The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)
FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!
Search the database pages:
Bivalvia | Unionida | Unionoidea | Unionidae | Unioninae | Unionini | Oxynaiina genus Schistodesmus Simpson, 1900: cladomics
5 Recent valid species (6 total species) Cladomics Summarynumber of species = 5 (6 total)
analyzed in 96 cladograms in 57 publications
maximum number of terminals in an analysis =
5 (mean = 1.95, median = 2) Basic Monophylynumber of cladograms that test basic monophyly
(i.e., more than 1 included species) = 52
maximum number of included terminals = 5, (mean =
2.75 median = 3)
proportion of cladograms that support basic monophyly
= 0.98
Pairwise Cladogram ClusteringThe graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.
Wang et al., 2025 [Publ 4488], fig. 3B ** Top Monophyly Cladogram ** iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 5 | 60 | 18 | 4 | 0 | +-------------------- Pseudocuneopsis
|
--+ +----------------- Nodularia
| |
+--+ +-- Tchangsinaia
| +-----------+
| | +-- Cuneopsis
+--+
| +----------- Schistodesmus_spinosus **
+--+
| +-------- Schistodesmus_tongpenensis **
+--+
| +----- Schistodesmus_xinyuensis **
+--+
| +-- Schistodesmus_lampreyanus **
+--+
+-- Schistodesmus_luqiaoensis **
Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14 ** Top Sister-Group Cladogram ** iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 629 | 30 | 6 | -- | +----- Nodularia
+-----+
| | +-- Schistodesmus_lampreyanus **
| +--+
| +-- Schistodesmus_spinosus **
--+
| +-- Tchangsinaia
| +-----+
| | +-- Inversiunio
+--+
| +----- Pseudocuneopsis
+--+
| +-- Diaurora
+--+
+-- Cuneopsis
Wu et al., 2026 [Publ 4811], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 5 | 90 | 15 | 4 | 0 | +-------------------- Pseudocuneopsis
|
--+ +----------------- Nodularia
| |
+--+ +-- Tchangsinaia
| +-----------+
| | +-- Cuneopsis
+--+
| +----------- Schistodesmus_spinosus **
+--+
| +-------- Schistodesmus_tongpenensis **
+--+
| +----- Schistodesmus_xinyuensis **
+--+
| +-- Schistodesmus_lampreyanus **
+--+
+-- Schistodesmus_luqiaoensis **
Hou et al., 2025 [Publ 4445], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 5 | 48 | 17 | 4 | 0 | +-------------------- Pseudocuneopsis
|
--+ +----------------- Nodularia
| |
+--+ +-- Tchangsinaia
| +-----------+
| | +-- Cuneopsis
+--+
| +----------- Schistodesmus_spinosus **
+--+
| +-------- Schistodesmus_tongpenensis **
+--+
| +----- Schistodesmus_xinyuensis **
+--+
| +-- Schistodesmus_lampreyanus **
+--+
+-- Schistodesmus_luqiaoensis **
Hou et al., 2025 [Publ 4445], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 5 | 2 | 2 | 1 | 2 | +-------------- Cuneopsis
--+
| +----------- Schistodesmus_tongpenensis **
+--+
| +-------- Schistodesmus_spinosus **
+--+
| +----- Schistodesmus_xinyuensis **
+--+
| +-- Schistodesmus_lampreyanus **
+--+
+-- Schistodesmus_luqiaoensis **
Huang et al., 2019 [Publ 3843], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 1 | 4 | 196 | 8 | 3 | 2 | +----- Nodularia
+--+
| | +-- Tchangsinaia
| +--+
--+ +-- Cuneopsis
|
| +----- Schistodesmus_spinosus **
| |
+--+ +-- Schistodesmus_xinyuensis **
| |
+--+-- Schistodesmus_tongpenensis **
|
+-- Schistodesmus_lampreyanus **
Huang et al., 2019 [Publ 3843], fig. S1 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 43 | 8 | 3 | 1 | +----------------- Nodularia
--+
| +-------------- Tchangsinaia
+--+
| +----------- Cuneopsis
+--+
| +-------- Schistodesmus_spinosus **
+--+
| +----- Schistodesmus_tongpenensis **
+--+
| +-- Schistodesmus_lampreyanus **
