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Page last updated
Thu 3 September 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Search the database pages:

Bivalvia | Unionida | Unionoidea | Unionidae | Parreysiinae | Lamellidentini

genus Lamellidens Simpson, 1900: cladomics
     13 Recent valid species (14 total species)

Cladomics Summary

number of species = 13 (14 total)
analyzed in 47 cladograms in 29 publications
maximum number of terminals in an analysis = 10 (mean = 3.66, median = 2)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 24
maximum number of included terminals = 10, (mean = 6.21 median = 6)
proportion of cladograms that support basic monophyly = 0.92

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.


PCoA Ordination


NJ Tree

Bolotov et al., 2024 [Publ 4369], fig. 2

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
101081052720
                       +-- Balwantia                
  +--------------------+                            
  |                    +-- Trapezidens              
  |                                                 
--+                    +-- Lamellidens_nongyangensis **
  |  +-----------------+                            
  |  |                 +-- Lamellidens_corrianus **    
  |  |                                              
  +--+                 +-- Lamellidens_savadiensis **  
     |  +--------------+                            
     |  |              +-- Lamellidens_chindwinensis **
     +--+                                           
        |  +-------------- Lamellidens_ferrugineus **  
        +--+                                        
           |  +----------- Lamellidens_marginalis **   
           +--+                                     
              |  +-------- Lamellidens_generosus **    
              +--+                                  
                 |  +----- Lamellidens_mainwaringi **  
                 +--+                               
                    |  +-- Lamellidens_jenkinsianus ** 
                    +--+                            
                       +-- Lamellidens_lamellatus **

Bolotov et al., 2022 [Publ 4252], fig. 2

iTtTbiriSoTroTroSiD
101071051619
  +----------------- Trapezidens              
  |                                           
--+  +-------------- Lamellidens_marginalis **   
  |  |                                        
  |  |  +----------- Lamellidens_ferrugineus **  
  +--+  |                                     
     |  |  +-------- Lamellidens_generosus **    
     |  +--+                                  
     |  |  |  +----- Lamellidens_mainwaringi **  
     +--+  +--+                               
        |     |  +-- Lamellidens_lamellatus **   
        |     +--+                            
        |        +-- Lamellidens_jenkinsianus ** 
        |                                     
        |  +-------- Lamellidens_nongyangensis **
        +--+                                  
           |  +----- Lamellidens_savadiensis **  
           +--+                               
              |  +-- Lamellidens_corrianus **    
              +--+                            
                 +-- Lamellidens_chindwinensis **

Bolotov et al., 2024 [Publ 4369], fig. S1

iTtTbiriSoTroTroSiD
101081053726
                 +-- Balwantia                
  +--------------+                            
  |              +-- Trapezidens              
  |                                           
  |        +-------- Lamellidens_savadiensis **  
--+  +-----+                                  
  |  |     |  +----- Lamellidens_chindwinensis **
  |  |     +--+                               
  |  |        |  +-- Lamellidens_corrianus **    
  |  |        +--+                            
  +--+           +-- Lamellidens_nongyangensis **
     |                                        
     |           +-- Lamellidens_marginalis **   
     |  +--------+                            
     |  |        +-- Lamellidens_ferrugineus **  
     +--+                                     
        |  +-------- Lamellidens_generosus **    
        +--+                                  
           |  +----- Lamellidens_mainwaringi **  
           +--+                               
              |  +-- Lamellidens_jenkinsianus ** 
              +--+                            
                 +-- Lamellidens_lamellatus **

Bolotov et al., 2022 [Publ 4252], fig. S1

iTtTbiriSoTroTroSiD
101081051616
  +----------------- Trapezidens              
  |                                           
  |        +-------- Lamellidens_savadiensis **  
  |  +-----+                                  
--+  |     |  +----- Lamellidens_chindwinensis **
  |  |     +--+                               
  |  |        |  +-- Lamellidens_corrianus **    
  |  |        +--+                            
  +--+           +-- Lamellidens_nongyangensis **
     |                                        
     |           +-- Lamellidens_marginalis **   
     |  +--------+                            
     |  |        +-- Lamellidens_ferrugineus **  
     +--+                                     
        |  +-------- Lamellidens_generosus **    
        +--+                                  
           |  +----- Lamellidens_mainwaringi **  
           +--+                               
              |  +-- Lamellidens_jenkinsianus ** 
              +--+                            
                 +-- Lamellidens_lamellatus **

