The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)
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Bivalvia | Unionida | Unionoidea | Unionidae | Gonideinae | Rectidentini | Contradentina genus Lens Simpson, 1900: cladomics
13 Recent valid species (13 total species) Cladomics Summarynumber of species = 13 (13 total)
analyzed in 71 cladograms in 46 publications
maximum number of terminals in an analysis =
11 (mean = 2.94, median = 1) Basic Monophylynumber of cladograms that test basic monophyly
(i.e., more than 1 included species) = 35
maximum number of included terminals = 11, (mean =
4.94 median = 4)
proportion of cladograms that support basic monophyly
= 0.97
Pairwise Cladogram ClusteringThe graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.
Jeratthitikul et al., 2019 [Publ 3911], fig. 4 ** Top Monophyly Cladogram ** iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 9 | 17 | 5 | 2 | 1 | 0 | +----------------------- Physunio
|
| +-- Lens_maenamensis **
--+ +-----------------+
| | +-- Lens_rolfbrandti **
| |
| | +----------------- Lens_contradens__12 **
+--+ |
| | +----- Lens_micropterus__3 **
| | |
| +-----------+ +-- Lens_contradens__10 **
| | | |
+--+ +--+-- Lens_micropterus__2 **
| |
| +-- Lens_contradens__11 **
|
| +-------------- Lens_contradens__9 **
| |
+--+ +----------- Lens_contradens__8 **
| |
+--+ +-------- Lens_contradens__7 **
| |
+--+ +----- Lens_contradens__6 **
| |
| | +-- Lens_contradens__1 **
+--+ |
| +-- Lens_contradens__2 **
| |
| +-- Lens_contradens__3 **
+--+
+-- Lens_contradens__4 **
|
+-- Lens_micropterus__1 **
|
+-- Lens_contradens__5 **
Pfeiffer et al., 2021 [Publ 4126], fig. 9 ** Top Sister-Group Cladogram ** iT| tT | bi | riS | oT | roT | roS | iD | | 8 | 8 | 6 | 8 | 21 | 7 | 4 | 15 | +-- Trapezoideus
+-----------------+
| +-- Physunio
|
--+ +----------------- Yaukthwa
| |
| | +-------------- Solenaia
+--+ |
| | +-------- Lens_misellus **
| | +--+
+--+ | | +----- Lens_inornatus **
| | +--+
| | | +-- Lens_contradens **
| | +--+
+--+ +-- Lens_micropterus **
|
| +-- Lens_maenamensis **
| +--+
| | +-- Lens_eximius **
+-----+
| +-- Lens_pallegoixi **
+--+
+-- Lens_comptus **
Zieritz et al., 2026 [Publ 4812], fig. 2a iT| tT | bi | riS | oT | roT | roS | iD | | 11 | 11 | 9 | 11 | 1 | 1 | 1 | 14 | +----------------- Solenaia
|
| +-- Lens_inornatus **
--+ +-----------+
| | +-- Lens_misellus **
| |
| | +----- Lens_contradens **
+--+ +-----+
| | | +-- Lens_lugens **
| | +--+
| | +-- Lens_micropterus **
+--+
| +----- Lens_pallegoixi **
| +--+
| | | +-- Lens_comptus **
| | +--+
+--+ +-- Lens_novoselovi **
|
| +----- Lens_maenamensis **
+--+
| +-- Lens_rolfbrandti **
+--+
+-- Lens_eximius **
Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14 iT| tT | bi | riS | oT | roT | roS | iD | | 11 | 11 | 10 | 12 | 620 | 20 | 3 | 14 | +-- Physunio_modelli
+--------------------------+
| +-- Physunio_pellucidus
|
--+ +-- Physunio_massini
| +-----------------------+
| | +-- Physunio_superbus
+--+
| +----------------------- Physunio
| |
+--+ +-------------------- Trapezoideus
| |
| | +----------------- Yaukthwa
+--+ |
| | +-- Lens_inornatus **
| | +-----+
| | | +-- Lens_misellus **
+--+ +-----+
| | | +----- Lens_contradens **
| | +--+
| | | +-- Lens_micropterus **
| | +--+
+--+ +-- Lens_lugens **
|
| +----- Lens_pallegoixi **
| +-----+
| | | +-- Lens_comptus **
| | +--+
+--+ +-- Lens_novoselovi **
|
| +-------- Lens_maenamensis **
+--+
| +----- Lens_eximius__2 **
+--+
| +-- Lens_eximius__1 **
+--+
+-- Lens_rolfbrandti **
Zieritz et al., 2024 [Publ 4381], fig. 7A iT| tT | bi | riS | oT | roT | roS | iD | | 11 | 11 | 9 | 11 | 1 | 1 | 1 | 14 | +----------------- Solenaia
|
