MUSSELpdb | valid gen. Bineurus cladomics

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Page last updated
Thu 17 July 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

genus Bineurus Simpson, 1900: cladomics
5 Recent valid species (5 total species)

Cladomics Summary

number of species = 5 (5 total)
analyzed in 40 cladograms in 27 publications
maximum number of terminals in an analysis = 5 (mean = 2.68, median = 3)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 34
maximum number of included terminals = 5, (mean = 2.97 median = 3)
proportion of cladograms that support basic monophyly = 0.91

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination

NJ Tree

Jeratthitikul et al., 2024 [Publ 4387], fig. 1

** Top Monophyly Cladogram **

tT	bi	riS	oT	roT	roS	iD
4	4	3	5	54	31	9	0

  +-------------------------- Pseudodon              
  |                                                  
  |                       +-- Isannaia               
  |                 +-----+                          
--+                 |     +-- Pilsbryoconcha         
  |  +--------------+                                
  |  |              |  +----- Namkongnaia            
  |  |              +--+                             
  |  |                 |  +-- Lannanaia              
  +--+                 +--+                          
     |                    +-- Songkhlanaia           
     |                                               
     |  +-------------------- Sundadontina           
     +--+                                            
        |  +----------------- Nyeinchanconcha        
        +--+                                         
           |  +-------------- Thaiconcha             
           +--+                                      
              |  +----------- Bineurus_loeiensis **     
              +--+                                   
                 |  +-------- Bineurus_panhai **        
                 +--+                                
                    |  +----- Bineurus_anodontinum__2 **
                    +--+                             
                       |  +-- Bineurus_anodontinum__1 **
                       +--+                          
                          +-- Bineurus_exilis **

Jeratthitikul et al., 2025 [Publ 4423], fig. 1

** Top Sister-Group Cladogram **

tT	bi	riS	oT	roT	roS	iD
5	5	3	5	45	33	9	4

                          +-- Lannanaia           
  +-----------------------+                       
  |                       +-- Pilsbryoconcha      
  |                                               
--+  +----------------------- Songkhlanaia        
  |  |                                            
  |  |              +-------- Sundadontina        
  +--+  +-----------+                             
     |  |           |  +----- Isannaia            
     |  |           +--+                          
     |  |              |  +-- Namkongnaia         
     +--+              +--+                       
        |                 +-- Pseudodon           
        |                                         
        |  +----------------- Nyeinchanconcha     
        +--+                                      
           |  +-------------- Thaiconcha          
           +--+                                   
              |  +----------- Bineurus_loeiensis **  
              +--+                                
                 |  +-------- Bineurus_exilis **     
                 +--+                             
                    |  +----- Bineurus_mouhotii **   
                    +--+                          
                       |  +-- Bineurus_anodontinum **
                       +--+                       
                          +-- Bineurus_panhai **

Bolotov et al., 2023 [Publ 4312], fig. S2

tT	bi	riS	oT	roT	roS	iD
5	5	3	5	36	31	7	2

  +-------------------------- Songkhlanaia        
  |                                               
  |                       +-- Nyeinchanconcha     
--+  +--------------------+                       
  |  |                    +-- Sundadontina        
  |  |                                            
  +--+                    +-- Pseudodon           
     |  +-----------------+                       
     |  |                 +-- Pilsbryoconcha      
     +--+                                         
        |  +----------------- Namkongnaia         
        +--+                                      
           |  +-------------- Thaiconcha          
           +--+                                   
              |  +----------- Bineurus_loeiensis **  
              +--+                                
                 |  +-------- Bineurus_panhai **     
                 +--+                             
                    |  +----- Bineurus_anodontinum **
                    +--+                          
                       |  +-- Bineurus_exilis **     
                       +--+                       
                          +-- Bineurus_mouhotii **

Jeratthitikul & Sutcharit, 2023 [Publ 4289], fig. 1

tT	bi	riS	oT	roT	roS	iD
5	5	3	5	8	6	6	2

     +----------- Bineurus_mouhotii **   
  +--+                                
  |  |  +-------- Bineurus_loeiensis **  
  |  +--+                             
  |     |  +----- Bineurus_panhai **     
  |     +--+                          
  |        |  +-- Bineurus_anodontinum **
--+        +--+                       
  |           +-- Bineurus_exilis **     
  |                                   
  |        +----- Namkongnaia         
  |     +--+                          
  |     |  |  +-- Sundadontina        
  |     |  +--+                       
  +-----+     +-- Nyeinchanconcha     
        |                             
        |  +----- Pseudodon           
        +--+                          
           |  +-- Pilsbryoconcha      
           +--+                       
              +-- Thaiconcha

