The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)
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Bivalvia | Unionida | Unionoidea | Unionidae | Parreysiinae | Lamellidentini genus Lamellidens Simpson, 1900: cladomics
13 Recent valid species (14 total species) Cladomics Summarynumber of species = 13 (14 total)
analyzed in 50 cladograms in 31 publications
maximum number of terminals in an analysis =
10 (mean = 3.68, median = 1.5) Basic Monophylynumber of cladograms that test basic monophyly
(i.e., more than 1 included species) = 25
maximum number of included terminals = 10, (mean =
6.36 median = 6)
proportion of cladograms that support basic monophyly
= 0.92
Pairwise Cladogram ClusteringThe graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.
Bolotov et al., 2024 [Publ 4369], fig. 2 ** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram ** iT| tT | bi | riS | oT | roT | roS | iD | | 10 | 10 | 8 | 10 | 52 | 7 | 2 | 0 | +-- Balwantia
+--------------------+
| +-- Trapezidens
|
--+ +-- Lamellidens_nongyangensis **
| +-----------------+
| | +-- Lamellidens_corrianus **
| |
+--+ +-- Lamellidens_savadiensis **
| +--------------+
| | +-- Lamellidens_chindwinensis **
+--+
| +-------------- Lamellidens_ferrugineus **
+--+
| +----------- Lamellidens_marginalis **
+--+
| +-------- Lamellidens_generosus **
+--+
| +----- Lamellidens_mainwaringi **
+--+
| +-- Lamellidens_jenkinsianus **
+--+
+-- Lamellidens_lamellatus **
Bolotov et al., 2022 [Publ 4252], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 10 | 10 | 7 | 10 | 51 | 6 | 1 | 9 | +----------------- Trapezidens
|
--+ +-------------- Lamellidens_marginalis **
| |
| | +----------- Lamellidens_ferrugineus **
+--+ |
| | +-------- Lamellidens_generosus **
| +--+
| | | +----- Lamellidens_mainwaringi **
+--+ +--+
| | +-- Lamellidens_lamellatus **
| +--+
| +-- Lamellidens_jenkinsianus **
|
| +-------- Lamellidens_nongyangensis **
+--+
| +----- Lamellidens_savadiensis **
+--+
| +-- Lamellidens_corrianus **
+--+
+-- Lamellidens_chindwinensis **
Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14 iT| tT | bi | riS | oT | roT | roS | iD | | 10 | 10 | 8 | 10 | 620 | 7 | 2 | 12 | +-------- Lamellidens_savadiensis **
+-----+
| | +----- Lamellidens_corrianus **
| +--+
| | +-- Lamellidens_chindwinensis **
| +--+
--+ +-- Lamellidens_nongyangensis **
|
| +-- Lamellidens_ferrugineus **
| +--------+
| | +-- Lamellidens_marginalis **
+--+
| +-------- Lamellidens_lamellatus **
+--+
| +----- Lamellidens_jenkinsianus **
+--+
| +-- Lamellidens_generosus **
+--+
+-- Lamellidens_mainwaringi **
Bolotov et al., 2024 [Publ 4369], fig. S1 iT| tT | bi | riS | oT | roT | roS | iD | | 10 | 10 | 8 | 10 | 53 | 7 | 2 | 6 | +-- Balwantia
+--------------+
| +-- Trapezidens
|
| +-------- Lamellidens_savadiensis **
--+ +-----+
| | | +----- Lamellidens_chindwinensis **
| | +--+
| | | +-- Lamellidens_corrianus **
| | +--+
+--+ +-- Lamellidens_nongyangensis **
|
| +-- Lamellidens_marginalis **
| +--------+
| | +-- Lamellidens_ferrugineus **
+--+
| +-------- Lamellidens_generosus **
+--+
| +----- Lamellidens_mainwaringi **
+--+
| +-- Lamellidens_jenkinsianus **
