MUSSELpdb | valid gen. Lens cladomics

	Home \| About \| Database \| Mussel of the Month \| Publications & Presentations


MUSSELpdb
	Introduction Browse d'basing


Taxonomy
	all genera Unionidae Margaritiferidae Hyriidae Etheriidae Mycetopodidae Iridinidae Species & Genus Richness by Taxonomy


Geography
	Summary North America Central America South America Afrotropics North Eurasia East Asia Australasia Species Richness & Endemism by Geography

Page last updated
Thu 3 September 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Search the database pages:

genus Lens Simpson, 1900: cladomics
13 Recent valid species (13 total species)

Cladomics Summary

number of species = 13 (13 total)
analyzed in 73 cladograms in 47 publications
maximum number of terminals in an analysis = 11 (mean = 2.89, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 35
maximum number of included terminals = 11, (mean = 4.94 median = 4)
proportion of cladograms that support basic monophyly = 0.97

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination

NJ Tree

Jeratthitikul et al., 2019 [Publ 3911], fig. 4

** Top Monophyly Cladogram **

tT	bi	riS	oT	roT	roS	iD
4	4	9	17	5	2	1	0

  +----------------------- Physunio           
  |                                           
  |                    +-- Lens_maenamensis **   
--+  +-----------------+                      
  |  |                 +-- Lens_rolfbrandti **   
  |  |                                        
  |  |  +----------------- Lens_contradens__12 **
  +--+  |                                     
     |  |           +----- Lens_micropterus__3 **
     |  |           |                         
     |  +-----------+  +-- Lens_contradens__10 **
     |  |           |  |                      
     +--+           +--+-- Lens_micropterus__2 **
        |              |                      
        |              +-- Lens_contradens__11 **
        |                                     
        |  +-------------- Lens_contradens__9 ** 
        |  |                                  
        +--+  +----------- Lens_contradens__8 ** 
           |  |                               
           +--+  +-------- Lens_contradens__7 ** 
              |  |                            
              +--+  +----- Lens_contradens__6 ** 
                 |  |                         
                 |  |  +-- Lens_contradens__1 ** 
                 +--+  |                      
                    |  +-- Lens_contradens__2 ** 
                    |  |                      
                    |  +-- Lens_contradens__3 ** 
                    +--+                      
                       +-- Lens_contradens__4 ** 
                       |                      
                       +-- Lens_micropterus__1 **
                       |                      
                       +-- Lens_contradens__5 **

Pfeiffer et al., 2021 [Publ 4126], fig. 9

** Top Sister-Group Cladogram **

tT	bi	riS	oT	roT	roS	iD
8	8	6	8	21	7	4	15

                    +-- Trapezoideus    
  +-----------------+                   
  |                 +-- Physunio        
  |                                     
--+  +----------------- Yaukthwa        
  |  |                                  
  |  |  +-------------- Solenaia        
  +--+  |                               
     |  |     +-------- Lens_misellus **   
     |  |  +--+                         
     +--+  |  |  +----- Lens_inornatus **  
        |  |  +--+                      
        |  |     |  +-- Lens_contradens ** 
        |  |     +--+                   
        +--+        +-- Lens_micropterus **
           |                            
           |        +-- Lens_maenamensis **
           |     +--+                   
           |     |  +-- Lens_eximius **    
           +-----+                      
                 |  +-- Lens_pallegoixi ** 
                 +--+                   
                    +-- Lens_comptus **

