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Page last updated
Thu 3 September 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

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Bivalvia | Unionida | Unionoidea | Unionidae | Gonideinae | Gonideini

genus Ptychorhynchus Simpson, 1900: cladomics
     4 Recent valid species (4 total species)

Cladomics Summary

number of species = 4 (4 total)
analyzed in 62 cladograms in 35 publications
maximum number of terminals in an analysis = 1 (mean = 1, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 0
maximum number of included terminals = , (mean = median = )
proportion of cladograms that support basic monophyly = 0

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
11016302012--
  +----------------- Parvasolenaia          
  |                                         
--+  +-------------- Pseudopostolata        
  |  |                                      
  +--+  +----------- Koreosolenaia          
     |  |                                   
     +--+        +-- Cosmopseudodon         
        |  +-----+                          
        |  |     +-- Guiunio                
        +--+                                
           |  +----- Ptychorhynchus_pfisteri **
           +--+                             
              |  +-- Obovalis               
              +--+                          
                 +-- Postolata

Wu et al., 2024 [Publ 4388], fig. 4

iTtTbiriSoTroTroSiD
1101833--
     +-- Cosmopseudodon         
  +--+                          
  |  +-- Ptychorhynchus_pfisteri **
--+                             
  |  +-- Postolata              
  +--+                          
     +-- Obovalis

Huang et al., 2019 [Publ 3843], fig. S1

iTtTbiriSoTroTroSiD
11014662--
  +----- Parvasolenaia_rivularis   
--+                                
  |  +-- Parvasolenaia_triangularis
  +--+                             
     +-- Ptychorhynchus_pfisteri **

Dai et al., 2023 [Publ 4285], fig. 3-ML

iTtTbiriSoTroTroSiD
110144139--
  +-------------- Sinosolenaia           
  |                                      
  |        +----- Gonidea                
--+  +-----+                             
  |  |     |  +-- Leguminaia             
  |  |     +--+                          
  |  |        +-- Microcondylaea         
  +--+                                   
     |        +-- Koreosolenaia          
     |  +-----+                          
     |  |     +-- Parvasolenaia          
     +--+                                
        |  +----- Postolata              
        +--+                             
           |  +-- Ptychorhynchus_pfisteri **
           +--+                          
              +-- Obovalis

Sano et al., 2017 [Publ 3799], fig. 2

iTtTbiriSoTroTroSiD
11013233--
  +-- Obovalis               
--+                          
  +-- Ptychorhynchus_pfisteri **

Wu et al., 2020 [Publ 4121], fig. 3

iTtTbiriSoTroTroSiD
11014821--
  +-- Ptychorhynchus_pfisteri **
--+                          
  +-- Sinosolenaia

Dai et al., 2025 [Publ 4440], fig. 3

iTtTbiriSoTroTroSiD
11018765--
  +----------- Inversidens            
  |                                   
--+        +-- Microcondylaea         
  |  +-----+                          
  |  |     +-- Sinosolenaia           
  +--+                                
     |  +----- Parvasolenaia          
     +--+                             
        |  +-- Postolata              
        +--+                          
           +-- Ptychorhynchus_pfisteri **

Froufe et al., 2019 [Publ 3936], fig. 6

iTtTbiriSoTroTroSiD
11014332--
  +----- Ptychorhynchus_pfisteri **
--+                             
  |  +-- Microcondylaea         
  +--+                          
     +-- Sinosolenaia

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
110119996--
  +-------- Sinosolenaia_carinata  
  |                                
  |  +----- Gonidea                
  +--+                             
  |  |  +-- Leguminaia             
  |  +--+                          
  |     +-- Microcondylaea         
--+                                
  |  +----- Ptychorhynchus_pfisteri **
  +--+                             
  |  |  +-- Parvasolenaia          
  |  +--+                          
  |     +-- Koreosolenaia          
  |                                
  |  +----- Sinosolenaia_recognita 
  +--+                             
     |  +-- Sinosolenaia_iridinea  
     +--+                          
        +-- Sinosolenaia_oleivora

Froufe et al., 2019 [Publ 3936], fig. 3

iTtTbiriSoTroTroSiD
11014332--
  +----- Ptychorhynchus_pfisteri **
--+                             
  |  +-- Microcondylaea         
  +--+                          
     +-- Sinosolenaia

