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Page last updated
Thu 17 July 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Bivalvia | Unionoida | Unionoidea | Unionidae | Gonideinae | Rectidentini

genus Rectidens Simpson, 1900: cladomics
     2 Recent valid species (2 total species)

Cladomics Summary

number of species = 2 (2 total)
analyzed in 48 cladograms in 30 publications
maximum number of terminals in an analysis = 2 (mean = 1.04, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 2
maximum number of included terminals = 2, (mean = 2 median = 2)
proportion of cladograms that support basic monophyly = 1

Zieritz et al., 2024 [Publ 4381], fig. 2

** Top Monophyly Cladogram **

iTtTbiriSoTroTroSiD
22024432--
  +-------- Ensidens             
--+                              
  |  +----- Hyriopsis            
  +--+                           
     |  +-- Rectidens_sumatrensis **
     +--+                        
        +-- Rectidens_lauris **

Konopleva et al., 2023 [Publ 4262], fig. 8

** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
110181142--
  +----------------------- Ensidens               
  |                                               
--+                    +-- Hyriopsis_desowitzi    
  |  +-----------------+                          
  |  |                 +-- Hyriopsis_delaportei   
  +--+                                            
     |  +----------------- Rectidens_sumatrensis **  
     |  |                                         
     +--+  +-------------- Hyriopsis_panhai       
        |  |                                      
        +--+  +----------- Hyriopsis_myersiana    
           |  |                                   
           +--+        +-- Hyriopsis_khoratensis  
              |  +-----+                          
              |  |     +-- Hyriopsis_bialata      
              +--+                                
                 |  +----- Hyriopsis_sakhonensis  
                 +--+                             
                    |  +-- Hyriopsis_phuphaniensis
                    +--+                          
                       +-- Hyriopsis_kratiensis

Zieritz et al., 2024 [Publ 4381], fig. 6A

iTtTbiriSoTroTroSiD
2202111--
  +----- Ensidens             
--+                           
  |  +-- Rectidens_sumatrensis **
  +--+                        
     +-- Rectidens_lauris **

Zieritz et al., 2021 [Publ 4218], fig. 5

iTtTbiriSoTroTroSiD
11013132--
  +----- Ensidens             
--+                           
  |  +-- Rectidens_sumatrensis **
  +--+                        
     +-- Hyriopsis

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
1101198102--
  +----- Hyriopsis_desowitzi  
  |                           
  +----- Hyriopsis_myersiana  
  |                           
  +----- Hyriopsis_panhai     
--+                           
  |  +-- Hyriopsis_bialata    
  +--+                        
  |  +-- Hyriopsis_khoratensis
  |                           
  |  +-- Rectidens_sumatrensis **
  +--+                        
     +-- Ensidens

Zieritz et al., 2021 [Publ 4231], fig. 3-M_4

iTtTbiriSoTroTroSiD
11012211--
  +-- Hyriopsis            
--+                        
  +-- Rectidens_sumatrensis **

Bolotov et al., 2020 [Publ 4100], fig. S1

iTtTbiriSoTroTroSiD
1101111112--
  +----------------- Ensidens             
--+                                       
  |  +-------------- Hyriopsis_desowitzi  
  +--+                                    
     |  +----------- Rectidens_sumatrensis **
     +--+                                 
        |  +-------- Hyriopsis_panhai     
        +--+                              
           |  +----- Hyriopsis_myersiana  
           +--+                           
              |  +-- Hyriopsis_khoratensis
              +--+                        
                 +-- Hyriopsis_bialata

Bolotov et al., 2017 [Publ 3761], fig. S1

iTtTbiriSoTroTroSiD
1101138102--
  +-------------- Ensidens             
  |                                    
--+           +-- Hyriopsis_bialata    
  |  +--------+                        
  |  |        +-- Hyriopsis_khoratensis
  +--+                                 
     |  +-------- Hyriopsis_panhai     
     +--+                              
        |  +----- Hyriopsis_myersiana  
        +--+                           
           |  +-- Rectidens_sumatrensis **
           +--+                        
              +-- Hyriopsis_desowitzi