+--+
+-- Lamellidens_lamellatus **
Bolotov et al., 2022 [Publ 4252], fig. S1 iT| tT | bi | riS | oT | roT | roS | iD | | 10 | 10 | 8 | 10 | 51 | 6 | 1 | 6 | +----------------- Trapezidens
|
| +-------- Lamellidens_savadiensis **
| +-----+
--+ | | +----- Lamellidens_chindwinensis **
| | +--+
| | | +-- Lamellidens_corrianus **
| | +--+
+--+ +-- Lamellidens_nongyangensis **
|
| +-- Lamellidens_marginalis **
| +--------+
| | +-- Lamellidens_ferrugineus **
+--+
| +-------- Lamellidens_generosus **
+--+
| +----- Lamellidens_mainwaringi **
+--+
| +-- Lamellidens_jenkinsianus **
+--+
+-- Lamellidens_lamellatus **
Bolotov et al., 2022 [Publ 4239], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 9 | 9 | 7 | 9 | 49 | 5 | 1 | 7 | +-------------------- Trapezidens
|
--+ +----------------- Lamellidens_corrianus **
| |
+--+ +-------------- Lamellidens_nongyangensis **
| |
+--+ +----------- Lamellidens_savadiensis **
| |
| | +-- Lamellidens_ferrugineus **
+--+ +-----+
| | +-- Lamellidens_marginalis **
| |
+--+ +-- Lamellidens_lamellatus **
| +--+
| | +-- Lamellidens_jenkinsianus **
+--+
| +-- Lamellidens_generosus **
+--+
+-- Lamellidens_mainwaringi **
Bolotov et al., 2022 [Publ 4239], fig. S2 iT| tT | bi | riS | oT | roT | roS | iD | | 9 | 9 | 7 | 9 | 49 | 5 | 1 | 9 | +----------------- Trapezidens
|
| +----- Lamellidens_savadiensis **
--+ +--------+
| | | +-- Lamellidens_corrianus **
| | +--+
| | +-- Lamellidens_nongyangensis **
+--+
| +-- Lamellidens_marginalis **
| +--------+
| | +-- Lamellidens_ferrugineus **
+--+
| +-------- Lamellidens_lamellatus **
+--+
| +----- Lamellidens_jenkinsianus **
+--+
| +-- Lamellidens_mainwaringi **
+--+
+-- Lamellidens_generosus **
Bolotov et al., 2022 [Publ 4239], fig. S1 iT| tT | bi | riS | oT | roT | roS | iD | | 9 | 9 | 7 | 9 | 49 | 5 | 1 | 9 | +----------------- Trapezidens
|
| +----- Lamellidens_savadiensis **
--+ +--------+
| | | +-- Lamellidens_corrianus **
| | +--+
| | +-- Lamellidens_nongyangensis **
+--+
| +-- Lamellidens_ferrugineus **
| +--------+
| | +-- Lamellidens_marginalis **
+--+
| +-------- Lamellidens_lamellatus **
+--+
| +----- Lamellidens_jenkinsianus **
+--+
| +-- Lamellidens_generosus **
+--+
+-- Lamellidens_mainwaringi **
Bolotov et al., 2017 [Publ 3764], fig. S1 iT| tT | bi | riS | oT | roT | roS | iD | | 8 | 38 | 28 | 9 | 117 | 3 | 1 | 43 | +-------------- Trapezidens
|
+--------+ +----------- Lamellidens_lamellatus **
| | |
| +--+ +-------- Lamellidens_jenkinsianus **
| | |
| | | +----- Lamellidens_generosus **
| +--+ |
| | +----- Lamellidens_mainwaringi **
| | |
| +--+ +-- Lamellidens_ferrugineus **
| +--+
--+ | +-- Lamellidens_marginalis **
| |
| | +-- Lamellidens_savadiensis **
| +--+
| +-- Lamellidens_corrianus__1 **
|
| +-------------------- Parreysia
| |
| | +----------------- Leoparreysia
| | |
| | | +-- Nitia
+--+ | |
| | +-- Coelatura_gabonensis
| | |
| | +-- Prisodontopsis_mweruensis
| | |
+--+ +-- Coelatura_kunenensis
| |
| +-----------+-- Coelatura_aegyptiaca__1
| | |
| | +-- Coelatura_aegyptiaca__2
| | |
| | +-- Prisodontopsis_aviculaeformis
| | |
| | +-- Coelatura_luapulaensis
+--+ |
| +-- Coelatura_choziensis
|