+--+
+-- Schistodesmus_xinyuensis **
Huang et al., 2019 [Publ 3843], fig. S4 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 196 | 8 | 3 | 1 | +-------- Schistodesmus_spinosus **
--+
| +----- Schistodesmus_tongpenensis **
+--+
| +-- Schistodesmus_xinyuensis **
+--+
+-- Schistodesmus_lampreyanus **
Huang et al., 2019 [Publ 3843], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 41 | 8 | 3 | 1 | +-------- Schistodesmus_spinosus **
+--------+
| | +----- Schistodesmus_tongpenensis **
| +--+
| | +-- Schistodesmus_xinyuensis **
--+ +--+
| +-- Schistodesmus_lampreyanus **
|
| +-------------- Nodularia
+--+
| +----------- Cuneopsis_rufescens
+--+
| +-------- Tchangsinaia
+--+
| +----- Cuneopsis_celtiformis
+--+
| +-- Cuneopsis_szechenyii
+--+
+-- Cuneopsis_heudei
Chen et al., 2025 [Publ 4487], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 38 | 10 | 6 | 2 | +-- Tchangsinaia
+--------------+
| +-- Inversiunio
|
--+ +-- Cuneopsis
| +-----------+
| | +-- Pseudocuneopsis
+--+
| +----------- Nodularia
+--+
| +-------- Diaurora
+--+
| +----- Schistodesmus_spinosus **
+--+
| +-- Schistodesmus_xinyuensis **
+--+
+-- Schistodesmus_tongpenensis **
Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 138 | 18 | 4 | 2 | +-- Tchangsinaia
+-----------+
| +-- Cuneopsis
--+
| +----------- Pseudocuneopsis
+--+
| +-------- Nodularia
+--+
| +----- Schistodesmus_spinosus **
+--+
| +-- Schistodesmus_lampreyanus **
+--+
+-- Schistodesmus_xinyuensis **
Chen et al., 2023 [Publ 4302], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 38 | 11 | 6 | 2 | +-------- Diaurora
--+
| +----- Schistodesmus_spinosus **
+--+
| +-- Schistodesmus_xinyuensis **
+--+
+-- Schistodesmus_tongpenensis **
Wu et al., 2023 [Publ 4339], fig. 4_S5-aa iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 93 | 9 | 4 | 2 | +-- Nodularia
+-----------+
| +-- Pseudocuneopsis
--+
| +----------- Tchangsinaia
+--+
| +-------- Cuneopsis
+--+
| +----- Schistodesmus_spinosus **
+--+
| +-- Schistodesmus_lampreyanus **
+--+
+-- Schistodesmus_xinyuensis **
Wu et al., 2024 [Publ 4350], fig. 8b_S2 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 89 | 9 | 4 | 2 | +----------- Nodularia
|
+----------- Pseudocuneopsis
--+
| +-- Tchangsinaia
| +-----+
| | +-- Cuneopsis
+--+
| +----- Schistodesmus_spinosus **
+--+
| +-- Schistodesmus_lampreyanus **
+--+
+-- Schistodesmus_xinyuensis **
Chen et al., 2025 [Publ 4487], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 38 | 10 | 6 | 2 | +-- Tchangsinaia
+-----------------+
| +-- Inversiunio
--+
| +----------------- Pseudocuneopsis
+--+
| +-------------- Cuneopsis
+--+
| +----------- Nodularia
+--+
| +-------- Diaurora
+--+
| +----- Schistodesmus_spinosus **
+--+
| +-- Schistodesmus_xinyuensis **
+--+
+-- Schistodesmus_tongpenensis **
Wu et al., 2022 [Publ 4238], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 71 | 9 | 4 | 2 | +-------------- Pseudocuneopsis
|
--+ +----------- Nodularia
| |
+--+ +-- Tchangsinaia
| +-----+
| | +-- Cuneopsis
+--+
| +----- Schistodesmus_spinosus **
+--+
| +-- Schistodesmus_xinyuensis **
+--+
+-- Schistodesmus_lampreyanus **
Huang et al., 2019 [Publ 3843], fig. S3 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 195 | 8 | 3 | 2 | +----------- Nodularia
|
--+ +-- Tchangsinaia
| +-----+
| | +-- Cuneopsis
+--+
| +----- Schistodesmus_tongpenensis **
+--+
| +-- Schistodesmus_xinyuensis **
+--+
+-- Schistodesmus_lampreyanus **
Wu et al., 2022 [Publ 4238], fig. 2e iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 15 | 6 | 4 | 2 | +----------------- Pseudocuneopsis
--+