Dai et al., 2025 [Publ 4440], fig. 3

iTtTbiriSoTroTroSiD
11018711--
  +-- Hyriopsis            
--+                        
  +-- Rectidens_sumatrensis **

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iTtTbiriSoTroTroSiD
11015622--
  +----- Ensidens             
--+                           
  |  +-- Hyriopsis            
  +--+                        
     +-- Rectidens_sumatrensis **

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
1101198102--
  +----- Hyriopsis_desowitzi  
  |                           
  +----- Rectidens_sumatrensis **
  |                           
--+----- Ensidens             
  |                           
  |  +-- Hyriopsis_bialata    
  |  |                        
  |  +-- Hyriopsis_myersiana  
  +--+                        
     +-- Hyriopsis_panhai     
     |                        
     +-- Hyriopsis_khoratensis

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iTtTbiriSoTroTroSiD
11015622--
  +----- Ensidens             
--+                           
  |  +-- Hyriopsis            
  +--+                        
     +-- Rectidens_sumatrensis **

Jeratthitikul et al., 2024 [Publ 4387], fig. 1

iTtTbiriSoTroTroSiD
11015722--
  +----- Ensidens             
--+                           
  |  +-- Hyriopsis            
  +--+                        
     +-- Rectidens_sumatrensis **

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iTtTbiriSoTroTroSiD
11017962--
  +----- Hyriopsis_desowitzi  
  |                           
  +----- Rectidens_sumatrensis **
--+                           
  +----- Ensidens             
  |                           
  |  +-- Hyriopsis_khoratensis
  +--+                        
     +-- Hyriopsis_myersiana

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

iTtTbiriSoTroTroSiD
11014711--
  +-- Hyriopsis            
--+                        
  +-- Rectidens_sumatrensis **

Wu et al., 2024 [Publ 4388], fig. 6

iTtTbiriSoTroTroSiD
11015311--
  +-- Hyriopsis            
--+                        
  +-- Rectidens_sumatrensis **

Liu et al., 2024 [Publ 4380], fig. 5

iTtTbiriSoTroTroSiD
11013632--
  +-------- Ensidens             
--+                              
  |  +----- Hyriopsis_bialata    
  +--+                           
     |  +-- Hyriopsis_desowitzi  
     +--+                        
        +-- Rectidens_sumatrensis **

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iTtTbiriSoTroTroSiD
11019411--
  +-- Hyriopsis            
--+                        
  +-- Rectidens_sumatrensis **

Konopleva et al., 2019 [Publ 4041], fig. 3

iTtTbiriSoTroTroSiD
110133112--
  +-- Ensidens_ingallsianus
  |                        
  +-- Ensidens_telus       
  |                        
  +-- Ensidens_dugasti     
  |                        
  +-- Hyriopsis_khoratensis
  |                        
  +-- Hyriopsis_bialata    
  |                        
  +-- Hyriopsis_myersiana  
  |                        
--+-- Hyriopsis_desowitzi  
  |                        
  +-- Rectidens_sumatrensis **
  |                        
  +-- Hyriopsis_panhai     
  |                        
  +-- Ensidens_jaculus__1  
  |                        
  +-- Ensidens_jaculus__2  
  |                        
  +-- Ensidens_spiculus    
  |                        
  +-- Ensidens_sagittarius

Zieritz et al., 2020 [Publ 4102], fig. 4

iTtTbiriSoTroTroSiD
110130102--
  +----------- Ensidens             
  |                                 
--+  +-------- Hyriopsis_desowitzi  
  |  |                              
  |  +-------- Rectidens_sumatrensis **
  +--+                              
     |  +----- Hyriopsis_panhai     
     |  |                           
     +--+  +-- Hyriopsis_khoratensis
        |  |                        
        +--+-- Hyriopsis_bialata    
           |                        
           +-- Hyriopsis_myersiana