Bolotov et al., 2023 [Publ 4312], fig. 2

tT	bi	riS	oT	roT	roS	iD
4	4	2	4	35	30	7	3

  +-------------------------- Pilsbryoconcha      
  |                                               
--+  +----------------------- Songkhlanaia        
  |  |                                            
  +--+                    +-- Namkongnaia         
     |  +-----------------+                       
     |  |                 +-- Pseudodon           
     +--+                                         
        |  +----------------- Sundadontina        
        +--+                                      
           |  +-------------- Nyeinchanconcha     
           +--+                                   
              |  +----------- Thaiconcha          
              +--+                                
                 |  +-------- Bineurus_loeiensis **  
                 +--+                             
                    |  +----- Bineurus_mouhotii **   
                    +--+                          
                       |  +-- Bineurus_anodontinum **
                       +--+                       
                          +-- Bineurus_exilis **

Konopleva et al., 2023 [Publ 4262], fig. 8

tT	bi	riS	oT	roT	roS	iD
4	4	2	4	78	22	7	3

  +----------------------------- Pilsbryoconcha      
--+                                                  
  |  +-------------------------- Songkhlanaia        
  +--+                                               
     |  +----------------------- Namkongnaia         
     +--+                                            
        |  +-------------------- Pseudodon           
        +--+                                         
           |  +----------------- Sundadontina        
           +--+                                      
              |  +-------------- Nyeinchanconcha     
              +--+                                   
                 |  +----------- Thaiconcha          
                 +--+                                
                    |  +-------- Bineurus_loeiensis **  
                    +--+                             
                       |  +----- Bineurus_mouhotii **   
                       +--+                          
                          |  +-- Bineurus_anodontinum **
                          +--+                       
                             +-- Bineurus_exilis **

Konopleva et al., 2021 [Publ 4213], fig. 1

tT	bi	riS	oT	roT	roS	iD
4	4	2	4	23	15	6	3

                 +----- Pilsbryoconcha_exilis__2
  +--------------+                              
  |              |  +-- Pilsbryoconcha_exilis__1
  |              +--+                           
  |                 +-- Namkongnaia             
--+                                             
  |                 +-- Pseudodon               
  |  +--------------+                           
  |  |              +-- Sundadontina            
  +--+                                          
     |  +-------------- Nyeinchanconcha         
     +--+                                       
        |  +----------- Thaiconcha              
        +--+                                    
           |  +-------- Bineurus_loeiensis **      
           +--+                                 
              |  +----- Bineurus_mouhotii **       
              +--+                              
                 |  +-- Bineurus_anodontinum **    
                 +--+                           
                    +-- Bineurus_exilis **

Bolotov et al., 2023 [Publ 4312], fig. S1

tT	bi	riS	oT	roT	roS	iD
4	4	2	4	35	30	7	3

  +-------------------- Pilsbryoconcha      
  |                                         
  |           +-------- Songkhlanaia        
--+  +--------+                             
  |  |        |  +----- Sundadontina        
  |  |        +--+                          
  |  |           |  +-- Namkongnaia         
  +--+           +--+                       
     |              +-- Pseudodon           
     |                                      
     |  +-------------- Nyeinchanconcha     
     +--+                                   
        |  +----------- Thaiconcha          
        +--+                                
           |  +-------- Bineurus_loeiensis **  
           +--+                             
              |  +----- Bineurus_mouhotii **   
              +--+                          
                 |  +-- Bineurus_anodontinum **
                 +--+                       
                    +-- Bineurus_exilis **

Huang et al., 2019 [Publ 3843], fig. S4

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	196	9	5	4

  +-------- Nyeinchanconcha       
  |                               
  +-------- Thaiconcha            
  |                               
  +-------- Sundadontina_cumingii 
  |                               
  +-------- Sundadontina_tumida__1
--+                               
  +-------- Sundadontina_tumida__2
  |                               
  +-------- Pilsbryoconcha        
  |                               
  +-------- Pseudodon             
  |                               
  |  +----- Bineurus_loeiensis **    
  +--+                            
     |  +-- Bineurus_mouhotii **     
     +--+                         
        +-- Bineurus_exilis **