Bolotov et al., 2020 [Publ 4100], fig. 1

iTtTbiriSoTroTroSiD
110111685--
-- Ptychorhynchus_pfisteri **

Dai et al., 2025 [Publ 4441], fig. 1

iTtTbiriSoTroTroSiD
1101461511--
  +-------------------- Sinosolenaia           
  |                                            
  |              +----- Gonidea                
--+  +-----------+                             
  |  |           |  +-- Leguminaia             
  |  |           +--+                          
  +--+              +-- Microcondylaea         
     |                                         
     |  +-------------- Pseudopostolata        
     |  |                                      
     +--+           +-- Koreosolenaia          
        |  +--------+                          
        |  |        +-- Parvasolenaia          
        +--+                                   
           |  +-------- Postolata              
           |  |                                
           +--+     +-- Guiunio                
              |  +--+                          
              |  |  +-- Cosmopseudodon         
              +--+                             
                 |  +-- Obovalis               
                 +--+                          
                    +-- Ptychorhynchus_pfisteri **

Wu et al., 2021 [Publ 4366], fig. 3-ML

iTtTbiriSoTroTroSiD
11017343--
     +-- Parvasolenaia          
  +--+                          
  |  +-- Ptychorhynchus_pfisteri **
--+                             
  |  +-- Microcondylaea         
  +--+                          
     +-- Sinosolenaia

Wu et al., 2018 [Publ 3836], fig. 2

iTtTbiriSoTroTroSiD
11015243--
  +-------- Sinosolenaia_carinata  
  |                                
--+  +----- Gonidea                
  |  |                             
  +--+----- Sinosolenaia_oleivora  
     |                             
     |  +-- Parvasolenaia          
     +--+                          
        +-- Ptychorhynchus_pfisteri **

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

iTtTbiriSoTroTroSiD
1101140107--
  +-------------------- Inversidens            
--+                                            
  |  +----------------- Microcondylaea         
  +--+                                         
     |  +-------------- Sinosolenaia           
     +--+                                      
        |  +----------- Koreosolenaia          
        +--+                                   
           |  +-------- Parvasolenaia          
           +--+                                
              |  +----- Ptychorhynchus_pfisteri **
              +--+                             
                 |  +-- Cosmopseudodon         
                 +--+                          
                    +-- Postolata

Huang et al., 2019 [Publ 3843], fig. 3

iTtTbiriSoTroTroSiD
11015532--
  +----- Sinosolenaia           
--+                             
  |  +-- Ptychorhynchus_pfisteri **
  +--+                          
     +-- Parvasolenaia

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iTtTbiriSoTroTroSiD
11019553--
  +-------- Microcondylaea         
--+                                
  |  +----- Sinosolenaia           
  +--+                             
     |  +-- Parvasolenaia          
     +--+                          
        +-- Ptychorhynchus_pfisteri **

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
110119786--
  +----------- Sinosolenaia           
  |                                   
  |     +----- Gonidea                
--+  +--+                             
  |  |  |  +-- Leguminaia             
  |  |  +--+                          
  +--+     +-- Microcondylaea         
     |                                
     |  +----- Ptychorhynchus_pfisteri **
     +--+                             
        |  +-- Parvasolenaia          
        +--+                          
           +-- Koreosolenaia

Dai et al., 2024 [Publ 4389], fig. 3

iTtTbiriSoTroTroSiD
1101451410--
  +-------------------- Sinosolenaia           
  |                                            
  |              +----- Gonidea                
--+  +-----------+                             
  |  |           |  +-- Leguminaia             
  |  |           +--+                          
  +--+              +-- Microcondylaea         
     |                                         
     |  +-------------- Pseudopostolata        
     |  |                                      
     +--+           +-- Parvasolenaia          
        |  +--------+                          
        |  |        +-- Koreosolenaia          
        +--+                                   
           |  +-------- Postolata              
           +--+                                
              |  +----- Ptychorhynchus_pfisteri **
              +--+                             
                 |  +-- Cosmopseudodon         
                 +--+                          
                    +-- Obovalis