Jeratthitikul et al., 2021 [Publ 4219], fig. 2

iTtTbiriSoTroTroSiD
11013522--
  +----- Ensidens             
--+                           
  |  +-- Hyriopsis            
  +--+                        
     +-- Rectidens_sumatrensis **

Jeratthitikul et al., 2021 [Publ 4212], fig. 4

iTtTbiriSoTroTroSiD
110120102--
  +----------------- Ensidens               
  |                                         
  |              +-- Hyriopsis_desowitzi    
--+  +-----------+                          
  |  |           +-- Hyriopsis_delaportei   
  |  |                                      
  +--+  +----------- Rectidens_sumatrensis **  
     |  |                                   
     |  |     +----- Hyriopsis_myersiana    
     +--+  +--+                             
        |  |  |  +-- Hyriopsis_bialata      
        |  |  +--+                          
        +--+     +-- Hyriopsis_panhai       
           |                                
           |  +----- Hyriopsis_khoratensis  
           +--+                             
              |  +-- Hyriopsis_phuphaniensis
              +--+                          
                 +-- Hyriopsis_sakhonensis

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iTtTbiriSoTroTroSiD
11019011--
  +-- Hyriopsis            
--+                        
  +-- Rectidens_sumatrensis **

Wu et al., 2024 [Publ 4350], fig. 5c

iTtTbiriSoTroTroSiD
11013632--
-- Rectidens_sumatrensis **

Jeratthitikul et al., 2021 [Publ 4219], fig. 1

iTtTbiriSoTroTroSiD
11013522--
  +----- Ensidens             
--+                           
  |  +-- Hyriopsis            
  +--+                        
     +-- Rectidens_sumatrensis **

Dai et al., 2023 [Publ 4285], fig. 5

iTtTbiriSoTroTroSiD
11015211--
  +-- Hyriopsis            
--+                        
  +-- Rectidens_sumatrensis **

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iTtTbiriSoTroTroSiD
11019011--
  +-- Hyriopsis            
--+                        
  +-- Rectidens_sumatrensis **

Matos et al., 2024 [Publ 4384], fig. 3

iTtTbiriSoTroTroSiD
11011911--
  +-- Hyriopsis            
--+                        
  +-- Rectidens_sumatrensis **

Zhang et al., 2024 [Publ 4397], fig. 5

iTtTbiriSoTroTroSiD
11012711--
  +-- Hyriopsis            
--+                        
  +-- Rectidens_sumatrensis **

Konopleva et al., 2019 [Publ 3900], fig. 1

iTtTbiriSoTroTroSiD
110151102--
  +-------------- Ensidens             
  |                                    
--+  +----------- Hyriopsis_desowitzi  
  |  |                                 
  +--+----------- Rectidens_sumatrensis **
     |                                 
     |  +-------- Hyriopsis_panhai     
     +--+                              
        |  +----- Hyriopsis_myersiana  
        +--+                           
           |  +-- Hyriopsis_khoratensis
           +--+                        
              +-- Hyriopsis_bialata

Bolotov et al., 2020 [Publ 4100], fig. 1

iTtTbiriSoTroTroSiD
1101117112--
  +-------------- Ensidens             
  |                                    
--+  +----------- Hyriopsis_desowitzi  
  |  |                                 
  +--+  +-------- Rectidens_sumatrensis **
     |  |                              
     +--+     +-- Hyriopsis_khoratensis
        |  +--+                        
        |  |  +-- Hyriopsis_bialata    
        +--+                           
           |  +-- Hyriopsis_myersiana  
           +--+                        
              +-- Hyriopsis_panhai

Pfeiffer et al., 2021 [Publ 4126], fig. 9

iTtTbiriSoTroTroSiD
110128122--
  +----- Ensidens             
--+                           
  |  +-- Rectidens_sumatrensis **
  +--+                        
     +-- Hyriopsis