| +-------------- Nodularia
+--+
| +----------- Tchangsinaia
+--+
| +-------- Cuneopsis
+--+
| +----- Schistodesmus_spinosus **
+--+
| +-- Schistodesmus_lampreyanus **
+--+
+-- Schistodesmus_xinyuensis **
Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 119 | 15 | 4 | 2 | +----------- Nodularia
|
--+ +-------- Cuneopsis
| |
| | +-- Tchangsinaia
+--+-----+
| +-- Inversiunio
|
| +----- Schistodesmus_spinosus **
+--+
| +-- Schistodesmus_xinyuensis **
+--+
+-- Schistodesmus_tongpenensis **
Wu et al., 2023 [Publ 4339], fig. 4_S5-nt iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 8 | 6 | 3 | 93 | 9 | 4 | 9 | +-------------- Pseudocuneopsis
|
--+ +----------- Nodularia
| |
+--+ +-- Schistodesmus_lampreyanus **
| +-----+
| | +-- Cuneopsis
+--+
| +----- Tchangsinaia
+--+
| +-- Schistodesmus_xinyuensis **
+--+
+-- Schistodesmus_spinosus **
Dai et al., 2025 [Publ 4440], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 0 | 3 | 42 | 10 | 6 | 3 | +----- Diaurora
|
--+ +-- Schistodesmus_xinyuensis **
| |
+--+-- Schistodesmus_spinosus **
|
+-- Schistodesmus_tongpenensis **
Chen et al., 2023 [Publ 4302], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 38 | 11 | 6 | 2 | +-------- Diaurora
--+
| +----- Schistodesmus_spinosus **
+--+
| +-- Schistodesmus_xinyuensis **
+--+
+-- Schistodesmus_tongpenensis **
Campbell & Lydeard, 2026 [Publ 4810], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 44 | 6 | 4 | 2 | +----------------- Pseudocuneopsis
--+
| +-------------- Nodularia
+--+
| +----------- Tchangsinaia
+--+
| +-------- Cuneopsis
+--+
| +----- Schistodesmus_spinosus **
+--+
| +-- Schistodesmus_lampreyanus **
+--+
+-- Schistodesmus_xinyuensis **
Wu et al., 2024 [Publ 4350], fig. 8a_S1 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 89 | 9 | 4 | 2 | +----------- Nodularia
|
+----------- Pseudocuneopsis
--+
| +-- Tchangsinaia
| +-----+
| | +-- Cuneopsis
+--+
| +----- Schistodesmus_spinosus **
+--+
| +-- Schistodesmus_lampreyanus **
+--+
+-- Schistodesmus_xinyuensis **
Dai et al., 2025 [Publ 4440], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 42 | 10 | 6 | 2 | +-------- Diaurora
--+
| +----- Schistodesmus_spinosus **
+--+
| +-- Schistodesmus_xinyuensis **
+--+
+-- Schistodesmus_tongpenensis **
Dai et al., 2025 [Publ 4440], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 85 | 13 | 4 | 2 | +-- Nodularia
+--------+
| +-- Pseudocuneopsis
|
--+ +-- Tchangsinaia
| +-----+
| | +-- Cuneopsis
+--+
| +----- Schistodesmus_spinosus **
+--+
| +-- Schistodesmus_xinyuensis **
+--+
+-- Schistodesmus_tongpenensis **
Dai et al., 2024 [Publ 4358], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 50 | 11 | 4 | 2 | +-------------- Pseudocuneopsis
|
--+ +----------- Nodularia
| |
+--+ +-- Tchangsinaia
| +-----+
| | +-- Cuneopsis
+--+
| +----- Schistodesmus_spinosus **
+--+
| +-- Schistodesmus_lampreyanus **
+--+
+-- Schistodesmus_xinyuensis **
Wu et al., 2022 [Publ 4238], fig. 2d iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 36 | 12 | 5 | 2 | +-------------- Pseudocuneopsis
|
--+ +----------- Nodularia
| |
| | +----- Schistodesmus_spinosus **
+--+ +--+
| | | +-- Schistodesmus_lampreyanus **
| | +--+
+--+ +-- Schistodesmus_xinyuensis **
|
| +----- Cuneopsis
+--+
| +-- Tchangsinaia
+--+
+-- Inversiunio
Wu et al., 2018 [Publ 3836], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 51 | 5 | 3 | -- | +-- Tchangsinaia
+-----+
| +-- Cuneopsis
--+
| +----- Nodularia
+--+
| +-- Schistodesmus_xinyuensis **
+--+
+-- Schistodesmus_spinosus **
Zhang et al., 2004 [Publ 2735], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 12 | 5 | 4 | -- | +----------------- Tchangsinaia
