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Thu 3 September 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

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genus Diplodon Spix in Wagner, 1827: cladomics
43 Recent valid species (43 total species)

Cladomics Summary

number of species = 43 (43 total)
analyzed in 35 cladograms in 22 publications
maximum number of terminals in an analysis = 3 (mean = 1.26, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 8
maximum number of included terminals = 3, (mean = 2.13 median = 2)
proportion of cladograms that support basic monophyly = 0.63

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination

NJ Tree

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

tT	bi	riS	oT	roT	roS	iD
3	26	32	4	629	0	0	0

           +----------------------- Diplodon_suavidicus__2 **
           |                                              
  +--------+                    +-- Triplodon             
  |        |  +-----------------+                         
  |        |  |                 +-- Prisodon              
  |        +--+                                           
  |           |  +----------------- Castalia_ambigua__4   
  |           |  |                                        
  |           +--+  +-------------- Callonaia             
  |              |  |                                     
  |              +--+  +----------- Castalia_stevensi     
  |                 |  |                                  
  |                 +--+  +-------- Castalia_ambigua__3   
  |                    |  |                               
  |                    +--+     +-- Castalia_ambigua__1   
  |                       |  +--+                         
--+                       |  |  +-- Castaliella           
  |                       +--+                            
  |                          |  +-- Castalia_ambigua__2   
  |                          +--+                         
  |                             +-- Castalia              
  |                                                       
  |                             +-- Hyridella_depressa    
  |                       +-----+                         
  |                       |     +-- Hyridella_drapeta     
  |  +--------------------+                               
  |  |                    |  +----- Diplodon_suavidicus__1 **
  |  |                    +--+                            
  |  |                       |  +-- Diplodon_chilensis **    
  |  |                       +--+                         
  +--+                          +-- Diplodon_gratus **       
     |                                                    
     |                          +-- Cucumerunio           
     |  +-----------------------+                         
     |  |                       +-- Hyridella_australis   
     |  |                                                 
     +--+  +----------------------- Echyridella           
        |  |                                              
        |  |              +-------- Velesunio__1          
        +--+  +-----------+                               
           |  |           |  +----- Alathyria_jacksoni__1 
           |  |           +--+                            
           |  |              |  +-- Velesunio_ambiguus    
           +--+              +--+                         
              |                 +-- Alathyria_profuga     
              |                                           
              |  +----------------- Velesunio__3          
              |  |                                        
              +--+  +-------------- Microdontia           
                 |  |                                     
                 +--+           +-- Lortiella             
                    |  +--------+                         
                    |  |        +-- Velesunio_angasi      
                    +--+                                  
                       |  +-------- Alathyria_pertexta__2 
                       +--+                               
                          |  +----- Velesunio__2          
                          +--+                            
                             |  +-- Alathyria_jacksoni__2 
                             +--+                         
                                +-- Alathyria_pertexta__1

Marshall et al., 2014 [Publ 3593], fig. 1

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	12	0	0	--

  +-- Diplodon_chilensis **
--+                     
  +-- Diplodon_gratus **

Santos-Neto et al., 2016 [Publ 3739], fig. 4

tT	bi	riS	oT	roT	roS	iD
2	19	0	3	18	0	0	21.5

     +----------------------- Diplodon_suavidicus__2 **
     |                                              
  +--+                    +-- Triplodon             
  |  |  +-----------------+                         
  |  |  |                 +-- Prisodon              
  |  +--+                                           
  |     |  +----------------- Castalia_ambigua__4   
  |     +--+                                        
  |        |  +-------------- Callonaia             
  |        +--+                                     
  |           |  +----------- Castalia_ambigua__3   
  |           +--+                                  
  |              |  +-------- Castalia_stevensi     
  |              +--+                               
--+                 |  +----- Castalia_ambigua__2   
  |                 +--+                            
  |                    |  +-- Castaliella           
  |                    +--+                         
  |                       +-- Castalia_ambigua__1   
  |                                                 
  |                 +-------- Westralunio           
  |                 |                               
  |     +-----------+     +-- Hyridella_drapeta     
  |     |           |  +--+                         
  |     |           |  |  +-- Hyridella_depressa    
  |     |           +--+                            
  |     |              |  +-- Diplodon_suavidicus__1 **
  +-----+              +--+                         
        |                 +-- Diplodon_gratus **       
        |                                           
        |  +----------------- Echyridella           
        |  |                                        
        +--+              +-- Cucumerunio           
           |  +-----------+                         
           |  |           +-- Hyridella_australis   
           +--+                                     
              |  +----------- Velesunio_ambiguus    
              +--+                                  
                 |  +-------- Alathyria             
                 +--+                               
                    |  +----- Velesunio_angasi      
                    +--+                            
                       |  +-- Lortiella             
                       +--+                         
                          +-- Microdontia