| +----- Lens_contradens **
--+ +--------+
| | | +-- Lens_lugens **
| | +--+
| | +-- Lens_micropterus **
+--+
| +-- Lens_inornatus **
| +--------+
| | +-- Lens_misellus **
| |
+--+ +----- Lens_maenamensis **
| +--+
| | | +-- Lens_eximius **
| | +--+
+--+ +-- Lens_rolfbrandti **
|
| +----- Lens_pallegoixi **
+--+
| +-- Lens_comptus **
+--+
+-- Lens_novoselovi **
Pfeiffer et al., 2021 [Publ 4126], fig. 12 iT| tT | bi | riS | oT | roT | roS | iD | | 10 | 10 | 7 | 10 | 1 | 1 | 1 | 14 | +-------------- Solenaia
|
| +-- Lens_contradens **
| +--------+
--+ | +-- Lens_micropterus **
| |
| | +-- Lens_inornatus **
| +--------+
+--+ +-- Lens_misellus **
|
| +----- Lens_maenamensis **
| +--+
| | | +-- Lens_rolfbrandti **
| | +--+
+--+ +-- Lens_eximius **
|
| +----- Lens_pallegoixi **
+--+
| +-- Lens_comptus **
+--+
+-- Lens_novoselovi **
Konopleva et al., 2023 [Publ 4262], fig. 8 iT| tT | bi | riS | oT | roT | roS | iD | | 9 | 9 | 7 | 9 | 73 | 10 | 3 | 14 | +-- Physunio
+--------------+
| +-- Trapezoideus
|
--+ +-------------- Yaukthwa
| |
| | +----- Lens_misellus **
+--+ +-----+
| | | +-- Lens_contradens **
| | +--+
| | +-- Lens_micropterus **
+--+
| +----- Lens_pallegoixi **
| +--+
| | | +-- Lens_novoselovi **
| | +--+
+--+ +-- Lens_comptus **
|
| +----- Lens_maenamensis **
+--+
| +-- Lens_eximius **
+--+
+-- Lens_rolfbrandti **
Bolotov et al., 2020 [Publ 4100], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 9 | 9 | 7 | 9 | 108 | 14 | 3 | 14 | +-- Trapezoideus
+--------------+
| +-- Physunio
|
--+ +-------------- Yaukthwa
| |
| | +----- Lens_misellus **
+--+ +-----+
| | | +-- Lens_contradens **
| | +--+
| | +-- Lens_micropterus **
+--+
| +----- Lens_pallegoixi **
| +--+
| | | +-- Lens_comptus **
| | +--+
+--+ +-- Lens_novoselovi **
|
| +----- Lens_maenamensis **
+--+
| +-- Lens_eximius **
+--+
+-- Lens_rolfbrandti **
Konopleva et al., 2019 [Publ 4041], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 9 | 9 | 7 | 9 | 25 | 10 | 3 | 14 | +----------------- Yaukthwa
|
| +-- Trapezoideus
--+ +-----------+
| | +-- Physunio
| |
| | +----- Lens_misellus **
+--+ +-----+
| | | +-- Lens_contradens **
| | +--+
| | +-- Lens_micropterus **
+--+
| +----- Lens_pallegoixi **
| +--+
| | | +-- Lens_novoselovi **
| | +--+
+--+ +-- Lens_comptus **
|
| +----- Lens_maenamensis **
+--+
| +-- Lens_eximius **
+--+
+-- Lens_rolfbrandti **
Bolotov et al., 2020 [Publ 4100], fig. S1 iT| tT | bi | riS | oT | roT | roS | iD | | 9 | 9 | 7 | 9 | 102 | 14 | 3 | 14 | +-------------------- Physunio
|
--+ +----------------- Trapezoideus
| |
+--+ +-------------- Yaukthwa
| |
| | +----- Lens_misellus **
+--+ +-----+
| | | +-- Lens_contradens **
| | +--+
| | +-- Lens_micropterus **
+--+
| +----- Lens_maenamensis **
| +--+
| | | +-- Lens_eximius **
| | +--+
+--+ +-- Lens_rolfbrandti **
|
| +----- Lens_pallegoixi **
+--+
| +-- Lens_novoselovi **
+--+
+-- Lens_comptus **
Konopleva et al., 2019 [Publ 3900], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 7 | 5 | 7 | 44 | 9 | 3 | 16 | +----------------------- Yaukthwa
|
--+ +-------------------- Physunio
| |
+--+ +----------------- Trapezoideus
| |
+--+ +-- Lens_contradens **
| +-----------+
| | +-- Lens_micropterus **
+--+
| +----------- Lens_misellus **
+--+
| +-------- Lens_eximius **
+--+
| +----- Lens_pallegoixi **
+--+
| +-- Lens_comptus **
+--+
+-- Lens_novoselovi **
Zieritz et al., 2020 [Publ 4102], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 4 | 6 | 25 | 8 | 3 | 18 | +-------------------- Physunio
|
--+ +----------------- Yaukthwa
| |
+--+ +-------------- Trapezoideus
| |
+--+ +-- Lens_contradens **
| +--------+
| | +-- Lens_misellus **
+--+
| +-------- Lens_eximius **
+--+
| +----- Lens_pallegoixi **