--+
| +-------------- Cuneopsis_heudei
+--+
| +----------- Nodularia
+--+
| +-------- Pseudocuneopsis
+--+
| +----- Cuneopsis_rufescens
+--+
| +-- Schistodesmus_xinyuensis **
+--+
+-- Schistodesmus_spinosus **
Wu et al., 2021 [Publ 4366], fig. 3-BI iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 72 | 5 | 4 | -- | +----------- Pseudocuneopsis
|
--+ +-------- Nodularia
| |
+--+ +-- Schistodesmus_xinyuensis **
| +--+
| | +-- Schistodesmus_lampreyanus **
+--+
| +-- Cuneopsis
+--+
+-- Tchangsinaia
Hou et al., 2025 [Publ 4401], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 26 | 23 | 4 | -- | +----------- Pseudocuneopsis
|
--+ +-------- Nodularia
| |
+--+ +-- Schistodesmus_xinyuensis **
| +--+
| | +-- Schistodesmus_spinosus **
+--+
| +-- Tchangsinaia
+--+
+-- Cuneopsis
Pavan-Kumar et al., 2022 [Publ 4263], fig. S5 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 37 | 2 | 2 | -- | +-- Nodularia
+--+
| +-- Tchangsinaia
--+
| +-- Schistodesmus_lampreyanus **
+--+
+-- Schistodesmus_xinyuensis **
Liu et al., 2023 [Publ 4310], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 22 | 9 | 5 | -- | +-------- Cuneopsis
|
| +----- Nodularia
+--+
| | +-- Schistodesmus_xinyuensis **
--+ +--+
| +-- Schistodesmus_spinosus **
|
| +-- Tchangsinaia
| |
+-----+-- Inversiunio
|
+-- Pseudocuneopsis
Wu et al., 2020 [Publ 4147], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 28 | 4 | 4 | -- | +----------- Pseudocuneopsis
|
--+ +-------- Nodularia
| |
+--+ +-- Schistodesmus_lampreyanus **
| +--+
| | +-- Schistodesmus_xinyuensis **
+--+
| +-- Cuneopsis
+--+
+-- Tchangsinaia
Fukata & Iigo, 2020 [Publ 4106], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 52 | 4 | 4 | -- | +----- Pseudocuneopsis
+--+
| | +-- Nodularia
| +--+
--+ +-- Tchangsinaia
|
| +----- Cuneopsis
+--+
| +-- Schistodesmus_lampreyanus **
+--+
+-- Schistodesmus_xinyuensis **
Wu et al., 2021 [Publ 4366], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 89 | 4 | 3 | -- | +----------- Nodularia
--+
| +-------- Cuneopsis
+--+
| +----- Tchangsinaia
+--+
| +-- Schistodesmus_spinosus **
+--+
+-- Schistodesmus_xinyuensis **
Pavan-Kumar et al., 2022 [Publ 4263], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 35 | 2 | 2 | -- | +-- Nodularia
+--+
| +-- Tchangsinaia
--+
| +-- Schistodesmus_lampreyanus **
+--+
+-- Schistodesmus_xinyuensis **
Kim et al., 2021 [Publ 4197], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 16 | 5 | 2 | -- | +-------- Nodularia
--+
| +----- Cuneopsis
+--+
| +-- Schistodesmus_xinyuensis **
+--+
+-- Schistodesmus_lampreyanus **
Wu et al., 2021 [Publ 4366], fig. 3-ML iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 72 | 5 | 4 | -- | +-------------- Pseudocuneopsis
--+
| +----------- Nodularia
+--+
| +-------- Tchangsinaia
+--+
| +----- Cuneopsis
+--+
| +-- Schistodesmus_xinyuensis **
+--+
+-- Schistodesmus_lampreyanus **
Fukata & Iigo, 2019 [Publ 4108], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 15 | 6 | 4 | -- | +----------- Pseudocuneopsis
|
--+ +-- Tchangsinaia
| +-----+
| | +-- Nodularia
+--+
| +----- Cuneopsis
+--+
| +-- Schistodesmus_lampreyanus **
+--+
+-- Schistodesmus_xinyuensis **
Liu et al., 2023 [Publ 4310], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 22 | 9 | 5 | -- | +----------- Cuneopsis
|
| +----- Nodularia
--+ +--+
| | | +-- Schistodesmus_xinyuensis **
| | +--+
+--+ +-- Schistodesmus_spinosus **
|
| +----- Tchangsinaia
+--+
| +-- Inversiunio
+--+
+-- Pseudocuneopsis