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

tT	bi	riS	oT	roT	roS	iD
11	11	10	12	621	20	3	14

                             +-- Physunio_modelli   
  +--------------------------+                      
  |                          +-- Physunio_pellucidus
  |                                                 
--+                          +-- Physunio_massini   
  |  +-----------------------+                      
  |  |                       +-- Physunio_superbus  
  +--+                                              
     |  +----------------------- Physunio           
     |  |                                           
     +--+  +-------------------- Trapezoideus       
        |  |                                        
        |  |  +----------------- Yaukthwa           
        +--+  |                                     
           |  |              +-- Lens_inornatus **     
           |  |        +-----+                      
           |  |        |     +-- Lens_misellus **      
           +--+  +-----+                            
              |  |     |  +----- Lens_contradens **    
              |  |     +--+                         
              |  |        |  +-- Lens_micropterus **   
              |  |        +--+                      
              +--+           +-- Lens_lugens **        
                 |                                  
                 |        +----- Lens_pallegoixi **    
                 |  +-----+                         
                 |  |     |  +-- Lens_comptus **       
                 |  |     +--+                      
                 +--+        +-- Lens_novoselovi **    
                    |                               
                    |  +-------- Lens_maenamensis **   
                    +--+                            
                       |  +----- Lens_eximius__2 **    
                       +--+                         
                          |  +-- Lens_eximius__1 **    
                          +--+                      
                             +-- Lens_rolfbrandti **

Zieritz et al., 2026 [Publ 4812], fig. 2a

tT	bi	riS	oT	roT	roS	iD
11	11	9	11	1	1	1	14

  +----------------- Solenaia        
  |                                  
  |              +-- Lens_inornatus **  
--+  +-----------+                   
  |  |           +-- Lens_misellus **   
  |  |                               
  |  |        +----- Lens_contradens ** 
  +--+  +-----+                      
     |  |     |  +-- Lens_lugens **     
     |  |     +--+                   
     |  |        +-- Lens_micropterus **
     +--+                            
        |     +----- Lens_pallegoixi ** 
        |  +--+                      
        |  |  |  +-- Lens_comptus **    
        |  |  +--+                   
        +--+     +-- Lens_novoselovi ** 
           |                         
           |  +----- Lens_maenamensis **
           +--+                      
              |  +-- Lens_rolfbrandti **
              +--+                   
                 +-- Lens_eximius **

Zieritz et al., 2024 [Publ 4381], fig. 7A

tT	bi	riS	oT	roT	roS	iD
11	11	9	11	1	1	1	14

  +----------------- Solenaia        
  |                                  
  |           +----- Lens_contradens ** 
--+  +--------+                      
  |  |        |  +-- Lens_lugens **     
  |  |        +--+                   
  |  |           +-- Lens_micropterus **
  +--+                               
     |           +-- Lens_inornatus **  
     |  +--------+                   
     |  |        +-- Lens_misellus **   
     |  |                            
     +--+     +----- Lens_maenamensis **
        |  +--+                      
        |  |  |  +-- Lens_eximius **    
        |  |  +--+                   
        +--+     +-- Lens_rolfbrandti **
           |                         
           |  +----- Lens_pallegoixi ** 
           +--+                      
              |  +-- Lens_comptus **    
              +--+                   
                 +-- Lens_novoselovi **

Pfeiffer et al., 2021 [Publ 4126], fig. 12

tT	bi	riS	oT	roT	roS	iD
10	10	7	10	1	1	1	14

  +-------------- Solenaia        
  |                               
  |           +-- Lens_contradens ** 
  |  +--------+                   
--+  |        +-- Lens_micropterus **
  |  |                            
  |  |        +-- Lens_inornatus **  
  |  +--------+                   
  +--+        +-- Lens_misellus **   
     |                            
     |     +----- Lens_maenamensis **
     |  +--+                      
     |  |  |  +-- Lens_rolfbrandti **
     |  |  +--+                   
     +--+     +-- Lens_eximius **    
        |                         
        |  +----- Lens_pallegoixi ** 
        +--+                      
           |  +-- Lens_comptus **    
           +--+                   
              +-- Lens_novoselovi **