Bolotov et al., 2017 [Publ 3764], fig. S1

iTtTbiriSoTroTroSiD
1101124102--
  +----------- Ensidens             
  |                                 
--+  +-------- Hyriopsis_desowitzi  
  |  |                              
  |  +-------- Rectidens_sumatrensis **
  +--+                              
     |  +----- Hyriopsis_panhai     
     |  |                           
     +--+  +-- Hyriopsis_myersiana  
        |  |                        
        +--+-- Hyriopsis_bialata    
           |                        
           +-- Hyriopsis_khoratensis

Pfeiffer et al., 2021 [Publ 4126], fig. 30

iTtTbiriSoTroTroSiD
1101661--
  +-- Rectidens_sumatrensis **
--+                        
  +-- Ensidens

Pfeiffer et al., 2018 [Publ 3827], fig. 1

iTtTbiriSoTroTroSiD
1101622--
  +-- Ensidens    
  |               
  +-- Hyriopsis__1
--+               
  +-- Hyriopsis__2
  |               
  +-- Rectidens **

Bolotov et al., 2017 [Publ 3761], fig. S2

iTtTbiriSoTroTroSiD
1101138102--
        +-- Hyriopsis_desowitzi  
        |                        
  +-----+-- Rectidens_sumatrensis **
  |     |                        
  |     +-- Ensidens             
--+                              
  |  +----- Hyriopsis_panhai     
  |  |                           
  +--+----- Hyriopsis_khoratensis
     |                           
     |  +-- Hyriopsis_myersiana  
     +--+                        
        +-- Hyriopsis_bialata

Pfeiffer et al., 2021 [Publ 4126], fig. 27

iTtTbiriSoTroTroSiD
11011091--
  +-- Rectidens_sumatrensis **
--+                        
  +-- Hyriopsis

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iTtTbiriSoTroTroSiD
110112011--
-- Rectidens_sumatrensis **

Wu et al., 2024 [Publ 4350], fig. 6-7

iTtTbiriSoTroTroSiD
11013632--
  +----- Ensidens             
--+                           
  |  +-- Rectidens_sumatrensis **
  +--+                        
     +-- Hyriopsis

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iTtTbiriSoTroTroSiD
11019411--
  +-- Hyriopsis            
--+                        
  +-- Rectidens_sumatrensis **

Wu et al., 2024 [Publ 4350], fig. 5a

iTtTbiriSoTroTroSiD
11013632--
  +----- Ensidens             
--+                           
  |  +-- Rectidens_sumatrensis **
  +--+                        
     +-- Hyriopsis

Wu et al., 2024 [Publ 4388], fig. 5

iTtTbiriSoTroTroSiD
11013732--
  +----- Ensidens             
--+                           
  |  +-- Rectidens_sumatrensis **
  +--+                        
     +-- Hyriopsis

Wu et al., 2024 [Publ 4350], fig. 5b

iTtTbiriSoTroTroSiD
11013632--
-- Rectidens_sumatrensis **

Zieritz et al., 2016 [Publ 3804], fig. S2-1

iTtTbiriSoTroTroSiD
1101551--
  +-- Rectidens_sumatrensis **
--+                        
  +-- Hyriopsis

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

iTtTbiriSoTroTroSiD
11017522--
  +-- Ensidens             
  |                        
--+-- Hyriopsis            
  |                        
  +-- Rectidens_sumatrensis **

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
1101196102--
  +-------------- Ensidens             
  |                                    
--+  +----------- Hyriopsis_desowitzi  
  |  |                                 
  +--+  +-------- Rectidens_sumatrensis **
     |  |                              
     +--+     +-- Hyriopsis_panhai     
        |  +--+                        
        |  |  +-- Hyriopsis_khoratensis
        +--+                           
           |  +-- Hyriopsis_bialata    
           +--+                        
              +-- Hyriopsis_myersiana

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

iTtTbiriSoTroTroSiD
11017522--
  +----- Ensidens             
--+                           
  |  +-- Rectidens_sumatrensis **
  +--+                        
     +-- Hyriopsis

Matos et al., 2024 [Publ 4383], fig. 3

iTtTbiriSoTroTroSiD
11011911--
  +-- Hyriopsis            
--+                        
  +-- Rectidens_sumatrensis **
 
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