Bolotov et al., 2022 [Publ 4239], fig. S2

iTtTbiriSoTroTroSiD
997949519
  +----------------- Trapezidens              
  |                                           
  |           +----- Lamellidens_savadiensis **  
--+  +--------+                               
  |  |        |  +-- Lamellidens_corrianus **    
  |  |        +--+                            
  |  |           +-- Lamellidens_nongyangensis **
  +--+                                        
     |           +-- Lamellidens_marginalis **   
     |  +--------+                            
     |  |        +-- Lamellidens_ferrugineus **  
     +--+                                     
        |  +-------- Lamellidens_lamellatus **   
        +--+                                  
           |  +----- Lamellidens_jenkinsianus ** 
           +--+                               
              |  +-- Lamellidens_mainwaringi **  
              +--+                            
                 +-- Lamellidens_generosus **

Bolotov et al., 2022 [Publ 4239], fig. 2

iTtTbiriSoTroTroSiD
997949517
  +-------------------- Trapezidens              
  |                                              
--+  +----------------- Lamellidens_corrianus **    
  |  |                                           
  +--+  +-------------- Lamellidens_nongyangensis **
     |  |                                        
     +--+  +----------- Lamellidens_savadiensis **  
        |  |                                     
        |  |        +-- Lamellidens_ferrugineus **  
        +--+  +-----+                            
           |  |     +-- Lamellidens_marginalis **   
           |  |                                  
           +--+     +-- Lamellidens_lamellatus **   
              |  +--+                            
              |  |  +-- Lamellidens_jenkinsianus ** 
              +--+                               
                 |  +-- Lamellidens_generosus **    
                 +--+                            
                    +-- Lamellidens_mainwaringi **

Bolotov et al., 2022 [Publ 4239], fig. S1

iTtTbiriSoTroTroSiD
997949519
  +----------------- Trapezidens              
  |                                           
  |           +----- Lamellidens_savadiensis **  
--+  +--------+                               
  |  |        |  +-- Lamellidens_corrianus **    
  |  |        +--+                            
  |  |           +-- Lamellidens_nongyangensis **
  +--+                                        
     |           +-- Lamellidens_ferrugineus **  
     |  +--------+                            
     |  |        +-- Lamellidens_marginalis **   
     +--+                                     
        |  +-------- Lamellidens_lamellatus **   
        +--+                                  
           |  +----- Lamellidens_jenkinsianus ** 
           +--+                               
              |  +-- Lamellidens_generosus **    
              +--+                            
                 +-- Lamellidens_mainwaringi **

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
8879193319
  +----------------------- Trapezidens               
  |                                                  
--+  +-------------------- Lamellidens_corrianus **     
  |  |                                               
  +--+  +----------------- Lamellidens_savadiensis **   
     |  |                                            
     +--+              +-- Lamellidens_lamellatus **    
        |  +-----------+                             
        |  |           +-- Lamellidens_jenkinsianus **  
        +--+                                         
           |  +----------- Lamellidens_generosus **     
           +--+                                      
              |  +-------- Lamellidens_mainwaringi__2 **
              +--+                                   
                 |  +----- Lamellidens_mainwaringi__1 **
                 +--+                                
                    |  +-- Lamellidens_ferrugineus **   
                    +--+                             
                       +-- Lamellidens_marginalis **