Konopleva et al., 2023 [Publ 4262], fig. 8

tT	bi	riS	oT	roT	roS	iD
9	9	7	9	73	10	3	14

                 +-- Physunio        
  +--------------+                   
  |              +-- Trapezoideus    
  |                                  
--+  +-------------- Yaukthwa        
  |  |                               
  |  |        +----- Lens_misellus **   
  +--+  +-----+                      
     |  |     |  +-- Lens_contradens ** 
     |  |     +--+                   
     |  |        +-- Lens_micropterus **
     +--+                            
        |     +----- Lens_pallegoixi ** 
        |  +--+                      
        |  |  |  +-- Lens_novoselovi ** 
        |  |  +--+                   
        +--+     +-- Lens_comptus **    
           |                         
           |  +----- Lens_maenamensis **
           +--+                      
              |  +-- Lens_eximius **    
              +--+                   
                 +-- Lens_rolfbrandti **

Bolotov et al., 2020 [Publ 4100], fig. S1

tT	bi	riS	oT	roT	roS	iD
9	9	7	9	102	14	3	14

  +-------------------- Physunio        
  |                                     
--+  +----------------- Trapezoideus    
  |  |                                  
  +--+  +-------------- Yaukthwa        
     |  |                               
     |  |        +----- Lens_misellus **   
     +--+  +-----+                      
        |  |     |  +-- Lens_contradens ** 
        |  |     +--+                   
        |  |        +-- Lens_micropterus **
        +--+                            
           |     +----- Lens_maenamensis **
           |  +--+                      
           |  |  |  +-- Lens_eximius **    
           |  |  +--+                   
           +--+     +-- Lens_rolfbrandti **
              |                         
              |  +----- Lens_pallegoixi ** 
              +--+                      
                 |  +-- Lens_novoselovi ** 
                 +--+                   
                    +-- Lens_comptus **

Konopleva et al., 2019 [Publ 4041], fig. 3

tT	bi	riS	oT	roT	roS	iD
9	9	7	9	25	10	3	14

  +----------------- Yaukthwa        
  |                                  
  |              +-- Trapezoideus    
--+  +-----------+                   
  |  |           +-- Physunio        
  |  |                               
  |  |        +----- Lens_misellus **   
  +--+  +-----+                      
     |  |     |  +-- Lens_contradens ** 
     |  |     +--+                   
     |  |        +-- Lens_micropterus **
     +--+                            
        |     +----- Lens_pallegoixi ** 
        |  +--+                      
        |  |  |  +-- Lens_novoselovi ** 
        |  |  +--+                   
        +--+     +-- Lens_comptus **    
           |                         
           |  +----- Lens_maenamensis **
           +--+                      
              |  +-- Lens_eximius **    
              +--+                   
                 +-- Lens_rolfbrandti **

Bolotov et al., 2020 [Publ 4100], fig. 1

tT	bi	riS	oT	roT	roS	iD
9	9	7	9	108	14	3	14

                 +-- Trapezoideus    
  +--------------+                   
  |              +-- Physunio        
  |                                  
--+  +-------------- Yaukthwa        
  |  |                               
  |  |        +----- Lens_misellus **   
  +--+  +-----+                      
     |  |     |  +-- Lens_contradens ** 
     |  |     +--+                   
     |  |        +-- Lens_micropterus **
     +--+                            
        |     +----- Lens_pallegoixi ** 
        |  +--+                      
        |  |  |  +-- Lens_comptus **    
        |  |  +--+                   
        +--+     +-- Lens_novoselovi ** 
           |                         
           |  +----- Lens_maenamensis **
           +--+                      
              |  +-- Lens_eximius **    
              +--+                   
                 +-- Lens_rolfbrandti **

Konopleva et al., 2019 [Publ 3900], fig. 1

tT	bi	riS	oT	roT	roS	iD
7	7	5	7	44	9	3	16

  +----------------------- Yaukthwa        
  |                                        
--+  +-------------------- Physunio        
  |  |                                     
  +--+  +----------------- Trapezoideus    
     |  |                                  
     +--+              +-- Lens_contradens ** 
        |  +-----------+                   
        |  |           +-- Lens_micropterus **
        +--+                               
           |  +----------- Lens_misellus **   
           +--+                            
              |  +-------- Lens_eximius **    
              +--+                         
                 |  +----- Lens_pallegoixi ** 
                 +--+                      
                    |  +-- Lens_comptus **    
                    +--+                   
                       +-- Lens_novoselovi **