Hou et al., 2024 [Publ 4400], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 74 | 22 | 4 | -- | +----------- Pseudocuneopsis
|
--+ +-------- Nodularia
| |
+--+ +-- Schistodesmus_xinyuensis **
| +--+
| | +-- Schistodesmus_spinosus **
+--+
| +-- Tchangsinaia
+--+
+-- Cuneopsis
Dai et al., 2024 [Publ 4386], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 40 | 9 | 4 | -- | +----------- Pseudocuneopsis
|
--+ +-------- Nodularia
| |
+--+ +-- Schistodesmus_lampreyanus **
| +--+
| | +-- Schistodesmus_xinyuensis **
+--+
| +-- Tchangsinaia
+--+
+-- Cuneopsis
Wu et al., 2021 [Publ 4366], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 67 | 5 | 4 | -- | +----------- Pseudocuneopsis
|
--+ +-------- Nodularia
| |
+--+ +-- Schistodesmus_xinyuensis **
| +--+
| | +-- Schistodesmus_lampreyanus **
+--+
| +-- Tchangsinaia
+--+
+-- Cuneopsis
Hou et al., 2025 [Publ 4432], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 26 | 23 | 4 | -- | +----------- Pseudocuneopsis
|
--+ +-------- Nodularia
| |
+--+ +-- Schistodesmus_xinyuensis **
| +--+
| | +-- Schistodesmus_spinosus **
+--+
| +-- Tchangsinaia
+--+
+-- Cuneopsis
Wu et al., 2024 [Publ 4379], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 52 | 17 | 4 | -- | +----------- Pseudocuneopsis
|
--+ +-------- Nodularia
| |
+--+ +-- Schistodesmus_spinosus **
| +--+
| | +-- Schistodesmus_xinyuensis **
+--+
| +-- Tchangsinaia
+--+
+-- Cuneopsis
Zhou et al., 2007 [Publ 2658], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 19 | 2 | 2 | -- | +-- Schistodesmus_xinyuensis **
--+
+-- Schistodesmus_spinosus **
Wu et al., 2024 [Publ 4379], fig. 4_6 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 21 | 10 | 1 | -- | +----- Nodularia
--+
| +-- Schistodesmus_xinyuensis **
+--+
+-- Schistodesmus_spinosus **
Chen et al., 2025 [Publ 4448], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 84 | 17 | 5 | -- | +-------------- Pseudocuneopsis
|
--+ +----------- Nodularia
| |
+--+ +-- Tchangsinaia
| +-----+
| | +-- Cuneopsis
+--+
| +----- Tongchuania
+--+
| +-- Schistodesmus_xinyuensis **
+--+
+-- Schistodesmus_lampreyanus **
Wu et al., 2021 [Publ 4366], fig. 2_I iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 89 | 4 | 3 | -- | +----------- Nodularia
--+
| +-------- Cuneopsis
+--+
| +----- Tchangsinaia
+--+
| +-- Schistodesmus_xinyuensis **
+--+
+-- Schistodesmus_spinosus **
Wu et al., 2023 [Publ 4296], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 21 | 8 | 5 | -- | +----------- Cuneopsis
|
| +----- Nodularia
--+ +--+
| | | +-- Schistodesmus_xinyuensis **
| | +--+
+--+ +-- Schistodesmus_spinosus **
|
| +----- Tchangsinaia
+--+
| +-- Inversiunio
+--+
+-- Pseudocuneopsis
Dai et al., 2024 [Publ 4386], fig. 5-BI iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 21 | 8 | 3 | -- | +----- Cuneopsis
|
--+----- Nodularia
|
| +-- Inversiunio
+--+
+-- Schistodesmus_xinyuensis **
Wu et al., 2020 [Publ 4121], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 48 | 4 | 4 | -- | +----------- Pseudocuneopsis
--+
| +-------- Nodularia
+--+
| +----- Tchangsinaia
+--+
| +-- Cuneopsis
+--+
+-- Schistodesmus_xinyuensis **
Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 75 | 5 | 3 | -- | +----------- Nodularia
--+
| +-------- Schistodesmus_xinyuensis **
+--+
| +----- Cuneopsis_szechenyii
+--+
| +-- Tchangsinaia
+--+
+-- Cuneopsis_rufescens
Froufe et al., 2019 [Publ 3936], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 43 | 2 | 2 | -- | +----- Nodularia
--+
| +-- Tchangsinaia
+--+
+-- Schistodesmus_lampreyanus **
Sano et al., 2020 [Publ 4138], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 67 | 6 | 3 | -- | +-------- Inversiunio