| +----- Indochinella
| +-----+
| | | +-- Scabies
| | +--+
+--+ +-- Radiatula
|
| +-------- Indonaia_shurtleffiana
| |
+--+ +----- Indonaia_rugosa
| |
| | +-- Indonaia_andersoniana
+--+--+
| +-- Indonaia_caerulea
|
| +-- Indonaia_bonneaudii
+--+
+-- Lamellidens_corrianus__2 **
Huang et al., 2019 [Publ 3843], fig. S4 iT| tT | bi | riS | oT | roT | roS | iD | | 8 | 8 | 3 | 9 | 193 | 3 | 1 | 6 | +-------------- Trapezidens
|
--+ +----------- Lamellidens_corrianus **
| |
+--+ +-------- Lamellidens_savadiensis **
| |
| | +----- Lamellidens_marginalis **
+--+ |
| +----- Lamellidens_ferrugineus **
| |
+--+ +-- Lamellidens_mainwaringi__1 **
| |
| +-- Lamellidens_mainwaringi__2 **
| |
+--+-- Lamellidens_lamellatus **
|
+-- Lamellidens_generosus **
|
+-- Lamellidens_jenkinsianus **
Huang et al., 2019 [Publ 3843], fig. S3 iT| tT | bi | riS | oT | roT | roS | iD | | 8 | 8 | 4 | 9 | 191 | 3 | 1 | 11 | +----------- Trapezidens
|
| +-- Lamellidens_corrianus **
--+ +-----+
| | +-- Lamellidens_savadiensis **
| |
| | +----- Lamellidens_lamellatus **
+--+ |
| +----- Lamellidens_jenkinsianus **
| |
| | +-- Lamellidens_marginalis **
+--+--+
| +-- Lamellidens_ferrugineus **
|
| +-- Lamellidens_mainwaringi__1 **
| |
+--+-- Lamellidens_mainwaringi__2 **
|
+-- Lamellidens_generosus **
Huang et al., 2019 [Publ 3843], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 8 | 8 | 7 | 9 | 193 | 3 | 1 | 9 | +----------------------- Trapezidens
|
--+ +-------------------- Lamellidens_corrianus **
| |
+--+ +----------------- Lamellidens_savadiensis **
| |
+--+ +-- Lamellidens_lamellatus **
| +-----------+
| | +-- Lamellidens_jenkinsianus **
+--+
| +----------- Lamellidens_generosus **
+--+
| +-------- Lamellidens_mainwaringi__2 **
+--+
| +----- Lamellidens_mainwaringi__1 **
+--+
| +-- Lamellidens_ferrugineus **
+--+
+-- Lamellidens_marginalis **
Bolotov et al., 2017 [Publ 3761], fig. S2 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 3 | 6 | 133 | 2 | 1 | 7 | +----------- Trapezidens
|
--+ +-------- Lamellidens_savadiensis **
| |
| +-------- Lamellidens_corrianus **
+--+
| +-- Lamellidens_marginalis **
| +--+
| | +-- Lamellidens_ferrugineus **
+--+
| +-- Lamellidens_jenkinsianus **
+--+
+-- Lamellidens_mainwaringi **
Bolotov et al., 2017 [Publ 3761], fig. S1 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 4 | 6 | 133 | 2 | 1 | 6 | +-------------- Trapezidens
|
--+ +----------- Lamellidens_corrianus **
| |
+--+ +-------- Lamellidens_savadiensis **
| |
+--+ +-- Lamellidens_jenkinsianus **
| +--+
| | +-- Lamellidens_mainwaringi **
+--+
| +-- Lamellidens_ferrugineus **
+--+
+-- Lamellidens_marginalis **
Konopleva et al., 2020 [Publ 4072], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 8 | 5 | 1 | 8 | +----------- Trapezidens
--+
| +-------- Lamellidens_savadiensis **
+--+
| +----- Lamellidens_generosus **
+--+
| +-- Lamellidens_mainwaringi **
+--+
+-- Lamellidens_ferrugineus **
Bolotov et al., 2022 [Publ 4250], fig. 8 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 45 | 5 | 1 | 6 | +----------- Trapezidens
--+
| +-------- Lamellidens_savadiensis **
+--+
| +----- Lamellidens_ferrugineus **
+--+