Zieritz et al., 2020 [Publ 4102], fig. 4

tT	bi	riS	oT	roT	roS	iD
6	6	4	6	25	8	3	18

  +-------------------- Physunio       
  |                                    
--+  +----------------- Yaukthwa       
  |  |                                 
  +--+  +-------------- Trapezoideus   
     |  |                              
     +--+           +-- Lens_contradens **
        |  +--------+                  
        |  |        +-- Lens_misellus **  
        +--+                           
           |  +-------- Lens_eximius **   
           +--+                        
              |  +----- Lens_pallegoixi **
              +--+                     
                 |  +-- Lens_comptus **   
                 +--+                  
                    +-- Lens_novoselovi **

Huang et al., 2019 [Publ 3843], fig. S4

tT	bi	riS	oT	roT	roS	iD
5	5	3	5	195	7	2	16

  +----------- Physunio_modelli 
  |                             
  +----------- Physunio         
  |                             
  +----------- Physunio_superbus
--+                             
  +----------- Yaukthwa         
  |                             
  |        +-- Lens_comptus **     
  |  +-----+                    
  |  |     +-- Lens_novoselovi **  
  +--+                          
     |  +----- Lens_eximius **     
     +--+                       
        |  +-- Lens_contradens **  
        +--+                    
           +-- Lens_micropterus **

Bolotov et al., 2018 [Publ 3826], fig. 2

tT	bi	riS	oT	roT	roS	iD
5	10	0	5	77	5	2	16

  +-- Lens_eximius **       
  |                      
  +-- Physunio_modelli__1
  |                      
  +-- Yaukthwa_nesemanni 
  |                      
  +-- Yaukthwa_panhai    
  |                      
  +-- Lens_comptus **       
  |                      
--+-- Lens_novoselovi **    
  |                      
  +-- Physunio_modelli__2
  |                      
  +-- Physunio           
  |                      
  +-- Yaukthwa_inlenensis
  |                      
  +-- Lens_contradens **    
  |                      
  +-- Lens_micropterus **

Bolotov et al., 2017 [Publ 3761], fig. S1

tT	bi	riS	oT	roT	roS	iD
5	5	3	5	133	7	2	16

  +-------------- Physunio        
  |                               
--+  +----------- Yaukthwa        
  |  |                            
  +--+        +-- Lens_contradens ** 
     |  +-----+                   
     |  |     +-- Lens_micropterus **
     +--+                         
        |  +----- Lens_eximius **    
        +--+                      
           |  +-- Lens_comptus **    
           +--+                   
              +-- Lens_novoselovi **

Huang et al., 2019 [Publ 3843], fig. 2

tT	bi	riS	oT	roT	roS	iD
5	5	3	5	195	7	2	16

  +-------------- Physunio        
  |                               
--+  +----------- Yaukthwa        
  |  |                            
  +--+        +-- Lens_contradens ** 
     |  +-----+                   
     |  |     +-- Lens_micropterus **
     +--+                         
        |  +----- Lens_eximius **    
        +--+                      
           |  +-- Lens_comptus **    
           +--+                   
              +-- Lens_novoselovi **

Huang et al., 2019 [Publ 3843], fig. S3

tT	bi	riS	oT	roT	roS	iD
5	5	3	5	193	7	2	16

  +-------------- Physunio        
  |                               
--+  +----------- Yaukthwa        
  |  |                            
  +--+        +-- Lens_contradens ** 
     |  +-----+                   
     |  |     +-- Lens_micropterus **
     +--+                         
        |  +----- Lens_eximius **    
        +--+                      
           |  +-- Lens_novoselovi ** 
           +--+                   
              +-- Lens_comptus **