--+
| +----- Nodularia
+--+
| +-- Schistodesmus_lampreyanus **
+--+
+-- Tchangsinaia
Xue et al., 2016 [Publ 3659], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 14 | 1 | 1 | -- | -- Schistodesmus_lampreyanus **
Guerra et al., 2019 [Publ 4168], fig. 1-M iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 21 | 0 | 0 | -- | -- Schistodesmus_lampreyanus **
Soroka & Burzynski, 2017 [Publ 3837], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 22 | 2 | 2 | -- | +----- Schistodesmus_lampreyanus **
--+
| +-- Tchangsinaia
+--+
+-- Nodularia
Huang et al., 2002 [Publ 2657], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 33 | 2 | 2 | -- | -- Schistodesmus_xinyuensis **
Wu et al., 2019 [Publ 3864], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 45 | 4 | 4 | -- | +----------- Pseudocuneopsis
--+
| +-------- Nodularia
+--+
| +----- Tchangsinaia
+--+
| +-- Cuneopsis
+--+
+-- Schistodesmus_xinyuensis **
Bolotov et al., 2017 [Publ 3761], fig. S2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 137 | 5 | 3 | -- | -- Schistodesmus_xinyuensis **
Sano et al., 2020 [Publ 4138], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 67 | 6 | 3 | -- | +-------- Inversiunio
--+
| +----- Nodularia
+--+
| +-- Tchangsinaia
+--+
+-- Schistodesmus_lampreyanus **
Bolotov et al., 2017 [Publ 3764], fig. S1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 123 | 5 | 3 | -- | -- Schistodesmus_xinyuensis **
Froufe et al., 2019 [Publ 3936], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 43 | 2 | 2 | -- | +----- Nodularia
--+
| +-- Tchangsinaia
+--+
+-- Schistodesmus_lampreyanus **
Zhou et al., 2016 [Publ 3710], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 19 | 0 | 0 | -- | -- Schistodesmus_lampreyanus **
Huang et al., 2019 [Publ 3843], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 55 | 2 | 2 | -- | +----- Nodularia
--+
| +-- Tchangsinaia
+--+
+-- Schistodesmus_lampreyanus **
Liu et al., 2017 [Publ 3939], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 14 | 6 | 2 | -- | +-- Schistodesmus_xinyuensis **
|
--+-- Cuneopsis
|
+-- Nodularia
Dai et al., 2024 [Publ 4358], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 5 | 5 | 2 | -- | +----- Pseudocuneopsis
--+
| +-- Schistodesmus_xinyuensis **
+--+
+-- Nodularia
Guerra et al., 2019 [Publ 4168], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 40 | 0 | 0 | -- | -- Schistodesmus_lampreyanus **
Wu et al., 2022 [Publ 4238], fig. 2a_S1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 11 | 10 | 5 | -- | +-------------- Nodularia
--+
| +----------- Pseudocuneopsis
+--+
| +-------- Schistodesmus **
+--+
| +----- Cuneopsis
+--+
| +-- Tchangsinaia
+--+
+-- Inversiunio
Cha et al., 2018 [Publ 3916], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 37 | 2 | 2 | -- | +----- Schistodesmus_lampreyanus **
--+
| +-- Nodularia
+--+
+-- Tchangsinaia
Aunins et al., 2018 [Publ 3863], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 39 | 3 | 3 | -- | +-------- Nodularia
--+
| +----- Tchangsinaia
+--+
| +-- Cuneopsis
+--+
+-- Schistodesmus_lampreyanus **
Araujo et al., 2017 [Publ 3801], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 13 | 0 | 0 | -- | -- Schistodesmus_lampreyanus **
Dai et al., 2024 [Publ 4386], fig. 5-ML iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 21 | 8 | 3 | -- | +-------- Nodularia
--+
| +----- Cuneopsis
+--+
| +-- Inversiunio
+--+
+-- Schistodesmus_xinyuensis **
Choi et al., 2020 [Publ 4169], fig. 2a iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 11 | 10 | 3 | -- | -- Schistodesmus_xinyuensis **