Chen et al., 2025 [Publ 4663], fig. 1

iTtTbiriSoTroTroSiD
1101471612--
  +----------------------- Sinosolenaia           
  |                                               
  |                 +----- Gonidea                
--+  +--------------+                             
  |  |              |  +-- Leguminaia             
  |  |              +--+                          
  +--+                 +-- Microcondylaea         
     |                                            
     |  +----------------- Pseudopostolata        
     |  |                                         
     +--+              +-- Koreosolenaia          
        |  +-----------+                          
        |  |           +-- Parvasolenaia          
        +--+                                      
           |  +----------- Postolata              
           |  |                                   
           +--+        +-- Guiunio                
              |  +-----+                          
              |  |     +-- Cosmopseudodon         
              +--+                                
                 |  +----- Obovalis               
                 +--+                             
                    |  +-- Tenuiunio              
                    +--+                          
                       +-- Ptychorhynchus_pfisteri **

Sano et al., 2020 [Publ 4138], fig. 3

iTtTbiriSoTroTroSiD
11016722--
  +----- Inversidens            
--+                             
  |  +-- Obovalis               
  +--+                          
     +-- Ptychorhynchus_pfisteri **

Wu et al., 2021 [Publ 4215], fig. 3

iTtTbiriSoTroTroSiD
11012376--
           +-- Ptychorhynchus_pfisteri **
  +--------+                          
  |        +-- Parvasolenaia          
--+                                   
  |  +-------- Inversidens            
  +--+                                
     |  +----- Gonidea                
     +--+                             
        |  +-- Microcondylaea         
        +--+                          
           +-- Leguminaia

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iTtTbiriSoTroTroSiD
1101121118--
     +-------- Sinosolenaia           
  +--+                                
  |  |  +----- Gonidea                
  |  +--+                             
  |     |  +-- Leguminaia             
--+     +--+                          
  |        +-- Microcondylaea         
  |                                   
  |  +-------- Inversidens            
  |  |                                
  +--+     +-- Obovalis               
     |  +--+                          
     |  |  +-- Ptychorhynchus_pfisteri **
     +--+                             
        |  +-- Parvasolenaia          
        +--+                          
           +-- Koreosolenaia

Fukata & Iigo, 2020 [Publ 4106], fig. 1

iTtTbiriSoTroTroSiD
11015332--
  +----- Sinosolenaia           
--+                             
  |  +-- Parvasolenaia          
  +--+                          
     +-- Ptychorhynchus_pfisteri **

Liu et al., 2024 [Publ 4380], fig. 5

iTtTbiriSoTroTroSiD
11013587--
  +-------------- Sinosolenaia           
  |                                      
  |           +-- Gonidea                
--+  +--------+                          
  |  |        +-- Microcondylaea         
  |  |                                   
  +--+        +-- Koreosolenaia          
     |  +-----+                          
     |  |     +-- Parvasolenaia          
     +--+                                
        |  +----- Postolata              
        +--+                             
           |  +-- Obovalis               
           +--+                          
              +-- Ptychorhynchus_pfisteri **

Bolotov et al., 2020 [Publ 4100], fig. S1

iTtTbiriSoTroTroSiD
110111085--
  +-- Sinosolenaia_iridinea    
  |                            
  +-- Sinosolenaia_recognita__1
  |                            
  +-- Sinosolenaia_carinata    
  |                            
  +-- Leguminaia               
  |                            
  +-- Microcondylaea           
--+                            
  +-- Gonidea                  
  |                            
  +-- Ptychorhynchus_pfisteri **  
  |                            
  +-- Parvasolenaia            
  |                            
  +-- Sinosolenaia_recognita__2
  |                            
  +-- Sinosolenaia_oleivora

Konopleva et al., 2023 [Publ 4269], fig. S1_4

iTtTbiriSoTroTroSiD
11015555--
  +----------- Sinosolenaia           
  |                                   
--+        +-- Ptychorhynchus_pfisteri **
  |  +-----+                          
  |  |     +-- Parvasolenaia          
  +--+                                
     |  +----- Gonidea                
     +--+                             
        |  +-- Microcondylaea         
        +--+                          
           +-- Leguminaia

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iTtTbiriSoTroTroSiD
11019553--
     +-- Ptychorhynchus_pfisteri **
  +--+                          
  |  +-- Microcondylaea         
--+                             
  |  +-- Parvasolenaia          
  +--+                          
     +-- Sinosolenaia