+--+
| +-- Lens_comptus **
+--+
+-- Lens_novoselovi **
Bolotov et al., 2017 [Publ 3764], fig. S1 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 5 | 119 | 7 | 2 | 16 | +-------------- Physunio
|
--+ +----------- Yaukthwa
| |
+--+ +-- Lens_contradens **
| +-----+
| | +-- Lens_micropterus **
+--+
| +----- Lens_eximius **
+--+
| +-- Lens_comptus **
+--+
+-- Lens_novoselovi **
Huang et al., 2019 [Publ 3843], fig. S4 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 5 | 195 | 7 | 2 | 16 | +----------- Physunio_modelli
|
+----------- Physunio
|
+----------- Physunio_superbus
--+
+----------- Yaukthwa
|
| +-- Lens_comptus **
| +-----+
| | +-- Lens_novoselovi **
+--+
| +----- Lens_eximius **
+--+
| +-- Lens_contradens **
+--+
+-- Lens_micropterus **
Huang et al., 2019 [Publ 3843], fig. S3 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 5 | 193 | 7 | 2 | 16 | +-------------- Physunio
|
--+ +----------- Yaukthwa
| |
+--+ +-- Lens_contradens **
| +-----+
| | +-- Lens_micropterus **
+--+
| +----- Lens_eximius **
+--+
| +-- Lens_novoselovi **
+--+
+-- Lens_comptus **
Bolotov et al., 2017 [Publ 3761], fig. S1 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 5 | 133 | 7 | 2 | 16 | +-------------- Physunio
|
--+ +----------- Yaukthwa
| |
+--+ +-- Lens_contradens **
| +-----+
| | +-- Lens_micropterus **
+--+
| +----- Lens_eximius **
+--+
| +-- Lens_comptus **
+--+
+-- Lens_novoselovi **
Bolotov et al., 2018 [Publ 3826], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 10 | 0 | 5 | 77 | 5 | 2 | 16 | +-- Lens_eximius **
|
+-- Physunio_modelli__1
|
+-- Yaukthwa_nesemanni
|
+-- Yaukthwa_panhai
|
+-- Lens_comptus **
|
--+-- Lens_novoselovi **
|
+-- Physunio_modelli__2
|
+-- Physunio
|
+-- Yaukthwa_inlenensis
|
+-- Lens_contradens **
|
+-- Lens_micropterus **
Huang et al., 2019 [Publ 3843], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 5 | 195 | 7 | 2 | 16 | +-------------- Physunio
|
--+ +----------- Yaukthwa
| |
+--+ +-- Lens_contradens **
| +-----+
| | +-- Lens_micropterus **
+--+
| +----- Lens_eximius **
+--+
| +-- Lens_comptus **
+--+
+-- Lens_novoselovi **
Bolotov et al., 2017 [Publ 3761], fig. S2 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 134 | 7 | 2 | 17 | +-- Physunio
+--------+
| +-- Yaukthwa
--+
| +-------- Lens_contradens **
+--+
| +----- Lens_eximius **
+--+
| +-- Lens_comptus **
+--+
+-- Lens_novoselovi **
Jeratthitikul et al., 2021 [Publ 4212], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 17 | 6 | 3 | 15 | +-- Trapezoideus
+-----------+
| +-- Physunio
--+
| +----------- Yaukthwa
+--+
| +-------- Lens_contradens **
+--+
| +----- Lens_novoselovi **
+--+
| +-- Lens_rolfbrandti **
+--+
+-- Lens_maenamensis **
Zieritz et al., 2024 [Publ 4381], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 42 | 5 | 3 | 17 | +-- Physunio
+--------+
| +-- Trapezoideus
--+
| +-------- Yaukthwa
+--+
| +----- Lens_eximius **
+--+
| +-- Lens_contradens **
+--+
+-- Lens_lugens **
Dai et al., 2023 [Publ 4285], fig. 3-ML iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 43 | 1 | 1 | -- | +----- Physunio
--+
| +-- Lens_contradens **
+--+
+-- Lens_comptus **
Zieritz et al., 2021 [Publ 4218], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 29 | 5 | 3 | -- | +----------- Physunio
--+
| +-------- Trapezoideus
+--+
| +----- Yaukthwa
+--+
| +-- Lens_contradens **
+--+
+-- Lens_eximius **
Pfeiffer et al., 2018 [Publ 3827], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 5 | 2 | 2 | -- | +----- Physunio
|
--+----- Trapezoideus
|
| +-- Lens_comptus **
+--+
+-- Lens_contradens **
Dai et al., 2025 [Publ 4441], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 45 | 1 | 1 | -- | +----- Physunio