Bolotov et al., 2017 [Publ 3764], fig. S1

tT	bi	riS	oT	roT	roS	iD
5	5	3	5	119	7	2	16

  +-------------- Physunio        
  |                               
--+  +----------- Yaukthwa        
  |  |                            
  +--+        +-- Lens_contradens ** 
     |  +-----+                   
     |  |     +-- Lens_micropterus **
     +--+                         
        |  +----- Lens_eximius **    
        +--+                      
           |  +-- Lens_comptus **    
           +--+                   
              +-- Lens_novoselovi **

Bolotov et al., 2017 [Publ 3761], fig. S2

tT	bi	riS	oT	roT	roS	iD
4	4	2	4	134	7	2	17

           +-- Physunio       
  +--------+                  
  |        +-- Yaukthwa       
--+                           
  |  +-------- Lens_contradens **
  +--+                        
     |  +----- Lens_eximius **   
     +--+                     
        |  +-- Lens_comptus **   
        +--+                  
           +-- Lens_novoselovi **

Jeratthitikul et al., 2021 [Publ 4212], fig. 4

tT	bi	riS	oT	roT	roS	iD
4	4	2	4	17	6	3	15

              +-- Trapezoideus    
  +-----------+                   
  |           +-- Physunio        
--+                               
  |  +----------- Yaukthwa        
  +--+                            
     |  +-------- Lens_contradens ** 
     +--+                         
        |  +----- Lens_novoselovi ** 
        +--+                      
           |  +-- Lens_rolfbrandti **
           +--+                   
              +-- Lens_maenamensis **

Zieritz et al., 2024 [Publ 4381], fig. 2

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	42	5	3	17

           +-- Physunio       
  +--------+                  
  |        +-- Trapezoideus   
--+                           
  |  +-------- Yaukthwa       
  +--+                        
     |  +----- Lens_eximius **   
     +--+                     
        |  +-- Lens_contradens **
        +--+                  
           +-- Lens_lugens **

Wu et al., 2024 [Publ 4388], fig. 5

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	34	5	3	--

        +-- Physunio       
  +-----+                  
  |     +-- Trapezoideus   
--+                        
  |  +----- Yaukthwa       
  +--+                     
     |  +-- Lens_contradens **
     +--+                  
        +-- Lens_eximius **

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	78	2	1	--

  +----- Physunio       
--+                     
  |  +-- Lens_contradens **
  +--+                  
     +-- Lens_comptus **

Wu et al., 2024 [Publ 4350], fig. 6-7

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	34	5	3	--

        +-- Physunio       
  +-----+                  
  |     +-- Trapezoideus   
--+                        
  |  +----- Yaukthwa       
  +--+                     
     |  +-- Lens_contradens **
     +--+                  
        +-- Lens_eximius **

Wu et al., 2024 [Publ 4350], fig. 5b

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	34	5	3	--

  +-------------- Yaukthwa_elongatula
  |                                  
--+  +----------- Trapezoideus       
  |  |                               
  +--+        +-- Physunio           
     |  +-----+                      
     |  |     +-- Yaukthwa_inlenensis
     +--+                            
        |  +----- Yaukthwa_paiensis  
        +--+                         
           |  +-- Lens_contradens **    
           +--+                      
              +-- Lens_eximius **

Zieritz et al., 2021 [Publ 4218], fig. 5

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	29	5	3	--

  +----------- Physunio       
--+                           
  |  +-------- Trapezoideus   
  +--+                        
     |  +----- Yaukthwa       
     +--+                     
        |  +-- Lens_contradens **
        +--+                  
           +-- Lens_eximius **

Dai et al., 2024 [Publ 4389], fig. 3

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	44	1	1	--

  +----- Physunio       
--+                     
  |  +-- Lens_comptus **   
  +--+                  
     +-- Lens_contradens **

Dai et al., 2023 [Publ 4285], fig. 3-ML

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	43	1	1	--

  +----- Physunio       
--+                     
  |  +-- Lens_contradens **
  +--+                  
     +-- Lens_comptus **