Bolotov et al., 2020 [Publ 4100], fig. S1

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	109	14	5	4

        +----- Bineurus_loeiensis **
  +-----+                        
  |     |  +-- Bineurus_exilis **   
  |     +--+                     
--+        +-- Bineurus_mouhotii ** 
  |                              
  |  +-------- Nyeinchanconcha   
  |  |                           
  +--+     +-- Thaiconcha        
     |  +--+                     
     |  |  +-- Pilsbryoconcha    
     +--+                        
        |  +-- Pseudodon         
        +--+                     
           +-- Sundadontina

Bolotov et al., 2017 [Publ 3761], fig. S1

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	136	9	5	4

  +----------------- Pilsbryoconcha       
  |                                       
  |              +-- Sundadontina_tumida  
--+  +-----------+                        
  |  |           +-- Pseudodon            
  |  |                                    
  +--+           +-- Nyeinchanconcha      
     |  +--------+                        
     |  |        +-- Sundadontina_cumingii
     +--+                                 
        |  +-------- Thaiconcha           
        +--+                              
           |  +----- Bineurus_exilis **      
           +--+                           
              |  +-- Bineurus_mouhotii **    
              +--+                        
                 +-- Bineurus_loeiensis **

Huang et al., 2019 [Publ 3843], fig. 2

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	196	9	5	4

  +----------------- Pilsbryoconcha    
  |                                    
--+              +-- Pseudodon         
  |  +-----------+                     
  |  |           +-- Sundadontina      
  +--+                                 
     |  +----------- Nyeinchanconcha   
     +--+                              
        |  +-------- Thaiconcha        
        +--+                           
           |  +----- Bineurus_loeiensis **
           +--+                        
              |  +-- Bineurus_mouhotii ** 
              +--+                     
                 +-- Bineurus_exilis **

Jeratthitikul et al., 2021 [Publ 4219], fig. 2

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	33	15	6	4

  +----------------- Pilsbryoconcha       
  |                                       
  |              +-- Namkongnaia          
  |        +-----+                        
  |        |     +-- Pseudodon            
--+  +-----+                              
  |  |     |  +----- Sundadontina_tumida  
  |  |     +--+                           
  |  |        |  +-- Sundadontina_brandti 
  |  |        +--+                        
  +--+           +-- Sundadontina_taskaevi
     |                                    
     |           +-- Nyeinchanconcha      
     |  +--------+                        
     |  |        +-- Sundadontina_cumingii
     +--+                                 
        |  +-------- Thaiconcha           
        +--+                              
           |  +----- Bineurus_mouhotii **    
           +--+                           
              |  +-- Bineurus_panhai **      
              +--+                        
                 +-- Bineurus_exilis **

Dai et al., 2023 [Publ 4285], fig. 3-ML

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	42	11	5	4

     +-------- Nyeinchanconcha   
  +--+                           
  |  |  +----- Bineurus_loeiensis **
  |  +--+                        
  |     |  +-- Bineurus_mouhotii ** 
  |     +--+                     
--+        +-- Bineurus_exilis **   
  |                              
  |        +-- Thaiconcha        
  |     +--+                     
  |     |  +-- Sundadontina      
  +-----+                        
        |  +-- Pilsbryoconcha    
        +--+                     
           +-- Pseudodon

Jeratthitikul et al., 2021 [Publ 4219], fig. 1

tT	bi	riS	oT	roT	roS	iD
3	3	0	3	33	15	6	4

  +-------------- Pilsbryoconcha   
  |                                
  |        +----- Sundadontina     
--+  +-----+                       
  |  |     |  +-- Namkongnaia      
  |  |     +--+                    
  +--+        +-- Pseudodon        
     |                             
     |  +-------- Nyeinchanconcha  
     |  |                          
     +--+  +----- Thaiconcha       
        |  |                       
        +--+  +-- Bineurus_panhai **  
           |  |                    
           +--+-- Bineurus_exilis **  
              |                    
              +-- Bineurus_mouhotii **