Hoeh et al., 2009 [Publ 2716], fig. 5

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	44	0	0	--

  +-- Diplodon_chilensis **
--+                     
  +-- Diplodon_gratus **

Graf & Cummings, 2006 [Publ 1968], fig. 2

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	45	0	0	--

  +-- Diplodon_chilensis **
--+                     
  +-- Diplodon_gratus **

Santos-Neto et al., 2016 [Publ 3739], fig. 2a_b

tT	bi	riS	oT	roT	roS	iD
2	19	0	3	18	0	0	20.5

                 +-------- Westralunio           
                 |                               
  +--------------+     +-- Hyridella_drapeta     
  |              |  +--+                         
  |              |  |  +-- Hyridella_depressa    
  |              +--+                            
  |                 |  +-- Diplodon_gratus **       
  |                 +--+                         
  |                    +-- Diplodon_suavidicus__2 **
  |                                              
  |                    +-- Hyridella_australis   
  |     +--------------+                         
  |     |              +-- Cucumerunio           
  +-----+                                        
  |     |  +-------------- Echyridella           
--+     +--+                                     
  |        |  +----------- Velesunio_ambiguus    
  |        +--+                                  
  |           |  +-------- Microdontia           
  |           +--+                               
  |              |  +----- Lortiella             
  |              +--+                            
  |                 |  +-- Alathyria             
  |                 +--+                         
  |                    +-- Velesunio_angasi      
  |                                              
  |  +-------------------- Diplodon_suavidicus__1 **
  |  |                                           
  +--+                 +-- Triplodon             
     |  +--------------+                         
     |  |              +-- Prisodon              
     +--+                                        
        |  +-------------- Castalia_stevensi     
        |  |                                     
        +--+           +-- Callonaia             
           |  +--------+                         
           |  |        +-- Castalia_ambigua__1   
           +--+                                  
              |  +-------- Castalia_ambigua__4   
              +--+                               
                 |  +----- Castalia_ambigua__2   
                 +--+                            
                    |  +-- Castaliella           
                    +--+                         
                       +-- Castalia_ambigua__3

Hoeh et al., 2009 [Publ 2716], fig. 3

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	45	0	0	--

  +-- Diplodon_chilensis **
--+                     
  +-- Diplodon_gratus **

Hoeh et al., 2009 [Publ 2716], fig. 4

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	44	0	0	--

  +-- Diplodon_chilensis **
--+                     
  +-- Diplodon_gratus **

Hoeh et al., 2001 [Publ 2652], fig. 14f2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	30	0	0	--

-- Diplodon_gratus **

Guerra et al., 2019 [Publ 4168], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	40	0	0	--

-- Diplodon_suavidicus **

Baker et al., 2004 [Publ 2685], fig. 2-3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	8	0	0	--

-- Diplodon_gratus **

Graf, 2002 [Publ 2651], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	17	0	0	--

-- Diplodon_chilensis **

Bolotov et al., 2016 [Publ 3743], fig. 4

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	29	0	0	--

-- Diplodon_suavidicus **

Baker et al., 2003 [Publ 2690], fig. 5

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	7	0	0	--

-- Diplodon_gratus **

Hoeh et al., 2002 [Publ 2653], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	30	0	0	--

-- Diplodon_gratus **

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	140	0	0	--

-- Diplodon_suavidicus **

Walker et al., 2006 [Publ 2666], fig. 6

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	35	0	0	--

-- Diplodon_gratus **

Graf et al., 2015 [Publ 3619], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	19	0	0	--

-- Diplodon_suavidicus **

Bolotov et al., 2017 [Publ 3761], fig. S2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	137	0	0	--

-- Diplodon_suavidicus **

Bolotov et al., 2017 [Publ 3761], fig. S1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	137	0	0	--

-- Diplodon_suavidicus **

Huang et al., 2019 [Publ 3843], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	199	0	0	--

-- Diplodon_suavidicus **

Roe & Hoeh, 2003 [Publ 2655], fig. 4f2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	30	0	0	--

-- Diplodon_gratus **

Bogan & Hoeh, 2000 [Publ 2650], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	33	0	0	--

-- Diplodon_gratus **

Hoeh et al., 2001 [Publ 2652], fig. 14f4

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	30	0	0	--

-- Diplodon_gratus **

Wu et al., 2021 [Publ 4366], fig. 6

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	90	0	0	--

-- Diplodon_suavidicus **

Konopleva et al., 2017 [Publ 3807], fig. 6

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	30	0	0	--

-- Diplodon_suavidicus **

Roe & Hoeh, 2003 [Publ 2655], fig. 4f3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	30	0	0	--

-- Diplodon_gratus **

Huang et al., 2019 [Publ 3843], fig. S3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	197	0	0	--

-- Diplodon_suavidicus **

Santos-Neto et al., 2016 [Publ 3739], fig. 3_5

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	6	0	0	--

-- Diplodon_suavidicus **

Guerra et al., 2019 [Publ 4168], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	40	0	0	--

-- Diplodon_suavidicus **

Huang et al., 2019 [Publ 3843], fig. S4

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	199	0	0	--

-- Diplodon_suavidicus **

Graf & Ó Foighil, 2000 [Publ 2649], fig. 1_2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	10	0	0	--

-- Diplodon_chilensis **

Bolotov et al., 2016 [Publ 3743], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	29	0	0	--

-- Diplodon_suavidicus **

Wu et al., 2021 [Publ 4366], fig. 2_I

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	90	0	0	--

-- Diplodon_suavidicus **

Graf, 2000 [Publ 2647], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	16	0	0	--

-- Diplodon **

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