Chen et al., 2025 [Publ 4448], fig. 3

iTtTbiriSoTroTroSiD
11018543--
     +-- Ptychorhynchus_pfisteri **
  +--+                          
  |  +-- Parvasolenaia          
--+                             
  |  +-- Microcondylaea         
  +--+                          
     +-- Sinosolenaia

Dai et al., 2024 [Publ 4358], fig. 4

iTtTbiriSoTroTroSiD
11015233--
  +-------- Microcondylaea         
--+                                
  |  +----- Sinosolenaia           
  +--+                             
     |  +-- Ptychorhynchus_pfisteri **
     +--+                          
        +-- Parvasolenaia

Liu et al., 2024 [Publ 4380], fig. 4

iTtTbiriSoTroTroSiD
11011196--
  +-------------- Sinosolenaia_carinata  
  |                                      
  |        +----- Inversidens            
--+  +-----+                             
  |  |     |  +-- Gonidea                
  |  |     +--+                          
  +--+        +-- Leguminaia             
     |                                   
     |  +-------- Sinosolenaia_oleivora  
     +--+                                
        |  +----- Postolata              
        +--+                             
           |  +-- Obovalis               
           +--+                          
              +-- Ptychorhynchus_pfisteri **

Sano et al., 2020 [Publ 4138], fig. 4

iTtTbiriSoTroTroSiD
11016722--
  +-- Obovalis               
--+                          
  +-- Ptychorhynchus_pfisteri **

Wu et al., 2024 [Publ 4350], fig. 6-7

iTtTbiriSoTroTroSiD
11013587--
  +----------------- Inversidens            
  |                                         
--+  +-------------- Sinosolenaia           
  |  |                                      
  +--+           +-- Gonidea                
     |  +--------+                          
     |  |        +-- Microcondylaea         
     +--+                                   
        |  +-------- Koreosolenaia          
        +--+                                
           |  +----- Parvasolenaia          
           +--+                             
              |  +-- Postolata              
              +--+                          
                 +-- Ptychorhynchus_pfisteri **

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iTtTbiriSoTroTroSiD
11019164--
  +-------- Inversidens            
  |                                
--+     +-- Parvasolenaia          
  |  +--+                          
  |  |  +-- Ptychorhynchus_pfisteri **
  +--+                             
     |  +-- Microcondylaea         
     +--+                          
        +-- Sinosolenaia

Dai et al., 2023 [Publ 4285], fig. 5

iTtTbiriSoTroTroSiD
11015254--
        +-- Microcondylaea         
  +-----+                          
  |     +-- Sinosolenaia           
--+                                
  |  +----- Parvasolenaia          
  +--+                             
     |  +-- Postolata              
     +--+                          
        +-- Ptychorhynchus_pfisteri **

Wu et al., 2021 [Publ 4366], fig. 3-BI

iTtTbiriSoTroTroSiD
11017343--
     +-- Parvasolenaia          
  +--+                          
  |  +-- Ptychorhynchus_pfisteri **
--+                             
  |  +-- Microcondylaea         
  +--+                          
     +-- Sinosolenaia

Zhang et al., 2024 [Publ 4397], fig. 5

iTtTbiriSoTroTroSiD
110126107--
  +----------------- Inversidens            
  |                                         
--+              +-- Microcondylaea         
  |  +-----------+                          
  |  |           +-- Sinosolenaia           
  +--+                                      
     |  +----------- Parvasolenaia          
     +--+                                   
        |  +-------- Koreosolenaia          
        +--+                                
           |  +----- Ptychorhynchus_pfisteri **
           +--+                             
              |  +-- Cosmopseudodon         
              +--+                          
                 +-- Postolata

Wu et al., 2024 [Publ 4350], fig. 5a

iTtTbiriSoTroTroSiD
11013587--
  +-------------- Sinosolenaia           
  |                                      
--+           +-- Gonidea                
  |  +--------+                          
  |  |        +-- Microcondylaea         
  +--+                                   
     |  +-------- Parvasolenaia          
     +--+                                
        |  +----- Koreosolenaia          
        +--+                             
           |  +-- Postolata              
           +--+                          
              +-- Ptychorhynchus_pfisteri **