| +-- Lamellidens_generosus **
+--+
+-- Lamellidens_mainwaringi **
Bolotov et al., 2022 [Publ 4250], fig. S4 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 5 | 5 | 1 | 6 | +----------- Trapezidens
--+
| +-------- Lamellidens_savadiensis **
+--+
| +----- Lamellidens_ferrugineus **
+--+
| +-- Lamellidens_mainwaringi **
+--+
+-- Lamellidens_generosus **
Bolotov et al., 2020 [Publ 4100], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 113 | 4 | 1 | 8 | +----------- Trapezidens
--+
| +-------- Lamellidens_savadiensis **
+--+
| +----- Lamellidens_generosus **
+--+
| +-- Lamellidens_ferrugineus **
+--+
+-- Lamellidens_mainwaringi **
Bolotov et al., 2018 [Publ 3826], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 78 | 2 | 1 | 6 | +----------- Trapezidens
--+
| +-------- Lamellidens_savadiensis **
+--+
| +----- Lamellidens_ferrugineus **
+--+
| +-- Lamellidens_generosus **
+--+
+-- Lamellidens_mainwaringi **
Bolotov et al., 2020 [Publ 4100], fig. S1 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 107 | 4 | 1 | 8 | +----------- Trapezidens
--+
| +-------- Lamellidens_savadiensis **
+--+
| +----- Lamellidens_generosus **
+--+
| +-- Lamellidens_ferrugineus **
+--+
+-- Lamellidens_mainwaringi **
Konopleva et al., 2017 [Publ 3807], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 2 | 2 | 1 | 7 | +-------- Trapezidens
--+
| +----- Lamellidens_savadiensis **
+--+
| +-- Lamellidens_mainwaringi **
+--+
+-- Lamellidens_jenkinsianus **
Wu et al., 2021 [Publ 4366], fig. 2_I iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 88 | 1 | 1 | 7 | +-------- Trapezidens
--+
| +----- Lamellidens_savadiensis **
+--+
| +-- Lamellidens_generosus **
+--+
+-- Lamellidens_ferrugineus **
Konopleva et al., 2017 [Publ 3807], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 28 | 2 | 1 | 7 | +-------- Trapezidens
--+
| +----- Lamellidens_savadiensis **
+--+
| +-- Lamellidens_jenkinsianus **
+--+
+-- Lamellidens_mainwaringi **
Wu et al., 2021 [Publ 4366], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 88 | 1 | 1 | 7 | +-------- Trapezidens
--+
| +----- Lamellidens_savadiensis **
+--+
| +-- Lamellidens_ferrugineus **
+--+
+-- Lamellidens_generosus **
Sonowal et al., 2021 [Publ 4221], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 10 | 0 | 2 | 14 | 0 | 0 | 16 | +-------------------------- Lamellidens_mainwaringi **
--+
| +----------------------- Lamprotula
+--+
| +-------------------- Potomida
+--+
| +----------------- Lampsilis
+--+
| +-------------- Elliptio
+--+
| +----------- Scriptolamprotula
+--+
| +-------- Unio_pictorum
+--+
| +----- Anodonta
+--+
| +-- Unio_ravoisieri
+--+
+-- Lamellidens_corrianus **
Araujo et al., 2017 [Publ 3801], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 13 | 0 | 0 | -- | -- Lamellidens_savadiensis **
Pavan-Kumar et al., 2022 [Publ 4263], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 36 | 0 | 0 | -- | -- Lamellidens_jenkinsianus **
Pfeiffer et al., 2018 [Publ 3827], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 27 | 1 | 1 | -- | +-- Trapezidens
--+
+-- Lamellidens_savadiensis **
