Lea = Pleuronaia barnesiana Pleurobema murrayense (I. Lea, 1868) = Pleurobema stabile Pleurobema nucleopsis (Conrad, 1849) = Pleurobema georgianum Pleurobema nux Lea = Pleurobema perovatum Pleurobema oviforme (Conrad, 1834) Pleurobema perovatum (Conrad, 1834) Pleurobema plenum (I. Lea, 1840) Pleurobema pyramidatum (I. Lea, 1840) = Pleurobema sintoxia Pleurobema pyriforme (I. Lea, 1857) Pleurobema riddellii (I. Lea, 1861) Pleurobema rubellum (Conrad, 1834) Pleurobema rubrum (Rafinesque, 1820) = Pleurobema sintoxia Pleurobema showalteri Lea = Pleurobema hartmanianum Pleurobema similans [sic] = Pleurobema perovatum Pleurobema sintoxia (Rafinesque, 1820) Pleurobema stabile Lea Pleurobema strodeanum (Wright, 1898) Pleurobema taitianum (I. Lea, 1834) Pleurobema tombigbeanum Frierson = Pleurobema taitianum Pleurobema troschelianum (I. Lea, 1852) = Pleurobema georgianum Pleurobema verum (I. Lea, 1861) Polymesoda caroliniana (Bosc, 1801) Polymesoda maritima (d’Orbigny, 1842) = Polymesoda floridana Popenaias buckleyi Lea = Elliptio jayensis Popenaias popei (Lea, 1857) = Popenaias popeii Potamilus alatus (Say, 1817) Potamilus amphichaenus (Frierson, 1898) Potamilus capax (Green, 1832) Potamilus inflatus (I. Lea, 1831) Potamilus ohiensis (Rafinesque, 1820) Potamilus purpuratus (Lamarck, 1819) Proptera laevissimus Lea = Potamilus ohiensis Ptychobranchus fasciolaris (Rafinesque, 1820) Ptychobranchus greenii (Conrad, 1834) Ptychobranchus jonesi (van der Schalie, 1934) Ptychobranchus occidentalis (Conrad, 1836) Ptychobranchus subtentum (Say, 1825) = Ptychobranchus subtentus Pyganodon cataracta (Say, 1817) Pyganodon fragilis (Lamarck, 1819) Pyganodon gibbosa (Say, 1824) Pyganodon grandis (Say, 1829) Pyganodon lacustris (I. Lea, 1852) Quadrula apiculata (Say, 1829) = Quadrula quadrula Quadrula asperata (I. Lea, 1861) = Pustulosa kieneriana Quadrula aurea (I. Lea, 1859) = Pustulosa pustulosa Quadrula couchiana (I. Lea, 1860) Quadrula cylindrica cylindrica (Say, 1817) = Theliderma cylindrica Quadrula cylindrica strigillata (Wright, 1898) = Theliderma cylindrica Quadrula fragosa (Conrad, 1835) Quadrula houstonensis (I. Lea, 1859) = Pustulosa pustulosa Quadrula intermedia (Conrad, 1836) = Theliderma intermedia Quadrula metanevra (Rafinesque, 1820) = Theliderma metanevra Quadrula nodulata (Rafinesque, 1820) = Pustulosa nodulata Quadrula petrina (Gould, 1855) = Pustulosa petrina Quadrula pustulosa mortoni (Conrad, 1835) = Pustulosa pustulosa Quadrula pustulosa pustulosa (I. Lea, 1831) = Pustulosa pustulosa Quadrula quadrula quadrula (Rafinesque, 1820) = Quadrula quadrula Quadrula refulgens (I. Lea, 1868) = Pustulosa pustulosa Quadrula rumphiana (I. Lea, 1852) = Quadrula quadrula Quadrula sparsa (I. Lea, 1841) = Theliderma sparsa Quadrula stapes (I. Lea, 1831) = Theliderma stapes Quadrula tuberosa (I. Lea, 1840) = Theliderma metanevra Quincuncina burkei