Dai et al., 2023 [Publ 4285], fig. 3-BI

iTtTbiriSoTroTroSiD
110144139--
  +-------------- Sinosolenaia           
  |                                      
  |        +----- Gonidea                
--+  +-----+                             
  |  |     |  +-- Leguminaia             
  |  |     +--+                          
  |  |        +-- Microcondylaea         
  +--+                                   
     |        +-- Koreosolenaia          
     |  +-----+                          
     |  |     +-- Parvasolenaia          
     +--+                                
        |  +----- Postolata              
        +--+                             
           |  +-- Ptychorhynchus_pfisteri **
           +--+                          
              +-- Obovalis

Wu et al., 2021 [Publ 4366], fig. 6

iTtTbiriSoTroTroSiD
11019076--
        +----- Sinosolenaia           
  +-----+                             
  |     |  +-- Ptychorhynchus_pfisteri **
  |     +--+                          
--+        +-- Parvasolenaia          
  |                                   
  |  +-------- Inversidens            
  +--+                                
     |  +----- Gonidea                
     +--+                             
        |  +-- Microcondylaea         
        +--+                          
           +-- Leguminaia

Wu et al., 2024 [Publ 4350], fig. 5b

iTtTbiriSoTroTroSiD
11013587--
  +-------------- Sinosolenaia           
  |                                      
--+           +-- Gonidea                
  |  +--------+                          
  |  |        +-- Microcondylaea         
  +--+                                   
     |  +-------- Koreosolenaia          
     +--+                                
        |  +----- Parvasolenaia          
        +--+                             
           |  +-- Postolata              
           +--+                          
              +-- Ptychorhynchus_pfisteri **

Wang et al., 2018 [Publ 3846], fig. 2

iTtTbiriSoTroTroSiD
11013021--
  +-- Ptychorhynchus_pfisteri **
--+                          
  +-- Sinosolenaia

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iTtTbiriSoTroTroSiD
11019164--
  +-------- Inversidens            
  |                                
--+     +-- Parvasolenaia          
  |  +--+                          
  |  |  +-- Ptychorhynchus_pfisteri **
  +--+                             
     |  +-- Microcondylaea         
     +--+                          
        +-- Sinosolenaia

Cha et al., 2018 [Publ 3916], fig. 1

iTtTbiriSoTroTroSiD
11013721--
  +-- Ptychorhynchus_pfisteri **
--+                          
  +-- Sinosolenaia

Huang et al., 2002 [Publ 2657], fig. 1

iTtTbiriSoTroTroSiD
11013311--
-- Ptychorhynchus_pfisteri **

Wu et al., 2019 [Publ 3864], fig. 3

iTtTbiriSoTroTroSiD
11014521--
  +-- Ptychorhynchus_pfisteri **
--+                          
  +-- Sinosolenaia

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

iTtTbiriSoTroTroSiD
11014732--
  +----- Ptychorhynchus_pfisteri **
--+                             
  |  +-- Microcondylaea         
  +--+                          
     +-- Sinosolenaia

Huang et al., 2019 [Publ 3843], fig. 1

iTtTbiriSoTroTroSiD
11014462--
  +----- Parvasolenaia          
--+                             
  |  +-- Ptychorhynchus_pfisteri **
  +--+                          
     +-- Sinosolenaia

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
110119996--
  +----------- Sinosolenaia           
  |                                   
  |     +----- Gonidea                
--+  +--+                             
  |  |  |  +-- Leguminaia             
  |  |  +--+                          
  +--+     +-- Microcondylaea         
     |                                
     |     +-- Ptychorhynchus_pfisteri **
     |     |                          
     +-----+-- Parvasolenaia          
           |                          
           +-- Koreosolenaia

Huang et al., 2002 [Publ 2657], fig. 2

iTtTbiriSoTroTroSiD
11013311--
-- Ptychorhynchus_pfisteri **

Wu et al., 2024 [Publ 4388], fig. 6

iTtTbiriSoTroTroSiD
11015275--
           +-- Microcondylaea         
  +--------+                          
  |        +-- Sinosolenaia           
--+                                   
  |  +-------- Parvasolenaia          
  +--+                                
     |  +----- Ptychorhynchus_pfisteri **
     +--+                             
        |  +-- Cosmopseudodon         
        +--+                          
           +-- Postolata