--+
| +-- Lens_comptus **
+--+
+-- Lens_contradens **
Dai et al., 2024 [Publ 4389], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 44 | 1 | 1 | -- | +----- Physunio
--+
| +-- Lens_comptus **
+--+
+-- Lens_contradens **
Chen et al., 2025 [Publ 4663], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 46 | 1 | 1 | -- | +----- Physunio
--+
| +-- Lens_comptus **
+--+
+-- Lens_contradens **
Liu et al., 2024 [Publ 4380], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 34 | 5 | 3 | -- | +-- Physunio
+-----+
| +-- Trapezoideus
--+
| +----- Yaukthwa
+--+
| +-- Lens_contradens **
+--+
+-- Lens_eximius **
Dai et al., 2023 [Publ 4285], fig. 3-BI iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 43 | 1 | 1 | -- | +----- Physunio
--+
| +-- Lens_contradens **
+--+
+-- Lens_comptus **
Wu et al., 2024 [Publ 4350], fig. 6-7 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 34 | 5 | 3 | -- | +-- Physunio
+-----+
| +-- Trapezoideus
--+
| +----- Yaukthwa
+--+
| +-- Lens_contradens **
+--+
+-- Lens_eximius **
Wu et al., 2024 [Publ 4350], fig. 5b iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 34 | 5 | 3 | -- | +-------------- Yaukthwa_elongatula
|
--+ +----------- Trapezoideus
| |
+--+ +-- Physunio
| +-----+
| | +-- Yaukthwa_inlenensis
+--+
| +----- Yaukthwa_paiensis
+--+
| +-- Lens_contradens **
+--+
+-- Lens_eximius **
Wu et al., 2024 [Publ 4350], fig. 5a iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 34 | 5 | 3 | -- | +-- Physunio
+-----+
| +-- Trapezoideus
--+
| +----- Yaukthwa
+--+
| +-- Lens_contradens **
+--+
+-- Lens_eximius **
Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 78 | 2 | 1 | -- | +----- Physunio
--+
| +-- Lens_contradens **
+--+
+-- Lens_comptus **
Wu et al., 2024 [Publ 4388], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 34 | 5 | 3 | -- | +-- Physunio
+-----+
| +-- Trapezoideus
--+
| +----- Yaukthwa
+--+
| +-- Lens_contradens **
+--+
+-- Lens_eximius **
Wu et al., 2024 [Publ 4350], fig. 5c iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 34 | 5 | 3 | -- | +----- Physunio
|
+----- Yaukthwa_paiensis
|
+----- Yaukthwa_inlenensis
--+
+----- Trapezoideus
|
+----- Yaukthwa_elongatula
|
| +-- Lens_contradens **
+--+
+-- Lens_eximius **
Jeratthitikul et al., 2021 [Publ 4219], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 34 | 2 | 2 | -- | +----- Lens_contradens **
--+
| +-- Trapezoideus
+--+
+-- Physunio
Zieritz et al., 2021 [Publ 4231], fig. 3-M_4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 22 | 1 | 1 | -- | +-- Lens_contradens **
--+
+-- Physunio
Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 75 | 2 | 2 | -- | +----- Lens_contradens **
--+
| +-- Trapezoideus
+--+
+-- Physunio
Dai et al., 2025 [Publ 4440], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 87 | 1 | 1 | -- | +-- Lens_contradens **
--+
+-- Physunio
Whelan et al., 2011 [Publ 3380], fig. 1-ML iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 35 | 0 | 0 | -- | Pfeiffer & Graf, 2015 [Publ 3620], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 39 | 2 | 1 | -- | +-- Lens_micropterus **
--+
+-- Physunio
Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 140 | 1 | 1 | -- | +-- Lens_contradens **
--+
+-- Physunio
Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 75 | 2 | 2 | -- | +----- Lens_contradens **
--+
| +-- Trapezoideus
+--+
+-- Physunio
Smith, 2021 [Publ 4331], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 19 | 0 | 0 | -- | Campbell & Lydeard, 2026 [Publ 4810], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 46 | 1 | 1 | -- | +-- Lens_contradens **
--+
+-- Physunio