Konopleva et al., 2019 [Publ 3900], fig. 1

tT	bi	riS	oT	roT	roS	iD
3	8	0	3	49	5	2	8

  +-- Bineurus_mouhotii **         
  |                             
  +-- Pseudodon_vagulus         
  |                             
  +-- Pseudodon_mekongi         
  |                             
  +-- Pilsbryoconcha_exilis     
--+                             
  +-- Pilsbryoconcha_carinifera 
  |                             
  +-- Bineurus_loeiensis **        
  |                             
  +-- Bineurus_exilis **           
  |                             
  +-- Pseudodon_vondembuschianus

Bolotov et al., 2020 [Publ 4100], fig. 1

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	115	14	5	4

        +----- Bineurus_loeiensis **
  +-----+                        
  |     |  +-- Bineurus_exilis **   
  |     +--+                     
--+        +-- Bineurus_mouhotii ** 
  |                              
  |  +-------- Nyeinchanconcha   
  |  |                           
  +--+     +-- Thaiconcha        
     |  +--+                     
     |  |  +-- Pilsbryoconcha    
     +--+                        
        |  +-- Pseudodon         
        +--+                     
           +-- Sundadontina

Bolotov et al., 2017 [Publ 3764], fig. S1

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	122	9	5	4

        +-- Nyeinchanconcha      
  +-----+                        
  |     +-- Sundadontina_cumingii
  |                              
  |     +-- Sundadontina_tumida  
  +-----+                        
  |     +-- Pseudodon            
--+                              
  |     +-- Thaiconcha           
  +-----+                        
  |     +-- Pilsbryoconcha       
  |                              
  |  +----- Bineurus_exilis **      
  +--+                           
     |  +-- Bineurus_mouhotii **    
     +--+                        
        +-- Bineurus_loeiensis **

Bolotov et al., 2017 [Publ 3761], fig. S2

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	136	10	5	4

  +-------- Thaiconcha        
--+                           
  |  +----- Bineurus_exilis **   
  +--+                        
     |  +-- Bineurus_mouhotii ** 
     +--+                     
        +-- Bineurus_loeiensis **

Huang et al., 2019 [Publ 3843], fig. S3

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	194	9	5	4

  +----------------- Pilsbryoconcha       
  |                                       
  |              +-- Sundadontina_tumida  
--+  +-----------+                        
  |  |           +-- Pseudodon            
  |  |                                    
  +--+           +-- Nyeinchanconcha      
     |  +--------+                        
     |  |        +-- Sundadontina_cumingii
     +--+                                 
        |  +-------- Thaiconcha           
        +--+                              
           |  +----- Bineurus_loeiensis **   
           +--+                           
              |  +-- Bineurus_mouhotii **    
              +--+                        
                 +-- Bineurus_exilis **

Dai et al., 2023 [Publ 4285], fig. 3-BI

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	42	11	5	4

           +-- Thaiconcha        
  +--------+                     
  |        +-- Sundadontina      
  |                              
  |        +-- Pilsbryoconcha    
--+--------+                     
  |        +-- Pseudodon         
  |                              
  |  +-------- Nyeinchanconcha   
  +--+                           
     |  +----- Bineurus_loeiensis **
     +--+                        
        |  +-- Bineurus_mouhotii ** 
        +--+                     
           +-- Bineurus_exilis **

Bolotov et al., 2018 [Publ 3826], fig. 2

tT	bi	riS	oT	roT	roS	iD
3	8	0	3	80	5	2	8

  +-- Bineurus_mouhotii **         
  |                             
  +-- Pseudodon_vagulus         
  |                             
  +-- Pseudodon_mekongi         
  |                             
  +-- Pilsbryoconcha_exilis     
--+                             
  +-- Pilsbryoconcha_carinifera 
  |                             
  +-- Bineurus_loeiensis **        
  |                             
  +-- Bineurus_exilis **           
  |                             
  +-- Pseudodon_vondembuschianus

Zieritz et al., 2021 [Publ 4218], fig. 5

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	30	5	4	--

  +-------------- Pilsbryoconcha      
--+                                   
  |  +----------- Pseudodon           
  +--+                                
     |  +-------- Sundadontina        
     +--+                             
        |  +----- Thaiconcha          
        +--+                          
           |  +-- Bineurus_loeiensis **  
           +--+                       
              +-- Bineurus_anodontinum **

Liu et al., 2024 [Publ 4380], fig. 5

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	35	4	3	--

  +----------- Pilsbryoconcha      
--+                                
  |  +-------- Pseudodon           
  +--+                             
     |  +----- Thaiconcha          
     +--+                          
        |  +-- Bineurus_anodontinum **
        +--+                       
           +-- Bineurus_loeiensis **