Dai et al., 2023 [Publ 4285], fig. 3-BI

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	43	1	1	--

  +----- Physunio       
--+                     
  |  +-- Lens_contradens **
  +--+                  
     +-- Lens_comptus **

Liu et al., 2024 [Publ 4380], fig. 5

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	34	5	3	--

        +-- Physunio       
  +-----+                  
  |     +-- Trapezoideus   
--+                        
  |  +----- Yaukthwa       
  +--+                     
     |  +-- Lens_contradens **
     +--+                  
        +-- Lens_eximius **

Wu et al., 2024 [Publ 4350], fig. 5a

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	34	5	3	--

        +-- Physunio       
  +-----+                  
  |     +-- Trapezoideus   
--+                        
  |  +----- Yaukthwa       
  +--+                     
     |  +-- Lens_contradens **
     +--+                  
        +-- Lens_eximius **

Pfeiffer et al., 2018 [Publ 3827], fig. 1

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	5	2	2	--

  +----- Physunio       
  |                     
--+----- Trapezoideus   
  |                     
  |  +-- Lens_comptus **   
  +--+                  
     +-- Lens_contradens **

Dai et al., 2025 [Publ 4441], fig. 1

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	45	1	1	--

  +----- Physunio       
--+                     
  |  +-- Lens_comptus **   
  +--+                  
     +-- Lens_contradens **

Chen et al., 2025 [Publ 4663], fig. 1

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	46	1	1	--

  +----- Physunio       
--+                     
  |  +-- Lens_comptus **   
  +--+                  
     +-- Lens_contradens **

Wu et al., 2024 [Publ 4350], fig. 5c

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	34	5	3	--

  +----- Physunio           
  |                         
  +----- Yaukthwa_paiensis  
  |                         
  +----- Yaukthwa_inlenensis
--+                         
  +----- Trapezoideus       
  |                         
  +----- Yaukthwa_elongatula
  |                         
  |  +-- Lens_contradens **    
  +--+                      
     +-- Lens_eximius **

Matos et al., 2024 [Publ 4383], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	19	1	1	--

  +-- Physunio       
--+                  
  +-- Lens_contradens **

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	95	1	1	--

  +-- Lens_contradens **
--+                  
  +-- Physunio

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	39	2	1	--

  +-- Lens_micropterus **
--+                   
  +-- Physunio

Wu et al., 2021 [Publ 4366], fig. 2_I

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	90	3	2	--

  +----- Physunio    
--+                  
  |  +-- Lens_eximius **
  +--+               
     +-- Yaukthwa

Konopleva et al., 2017 [Publ 3807], fig. 6

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	30	2	1	--

  +-- Lens_micropterus **
--+                   
  +-- Physunio

Wu et al., 2026 [Publ 4811], fig. 4

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	95	1	1	--

  +-- Lens_contradens **
--+                  
  +-- Physunio

Wu et al., 2021 [Publ 4366], fig. 6

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	90	3	2	--

  +----- Physunio    
--+                  
  |  +-- Lens_eximius **
  +--+               
     +-- Yaukthwa

Wu et al., 2024 [Publ 4388], fig. 6

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	52	1	1	--

  +-- Lens_contradens **
--+                  
  +-- Physunio

Dai et al., 2025 [Publ 4440], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	87	1	1	--

  +-- Lens_contradens **
--+                  
  +-- Physunio

Jeratthitikul et al., 2024 [Publ 4387], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	58	2	2	--

  +----- Physunio       
--+                     
  |  +-- Lens_contradens **
  +--+                  
     +-- Trapezoideus

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	75	2	2	--

  +----- Lens_contradens **
--+                     
  |  +-- Trapezoideus   
  +--+                  
     +-- Physunio

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	47	1	1	--

  +-- Lens_contradens **
--+                  
  +-- Physunio

Graf, 2002 [Publ 2651], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	17	0	0	--