Wu et al., 2023 [Publ 4339], fig. 4_S5-aa iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 95 | 1 | 1 | -- | +-- Lens_contradens **
--+
+-- Physunio
Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 121 | 2 | 2 | -- | Hoeh et al., 2009 [Publ 2716], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 45 | 0 | 0 | -- | Hoeh et al., 2009 [Publ 2716], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 46 | 0 | 0 | -- | Wu et al., 2023 [Publ 4339], fig. 4_S5-nt iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 95 | 1 | 1 | -- | +-- Lens_contradens **
--+
+-- Physunio
Graf, 2002 [Publ 2651], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 17 | 0 | 0 | -- | Chen et al., 2025 [Publ 4448], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 85 | 1 | 1 | -- | +-- Lens_contradens **
--+
+-- Physunio
Wu et al., 2024 [Publ 4350], fig. 8a_S1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 91 | 1 | 1 | -- | +-- Lens_contradens **
--+
+-- Physunio
Matos et al., 2024 [Publ 4384], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 19 | 1 | 1 | -- | +-- Physunio
--+
+-- Lens_contradens **
Pfeiffer et al., 2019 [Publ 3937], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 56 | 4 | 4 | -- | +-- Trapezoideus
+-----+
| +-- Physunio
--+
| +----- Yaukthwa
+--+
| +-- Lens_contradens **
+--+
+-- Solenaia
Dai et al., 2023 [Publ 4285], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 52 | 1 | 1 | -- | +-- Lens_contradens **
--+
+-- Physunio
Zhang et al., 2024 [Publ 4397], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 26 | 1 | 1 | -- | +-- Lens_contradens **
--+
+-- Physunio
Hoeh et al., 2009 [Publ 2716], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 45 | 0 | 0 | -- | Wu et al., 2021 [Publ 4366], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 90 | 3 | 2 | -- | +----- Physunio
--+
| +-- Lens_eximius **
+--+
+-- Yaukthwa
Konopleva et al., 2017 [Publ 3807], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 30 | 2 | 1 | -- | +-- Lens_micropterus **
--+
+-- Physunio
Jeratthitikul et al., 2024 [Publ 4387], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 58 | 2 | 2 | -- | +----- Physunio
--+
| +-- Lens_contradens **
+--+
+-- Trapezoideus
Wu et al., 2024 [Publ 4350], fig. 8b_S2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 91 | 1 | 1 | -- | +-- Lens_contradens **
--+
+-- Physunio
Matos et al., 2024 [Publ 4383], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 19 | 1 | 1 | -- | +-- Physunio
--+
+-- Lens_contradens **
Pfeiffer et al., 2019 [Publ 3937], fig. 4-5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 56 | 4 | 4 | -- | +-- Trapezoideus
+-----+
| +-- Physunio
--+
| +----- Yaukthwa
+--+
| +-- Lens_contradens **
+--+
+-- Solenaia
Zieritz et al., 2021 [Publ 4231], fig. 3-F_5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 47 | 1 | 1 | -- | +-- Lens_contradens **
--+
+-- Physunio
Wu et al., 2024 [Publ 4388], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 52 | 1 | 1 | -- | +-- Lens_contradens **
--+
+-- Physunio
Graf & Cummings, 2006 [Publ 1968], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 46 | 0 | 0 | -- | Wu et al., 2021 [Publ 4366], fig. 2_I iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 90 | 3 | 2 | -- | +----- Physunio
--+
| +-- Lens_eximius **
+--+
+-- Yaukthwa
Wu et al., 2026 [Publ 4811], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 94 | 1 | 1 | -- | +-- Lens_contradens **
--+
+-- Physunio
Whelan et al., 2011 [Publ 3380], fig. 1-MP iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 35 | 0 | 0 | -- | Jeratthitikul et al., 2021 [Publ 4219], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 33 | 2 | 2 | -- | +----- Lens_contradens **
--+
| +-- Trapezoideus
+--+
+-- Physunio
|