Bolotov et al., 2017 [Publ 3764], fig. S1

iTtTbiriSoTroTroSiD
8382891173143
           +-------------- Trapezidens                  
           |                                            
  +--------+  +----------- Lamellidens_lamellatus **       
  |        |  |                                         
  |        +--+  +-------- Lamellidens_jenkinsianus **     
  |           |  |                                      
  |           |  |  +----- Lamellidens_generosus **        
  |           +--+  |                                   
  |              |  +----- Lamellidens_mainwaringi **      
  |              |  |                                   
  |              +--+  +-- Lamellidens_ferrugineus **      
  |                 +--+                                
--+                 |  +-- Lamellidens_marginalis **       
  |                 |                                   
  |                 |  +-- Lamellidens_savadiensis **      
  |                 +--+                                
  |                    +-- Lamellidens_corrianus__1 **     
  |                                                     
  |  +-------------------- Parreysia                    
  |  |                                                  
  |  |  +----------------- Leoparreysia                 
  |  |  |                                               
  |  |  |              +-- Nitia                        
  +--+  |              |                                
     |  |              +-- Coelatura_gabonensis         
     |  |              |                                
     |  |              +-- Prisodontopsis_mweruensis    
     |  |              |                                
     +--+              +-- Coelatura_kunenensis         
        |              |                                
        |  +-----------+-- Coelatura_aegyptiaca__1      
        |  |           |                                
        |  |           +-- Coelatura_aegyptiaca__2      
        |  |           |                                
        |  |           +-- Prisodontopsis_aviculaeformis
        |  |           |                                
        |  |           +-- Coelatura_luapulaensis       
        +--+           |                                
           |           +-- Coelatura_choziensis         
           |                                            
           |        +----- Indochinella                 
           |  +-----+                                   
           |  |     |  +-- Scabies                      
           |  |     +--+                                
           +--+        +-- Radiatula                    
              |                                         
              |  +-------- Indonaia_shurtleffiana       
              |  |                                      
              +--+  +----- Indonaia_rugosa              
                 |  |                                   
                 |  |  +-- Indonaia_andersoniana        
                 +--+--+                                
                    |  +-- Indonaia_caerulea            
                    |                                   
                    |  +-- Indonaia_bonneaudii          
                    +--+                                
                       +-- Lamellidens_corrianus__2 **

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
8839193316
  +-------------- Trapezidens               
  |                                         
--+  +----------- Lamellidens_corrianus **     
  |  |                                      
  +--+  +-------- Lamellidens_savadiensis **   
     |  |                                   
     |  |  +----- Lamellidens_marginalis **    
     +--+  |                                
        |  +----- Lamellidens_ferrugineus **   
        |  |                                
        +--+  +-- Lamellidens_mainwaringi__1 **
           |  |                             
           |  +-- Lamellidens_mainwaringi__2 **
           |  |                             
           +--+-- Lamellidens_lamellatus **    
              |                             
              +-- Lamellidens_generosus **     
              |                             
              +-- Lamellidens_jenkinsianus **

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
88491913111
  +----------- Trapezidens               
  |                                      
  |        +-- Lamellidens_corrianus **     
--+  +-----+                             
  |  |     +-- Lamellidens_savadiensis **   
  |  |                                   
  |  |  +----- Lamellidens_lamellatus **    
  +--+  |                                
     |  +----- Lamellidens_jenkinsianus **  
     |  |                                
     |  |  +-- Lamellidens_marginalis **    
     +--+--+                             
        |  +-- Lamellidens_ferrugineus **   
        |                                
        |  +-- Lamellidens_mainwaringi__1 **
        |  |                             
        +--+-- Lamellidens_mainwaringi__2 **
           |                             
           +-- Lamellidens_generosus **

Bolotov et al., 2017 [Publ 3761], fig. S2

iTtTbiriSoTroTroSiD
6636133217
  +----------- Trapezidens             
  |                                    
--+  +-------- Lamellidens_savadiensis ** 
  |  |                                 
  |  +-------- Lamellidens_corrianus **   
  +--+                                 
     |     +-- Lamellidens_marginalis **  
     |  +--+                           
     |  |  +-- Lamellidens_ferrugineus ** 
     +--+                              
        |  +-- Lamellidens_jenkinsianus **
        +--+                           
           +-- Lamellidens_mainwaringi **

Bolotov et al., 2017 [Publ 3761], fig. S1

iTtTbiriSoTroTroSiD
6646133216
  +-------------- Trapezidens             
  |                                       
--+  +----------- Lamellidens_corrianus **   
  |  |                                    
  +--+  +-------- Lamellidens_savadiensis ** 
     |  |                                 
     +--+     +-- Lamellidens_jenkinsianus **
        |  +--+                           
        |  |  +-- Lamellidens_mainwaringi ** 
        +--+                              
           |  +-- Lamellidens_ferrugineus ** 
           +--+                           
              +-- Lamellidens_marginalis **