-- Lens_contradens **

Pfeiffer et al., 2019 [Publ 3937], fig. 6

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	56	4	4	--

        +-- Trapezoideus   
  +-----+                  
  |     +-- Physunio       
--+                        
  |  +----- Yaukthwa       
  +--+                     
     |  +-- Lens_contradens **
     +--+                  
        +-- Solenaia

Whelan et al., 2011 [Publ 3380], fig. 1-ML

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	35	0	0	--

-- Lens_contradens **

Klymus et al., 2025 [Publ 4814], fig. 3B

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	35	1	1	--

  +-- Lens_contradens **
--+                  
  +-- Physunio

Wu et al., 2024 [Publ 4350], fig. 8a_S1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	91	1	1	--

  +-- Lens_contradens **
--+                  
  +-- Physunio

Jeratthitikul et al., 2021 [Publ 4219], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	33	2	2	--

  +----- Lens_contradens **
--+                     
  |  +-- Trapezoideus   
  +--+                  
     +-- Physunio

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	95	1	1	--

  +-- Lens_contradens **
--+                  
  +-- Physunio

Whelan et al., 2011 [Publ 3380], fig. 1-MP

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	35	0	0	--

-- Lens_contradens **

Campbell & Lydeard, 2026 [Publ 4810], fig. 4

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	46	1	1	--

  +-- Lens_contradens **
--+                  
  +-- Physunio

Hoeh et al., 2009 [Publ 2716], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	46	0	0	--

-- Lens_contradens **

Graf & Cummings, 2006 [Publ 1968], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	46	0	0	--

-- Lens_contradens **

Chen et al., 2025 [Publ 4448], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	85	1	1	--

  +-- Lens_contradens **
--+                  
  +-- Physunio

Hoeh et al., 2009 [Publ 2716], fig. 5

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	45	0	0	--

-- Lens_contradens **

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	56	4	4	--

        +-- Trapezoideus   
  +-----+                  
  |     +-- Physunio       
--+                        
  |  +----- Yaukthwa       
  +--+                     
     |  +-- Lens_contradens **
     +--+                  
        +-- Solenaia

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	121	2	2	--

-- Lens_contradens **

Matos et al., 2024 [Publ 4384], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	19	1	1	--

  +-- Physunio       
--+                  
  +-- Lens_contradens **

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	140	1	1	--

  +-- Lens_contradens **
--+                  
  +-- Physunio

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	75	2	2	--

  +----- Lens_contradens **
--+                     
  |  +-- Trapezoideus   
  +--+                  
     +-- Physunio

Zieritz et al., 2021 [Publ 4231], fig. 3-M_4

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	22	1	1	--

  +-- Lens_contradens **
--+                  
  +-- Physunio

Klymus et al., 2025 [Publ 4814], fig. 3A

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	35	1	1	--

  +-- Lens_contradens **
--+                  
  +-- Physunio

Jeratthitikul et al., 2021 [Publ 4219], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	34	2	2	--

  +----- Lens_contradens **
--+                     
  |  +-- Trapezoideus   
  +--+                  
     +-- Physunio

Dai et al., 2023 [Publ 4285], fig. 5

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	52	1	1	--

  +-- Lens_contradens **
--+                  
  +-- Physunio

Smith, 2021 [Publ 4331], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	19	0	0	--

-- Lens_contradens **

Zhang et al., 2024 [Publ 4397], fig. 5

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	26	1	1	--

  +-- Lens_contradens **
--+                  
  +-- Physunio

Hoeh et al., 2009 [Publ 2716], fig. 4

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	45	0	0	--

-- Lens_contradens **

Wu et al., 2024 [Publ 4350], fig. 8b_S2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	91	1	1	--

  +-- Lens_contradens **
--+                  
  +-- Physunio

"Making the world a better place, one mollusk at a time."
	The MUSSEL Project — Home Page https://musselpdb.org/. Site developed and maintained by Dan Graf & Kevin Cummings. Funded by The National Science Foundation.