Whelan et al., 2011 [Publ 3380], fig. 1-ML iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 35 | 1 | 1 | -- | +-- Lamellidens_savadiensis **
--+
+-- Trapezidens
Pfeiffer & Graf, 2015 [Publ 3620], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 39 | 1 | 1 | -- | +-- Trapezidens
--+
+-- Lamellidens_savadiensis **
Pavan-Kumar et al., 2022 [Publ 4263], fig. S5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 38 | 0 | 0 | -- | -- Lamellidens_jenkinsianus **
Jeratthitikul et al., 2019 [Publ 3922], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 22 | 1 | 1 | -- | +-- Lamellidens_mainwaringi **
--+
+-- Trapezidens
Chen et al., 2026 [Publ 4833], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 26 | 1 | 1 | -- | -- Lamellidens_savadiensis **
Wu et al., 2023 [Publ 4339], fig. 4_S5-aa iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 94 | 0 | 0 | -- | -- Lamellidens_jenkinsianus **
Jadhav & Jamkhedkar, 2009 [Publ 3697], fig. 11 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 24 | 0 | 0 | -- | -- Lamellidens_corrianus **
Konopleva et al., 2023 [Publ 4269], fig. S1_4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 55 | 1 | 1 | -- | +-- Trapezidens
--+
+-- Lamellidens_savadiensis **
Imsong & Murali, 2023 [Publ 4370], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 2 | 0 | 0 | -- | -- Lamellidens_nongyangensis **
Wu et al., 2023 [Publ 4339], fig. 4_S5-nt iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 94 | 0 | 0 | -- | -- Lamellidens_jenkinsianus **
Wu et al., 2024 [Publ 4350], fig. 8a_S1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 90 | 0 | 0 | -- | -- Lamellidens_jenkinsianus **
Choi et al., 2020 [Publ 4169], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 9 | 0 | 0 | -- | -- Lamellidens_generosus **
Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 139 | 0 | 0 | -- | -- Lamellidens_jenkinsianus **
Whelan et al., 2011 [Publ 3380], fig. 1-MP iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 35 | 1 | 1 | -- | +-- Lamellidens_savadiensis **
--+
+-- Trapezidens
Zieritz et al., 2024 [Publ 4381], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 44 | 0 | 0 | -- | -- Lamellidens_corrianus **
Sonowal et al., 2021 [Publ 4221], fig. 3b iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 14 | 0 | 0 | -- | -- Lamellidens_mainwaringi **
Pfeiffer & Graf, 2013 [Publ 3531], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 30 | 0 | 0 | -- | -- Lamellidens_savadiensis **
Pfeiffer et al., 2019 [Publ 3937], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 56 | 1 | 1 | -- | +-- Trapezidens
--+
+-- Lamellidens_savadiensis **
Wu et al., 2024 [Publ 4350], fig. 8b_S2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 90 | 0 | 0 | -- | -- Lamellidens_jenkinsianus **
Pfeiffer et al., 2019 [Publ 3937], fig. 4-5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 56 | 1 | 1 | -- | +-- Trapezidens
--+
+-- Lamellidens_savadiensis **
Wu et al., 2024 [Publ 4388], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 52 | 0 | 0 | -- | -- Lamellidens_marginalis **
Sonowal et al., 2021 [Publ 4221], fig. 3a iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 14 | 0 | 0 | -- | -- Lamellidens_corrianus **
|