Konopleva et al., 2020 [Publ 4072], fig. 2

iTtTbiriSoTroTroSiD
44248518
  +----------- Trapezidens            
--+                                   
  |  +-------- Lamellidens_savadiensis **
  +--+                                
     |  +----- Lamellidens_generosus **  
     +--+                             
        |  +-- Lamellidens_mainwaringi **
        +--+                          
           +-- Lamellidens_ferrugineus **

Bolotov et al., 2020 [Publ 4100], fig. S1

iTtTbiriSoTroTroSiD
4424108418
  +----------- Trapezidens            
--+                                   
  |  +-------- Lamellidens_savadiensis **
  +--+                                
     |  +----- Lamellidens_generosus **  
     +--+                             
        |  +-- Lamellidens_ferrugineus **
        +--+                          
           +-- Lamellidens_mainwaringi **

Bolotov et al., 2022 [Publ 4250], fig. S4

iTtTbiriSoTroTroSiD
44245516
  +----------- Trapezidens            
--+                                   
  |  +-------- Lamellidens_savadiensis **
  +--+                                
     |  +----- Lamellidens_ferrugineus **
     +--+                             
        |  +-- Lamellidens_mainwaringi **
        +--+                          
           +-- Lamellidens_generosus **

Bolotov et al., 2018 [Publ 3826], fig. 2

iTtTbiriSoTroTroSiD
442479216
  +----------- Trapezidens            
--+                                   
  |  +-------- Lamellidens_savadiensis **
  +--+                                
     |  +----- Lamellidens_ferrugineus **
     +--+                             
        |  +-- Lamellidens_generosus **  
        +--+                          
           +-- Lamellidens_mainwaringi **

Bolotov et al., 2020 [Publ 4100], fig. 1

iTtTbiriSoTroTroSiD
4424114418
  +----------- Trapezidens            
--+                                   
  |  +-------- Lamellidens_savadiensis **
  +--+                                
     |  +----- Lamellidens_generosus **  
     +--+                             
        |  +-- Lamellidens_ferrugineus **
        +--+                          
           +-- Lamellidens_mainwaringi **

Bolotov et al., 2022 [Publ 4250], fig. 8

iTtTbiriSoTroTroSiD
442445516
  +----------- Trapezidens            
--+                                   
  |  +-------- Lamellidens_savadiensis **
  +--+                                
     |  +----- Lamellidens_ferrugineus **
     +--+                             
        |  +-- Lamellidens_generosus **  
        +--+                          
           +-- Lamellidens_mainwaringi **

Wu et al., 2021 [Publ 4366], fig. 2_I

iTtTbiriSoTroTroSiD
331388117
  +-------- Trapezidens            
--+                                
  |  +----- Lamellidens_savadiensis **
  +--+                             
     |  +-- Lamellidens_generosus **  
     +--+                          
        +-- Lamellidens_ferrugineus **

Konopleva et al., 2017 [Publ 3807], fig. 5

iTtTbiriSoTroTroSiD
33132217
  +-------- Trapezidens             
--+                                 
  |  +----- Lamellidens_savadiensis ** 
  +--+                              
     |  +-- Lamellidens_mainwaringi ** 
     +--+                           
        +-- Lamellidens_jenkinsianus **

Wu et al., 2021 [Publ 4366], fig. 6

iTtTbiriSoTroTroSiD
331388117
  +-------- Trapezidens            
--+                                
  |  +----- Lamellidens_savadiensis **
  +--+                             
     |  +-- Lamellidens_ferrugineus **
     +--+                          
        +-- Lamellidens_generosus **

Konopleva et al., 2017 [Publ 3807], fig. 6

iTtTbiriSoTroTroSiD
331328217
  +-------- Trapezidens             
--+                                 
  |  +----- Lamellidens_savadiensis ** 
  +--+                              
     |  +-- Lamellidens_jenkinsianus **
     +--+                           
        +-- Lamellidens_mainwaringi **