Walker, 1922 = Fusconaia burkei Quincuncina infucata (Conrad, 1834) = Pustulosa infucata Quincuncina mitchelli (Simpson, 1895) = Fusconaia mitchelli Simpsonaias ambigua (Say, 1825) Sphaerium corneum (Linnaeus, 1758) Sphaerium fabale (Prime, 1852) Sphaerium nitidum Westerlund, 1876 Sphaerium occidentale (J. Lewis, 1856) Sphaerium patella (Gould, 1850) Sphaerium rhomboideum (Say, 1822) Sphaerium simile (Say, 1817) Sphaerium striatinum (Lamarck, 1818) Strophitus alabamensis Lea = Pseudodontoideus connasaugaensis Strophitus connasaugaensis (I. Lea, 1858) = Pseudodontoideus connasaugaensis Strophitus subvexus (Conrad, 1834) = Pseudodontoideus subvexus Strophitus undulatus tennesseensis Frierson = Strophitus undulatus Strophitus undulatus undulatus (Say, 1817) = Strophitus undulatus Toxolasma corvunculus (I. Lea, 1868) Toxolasma cylindrellus (I. Lea, 1868) Toxolasma lividus Rafinesque, 1831 = Toxolasma lividum Toxolasma mearnsi (Simpson, 1900) = Toxolasma texasiense Toxolasma parvus (Barnes, 1823) = Toxolasma parvum Toxolasma paulus (I. Lea, 1840) = Toxolasma paulum Toxolasma pullus (Conrad, 1838) Toxolasma texasiensis (I. Lea, 1857) = Toxolasma texasiense Tritogonia nervosa Rafinesque = Megalonaias nervosa Tritogonia verrucosa (Rafinesque, 1820) = Quadrula verrucosa Truncilla cognata (I. Lea, 1860) Truncilla donaciformis (I. Lea, 1828) Truncilla macrodon (I. Lea, 1859) Truncilla truncata Rafinesque, 1820 Uniomerus carolinianus (Bosc, 1801) Uniomerus declivis (Say, 1831) Uniomerus excultus (Conrad, 1838) = Uniomerus declivis Uniomerus obesus (Lea, 1831) = Uniomerus carolinianus Uniomerus tetralasmus (Say, 1831) Utterbackia imbecillis (Say, 1829) Utterbackia peggyae (R.I. Johnson, 1965) Utterbackia peninsularis Bogan & Hoeh, 1995 Venustaconcha ellipsiformis (Conrad, 1836) Venustaconcha pleasii (Marsh, 1891) Villosa amygdala (I. Lea, 1843) = Villosa amygdalum Villosa arkansasensis (I. Lea, 1862) = Obovaria arkansasensis Villosa brevicula Call = Lampsilis reeveiana Villosa choctawensis Athearn, 1964 = Obovaria choctawensis Villosa constricta (Conrad, 1838) = Venustaconcha constricta Villosa delumbis (Conrad, 1834) Villosa fabalis (I. Lea, 1831) = Paetulunio fabalis Villosa iris (I. Lea, 1829) = Cambarunio iris Villosa lienosa (Conrad, 1834) = Leaunio lienosus Villosa nebulosa (Conrad, 1834) = Cambarunio nebulosus Villosa ortmanni (Walker, 1925) = Leaunio ortmanni Villosa perpurpurea (I. Lea, 1861) = Venustaconcha trabalis Villosa taeniata (Conrad, 1834) = Cambarunio taeniatus Villosa trabalis (Conrad, 1834) = Venustaconcha trabalis Villosa vanuxemensis umbrans (I. Lea, 1857) = Leaunio umbrans Villosa vanuxemensis vanuxemensis (I. Lea, 1838) = Leaunio vanuxemensis Villosa vaughaniana (I. Lea, 1838) = Sagittunio vaughanianus Villosa vibex (Conrad, 1834) Villosa villosa (Wright, 1898)
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