Guerra et al., 2019 [Publ 4168], fig. 2-M iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 21 | 0 | 0 | -- | -- Schistodesmus_lampreyanus **
Lopes-Lima et al., 2017 [Publ 3792], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 31 | 2 | 2 | -- | +----- Schistodesmus_lampreyanus **
--+
| +-- Tchangsinaia
+--+
+-- Nodularia
Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 75 | 5 | 3 | -- | +----------- Nodularia
--+
| +-------- Schistodesmus_xinyuensis **
+--+
| +----- Cuneopsis_szechenyii
+--+
| +-- Tchangsinaia
+--+
+-- Cuneopsis_rufescens
Chen et al., 2020 [Publ 4170], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 33 | 0 | 0 | -- | -- Schistodesmus_lampreyanus **
Sano et al., 2020 [Publ 4138], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 67 | 6 | 3 | -- | +-------- Nodularia
--+
| +----- Inversiunio
+--+
| +-- Schistodesmus_lampreyanus **
+--+
+-- Tchangsinaia
Dai et al., 2023 [Publ 4285], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 52 | 4 | 4 | -- | +-- Pseudocuneopsis
+-----+
| +-- Nodularia
--+
| +----- Schistodesmus_xinyuensis **
+--+
| +-- Cuneopsis
+--+
+-- Tchangsinaia
Ouyang et al., 2015 [Publ 4493], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 30 | 6 | 3 | -- | -- Schistodesmus_xinyuensis **
Huang et al., 2002 [Publ 2657], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 33 | 2 | 2 | -- | -- Schistodesmus_xinyuensis **
Wu et al., 2024 [Publ 4388], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 52 | 6 | 4 | -- | +----------- Pseudocuneopsis
--+
| +-------- Nodularia
+--+
| +----- Tchangsinaia
+--+
| +-- Cuneopsis
+--+
+-- Schistodesmus_xinyuensis **
Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 79 | 5 | 3 | -- | +-------- Nodularia
--+
| +----- Schistodesmus_xinyuensis **
+--+
| +-- Tchangsinaia
+--+
+-- Cuneopsis
Sano et al., 2020 [Publ 4138], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 33 | 5 | 3 | -- | +-- Schistodesmus_xinyuensis **
--+
+-- Inversiunio
Bogan et al., 2023 [Publ 4295], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 30 | 0 | 0 | -- | -- Schistodesmus_xinyuensis **
Sano et al., 2017 [Publ 3799], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 32 | 7 | 3 | -- | -- Schistodesmus_xinyuensis **
Guerra et al., 2019 [Publ 4168], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 40 | 0 | 0 | -- | -- Schistodesmus_lampreyanus **
Bolotov et al., 2017 [Publ 3761], fig. S1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 137 | 5 | 3 | -- | +-------- Nodularia
--+
| +----- Cuneopsis
+--+
| +-- Tchangsinaia
+--+
+-- Schistodesmus_xinyuensis **
Ouyang et al., 2015 [Publ 4493], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 30 | 6 | 3 | -- | +----------------- Nodularia
|
| +----- Cuneopsis_rufescens__2
--+ +--------+
| | | +-- Schistodesmus_xinyuensis **
| | +--+
+--+ +-- Cuneopsis_celtiformis__2
|
| +----------- Tchangsinaia
+--+
| +-------- Cuneopsis_celtiformis__1
+--+
| +----- Cuneopsis_szechenyii__2
+--+
| +-- Cuneopsis_szechenyii__1
+--+
+-- Cuneopsis_rufescens__1
Huang et al., 2019 [Publ 3843], fig. 3-M iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 22 | 0 | 0 | -- | -- Schistodesmus_lampreyanus **
Zieritz et al., 2021 [Publ 4231], fig. 3-F_5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 47 | 2 | 2 | -- | +----- Nodularia
--+
| +-- Schistodesmus_lampreyanus **
+--+
+-- Tchangsinaia
Fonseca et al., 2016 [Publ 3775], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 30 | 2 | 2 | -- | +----- Schistodesmus_lampreyanus **
--+
| +-- Nodularia
+--+
+-- Tchangsinaia
|