Sonowal et al., 2021 [Publ 4221], fig. 4

iTtTbiriSoTroTroSiD
21002140016
  +-------------------------- Lamellidens_mainwaringi **
--+                                                  
  |  +----------------------- Lamprotula             
  +--+                                               
     |  +-------------------- Potomida               
     +--+                                            
        |  +----------------- Lampsilis              
        +--+                                         
           |  +-------------- Elliptio               
           +--+                                      
              |  +----------- Aculamprotula          
              +--+                                   
                 |  +-------- Unio_pictorum          
                 +--+                                
                    |  +----- Anodonta               
                    +--+                             
                       |  +-- Unio_ravoisieri        
                       +--+                          
                          +-- Lamellidens_corrianus **

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

iTtTbiriSoTroTroSiD
11013800--
-- Lamellidens_jenkinsianus **

Whelan et al., 2011 [Publ 3380], fig. 1-MP

iTtTbiriSoTroTroSiD
11013511--
  +-- Lamellidens_savadiensis **
--+                          
  +-- Trapezidens

Choi et al., 2020 [Publ 4169], fig. 5

iTtTbiriSoTroTroSiD
1101900--
-- Lamellidens_generosus **

Sonowal et al., 2021 [Publ 4221], fig. 3a

iTtTbiriSoTroTroSiD
11011400--
-- Lamellidens_corrianus **

Sonowal et al., 2021 [Publ 4221], fig. 3b

iTtTbiriSoTroTroSiD
11011400--
-- Lamellidens_mainwaringi **

Araujo et al., 2017 [Publ 3801], fig. 2

iTtTbiriSoTroTroSiD
11011300--
-- Lamellidens_savadiensis **

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iTtTbiriSoTroTroSiD
11019500--
-- Lamellidens_jenkinsianus **

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iTtTbiriSoTroTroSiD
11019100--
-- Lamellidens_jenkinsianus **

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

iTtTbiriSoTroTroSiD
11013911--
  +-- Trapezidens            
--+                          
  +-- Lamellidens_savadiensis **

Pfeiffer et al., 2018 [Publ 3827], fig. 2

iTtTbiriSoTroTroSiD
11012711--
  +-- Trapezidens            
--+                          
  +-- Lamellidens_savadiensis **

Konopleva et al., 2023 [Publ 4269], fig. S1_4

iTtTbiriSoTroTroSiD
11015511--
  +-- Trapezidens            
--+                          
  +-- Lamellidens_savadiensis **

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iTtTbiriSoTroTroSiD
11019500--
-- Lamellidens_jenkinsianus **

Jeratthitikul et al., 2019 [Publ 3922], fig. 1

iTtTbiriSoTroTroSiD
11012211--
  +-- Lamellidens_mainwaringi **
--+                          
  +-- Trapezidens

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iTtTbiriSoTroTroSiD
11015611--
  +-- Trapezidens            
--+                          
  +-- Lamellidens_savadiensis **

Imsong & Murali, 2023 [Publ 4370], fig. 2

iTtTbiriSoTroTroSiD
1101200--
-- Lamellidens_nongyangensis **

Jadhav & Jamkhedkar, 2009 [Publ 3697], fig. 11

iTtTbiriSoTroTroSiD
11012400--
-- Lamellidens_corrianus **

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iTtTbiriSoTroTroSiD
11015611--
  +-- Trapezidens            
--+                          
  +-- Lamellidens_savadiensis **

Whelan et al., 2011 [Publ 3380], fig. 1-ML

iTtTbiriSoTroTroSiD
11013511--
  +-- Lamellidens_savadiensis **
--+                          
  +-- Trapezidens

Zieritz et al., 2024 [Publ 4381], fig. 2

iTtTbiriSoTroTroSiD
11014500--
-- Lamellidens_corrianus **

Wu et al., 2024 [Publ 4388], fig. 6

iTtTbiriSoTroTroSiD
11015200--
-- Lamellidens_marginalis **

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iTtTbiriSoTroTroSiD
11019100--
-- Lamellidens_jenkinsianus **

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

iTtTbiriSoTroTroSiD
11013000--
-- Lamellidens_savadiensis **

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

iTtTbiriSoTroTroSiD
11013600--
-- Lamellidens_jenkinsianus **
 
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