Wu et al., 2021 [Publ 4366], fig. 2_I

iTtTbiriSoTroTroSiD
11019076--
        +----- Gonidea                
  +-----+                             
  |     |  +-- Leguminaia             
  |     +--+                          
--+        +-- Microcondylaea         
  |                                   
  |  +-------- Inversidens            
  +--+                                
     |  +----- Sinosolenaia           
     +--+                             
        |  +-- Parvasolenaia          
        +--+                          
           +-- Ptychorhynchus_pfisteri **

Wu et al., 2026 [Publ 4811], fig. 4

iTtTbiriSoTroTroSiD
110194127--
  +-------------------- Inversidens            
--+                                            
  |  +----------------- Sinosolenaia           
  +--+                                         
     |  +-------------- Parvasolenaia          
     +--+                                      
        |  +----------- Pseudopostolata        
        +--+                                   
           |  +-------- Koreosolenaia          
           +--+                                
              |  +----- Ptychorhynchus_pfisteri **
              +--+                             
                 |  +-- Cosmopseudodon         
                 +--+                          
                    +-- Postolata

Wu et al., 2024 [Publ 4388], fig. 5

iTtTbiriSoTroTroSiD
11013588--
  +-------------------- Sinosolenaia           
  |                                            
--+                 +-- Gonidea                
  |  +--------------+                          
  |  |              +-- Microcondylaea         
  +--+                                         
     |  +-------------- Koreosolenaia          
     +--+                                      
        |  +----------- Parvasolenaia          
        +--+                                   
           |  +-------- Postolata              
           +--+                                
              |  +----- Cosmopseudodon         
              +--+                             
                 |  +-- Ptychorhynchus_pfisteri **
                 +--+                          
                    +-- Obovalis

Aunins et al., 2018 [Publ 3863], fig. 1

iTtTbiriSoTroTroSiD
11013921--
  +----- Sinosolenaia_oleivora  
--+                             
  |  +-- Ptychorhynchus_pfisteri **
  +--+                          
     +-- Sinosolenaia_carinata

Wu et al., 2021 [Publ 4366], fig. 4

iTtTbiriSoTroTroSiD
11016843--
     +-- Parvasolenaia          
  +--+                          
  |  +-- Ptychorhynchus_pfisteri **
--+                             
  |  +-- Microcondylaea         
  +--+                          
     +-- Sinosolenaia

Sano et al., 2020 [Publ 4138], fig. 1

iTtTbiriSoTroTroSiD
11013333--
  +-------- Sinosolenaia           
  |                                
--+  +----- Inversidens            
  |  |                             
  +--+  +-- Obovalis_omiensis__1   
     |  |                          
     +--+-- Obovalis_omiensis__2   
        |                          
        +-- Ptychorhynchus_pfisteri **

Wu et al., 2022 [Publ 4238], fig. 3

iTtTbiriSoTroTroSiD
11017343--
     +-- Ptychorhynchus_pfisteri **
  +--+                          
  |  +-- Parvasolenaia          
--+                             
  |  +-- Microcondylaea         
  +--+                          
     +-- Sinosolenaia

Wu et al., 2024 [Publ 4350], fig. 5c

iTtTbiriSoTroTroSiD
11013587--
  +-------------- Sinosolenaia           
  |                                      
--+  +----------- Gonidea                
  |  |                                   
  +--+----------- Microcondylaea         
     |                                   
     |  +-------- Koreosolenaia          
     +--+                                
        |  +----- Parvasolenaia          
        +--+                             
           |  +-- Postolata              
           +--+                          
              +-- Ptychorhynchus_pfisteri **

Wang et al., 2018 [Publ 3865], fig. 2

iTtTbiriSoTroTroSiD
11013021--
  +-- Ptychorhynchus_pfisteri **
--+                          
  +-- Sinosolenaia

Sano et al., 2020 [Publ 4138], fig. 2

iTtTbiriSoTroTroSiD
11016722--
  +-- Ptychorhynchus_pfisteri **
--+                          
  +-- Obovalis

Zhou et al., 2017 [Publ 3788], fig. 2

iTtTbiriSoTroTroSiD
11011521--
  +-- Ptychorhynchus_pfisteri **
--+                          
  +-- Sinosolenaia
 
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