Dai et al., 2024 [Publ 4389], fig. 3

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	45	9	5	--

  +----------------- Pilsbryoconcha      
--+                                      
  |  +-------------- Pseudodon           
  +--+                                   
     |  +----------- Sundadontina        
     +--+                                
        |  +-------- Nyeinchanconcha     
        +--+                             
           |  +----- Thaiconcha          
           +--+                          
              |  +-- Bineurus_loeiensis **  
              +--+                       
                 +-- Bineurus_anodontinum **

Wu et al., 2024 [Publ 4350], fig. 5c

tT	bi	riS	oT	roT	roS	iD
2	5	0	2	35	4	3	6

  +-------- Pilsbryoconcha      
  |                             
--+  +----- Bineurus_anodontinum **
  |  |                          
  +--+----- Bineurus_loeiensis **  
     |                          
     |  +-- Thaiconcha          
     +--+                       
        +-- Pseudodon

Dai et al., 2025 [Publ 4441], fig. 1

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	46	9	5	--

  +----------------- Pilsbryoconcha    
--+                                    
  |  +-------------- Pseudodon         
  +--+                                 
     |  +----------- Sundadontina      
     +--+                              
        |  +-------- Nyeinchanconcha   
        +--+                           
           |  +----- Thaiconcha        
           +--+                        
              |  +-- Bineurus_loeiensis **
              +--+                     
                 +-- Bineurus_mouhotii **

Wu et al., 2024 [Publ 4350], fig. 6-7

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	35	4	3	--

  +----------- Pilsbryoconcha      
--+                                
  |  +-------- Pseudodon           
  +--+                             
     |  +----- Thaiconcha          
     +--+                          
        |  +-- Bineurus_anodontinum **
        +--+                       
           +-- Bineurus_loeiensis **

Wu et al., 2024 [Publ 4350], fig. 5a

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	35	4	3	--

  +----------- Pilsbryoconcha      
--+                                
  |  +-------- Pseudodon           
  +--+                             
     |  +----- Thaiconcha          
     +--+                          
        |  +-- Bineurus_anodontinum **
        +--+                       
           +-- Bineurus_loeiensis **

Wu et al., 2024 [Publ 4388], fig. 5

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	36	4	3	--

  +----------- Pilsbryoconcha      
--+                                
  |  +-------- Pseudodon           
  +--+                             
     |  +----- Thaiconcha          
     +--+                          
        |  +-- Bineurus_anodontinum **
        +--+                       
           +-- Bineurus_loeiensis **

Wu et al., 2024 [Publ 4350], fig. 5b

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	35	4	3	--

  +-------- Pilsbryoconcha      
  |                             
--+     +-- Bineurus_anodontinum **
  |  +--+                       
  |  |  +-- Bineurus_loeiensis **  
  +--+                          
     |  +-- Thaiconcha          
     +--+                       
        +-- Pseudodon

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	74	13	6	--

        +----- Pilsbryoconcha   
  +-----+                       
  |     |  +-- Namkongnaia      
  |     +--+                    
  |        +-- Pseudodon        
--+                             
  |        +-- Nyeinchanconcha  
  |  +-----+                    
  |  |     +-- Sundadontina     
  +--+                          
     |  +----- Thaiconcha       
     +--+                       
        |  +-- Bineurus_panhai **  
        +--+                    
           +-- Bineurus_mouhotii **

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	74	13	6	--

        +----- Pilsbryoconcha   
  +-----+                       
  |     |  +-- Namkongnaia      
  |     +--+                    
  |        +-- Pseudodon        
--+                             
  |        +-- Nyeinchanconcha  
  |  +-----+                    
  |  |     +-- Sundadontina     
  +--+                          
     |  +----- Thaiconcha       
     +--+                       
        |  +-- Bineurus_panhai **  
        +--+                    
           +-- Bineurus_mouhotii **

Zieritz et al., 2024 [Publ 4381], fig. 2

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	44	6	4	--

           +-- Pilsbryoconcha      
  +--------+                       
  |        +-- Pseudodon           
--+                                
  |  +-------- Sundadontina        
  +--+                             
     |  +----- Thaiconcha          
     +--+                          
        |  +-- Bineurus_loeiensis **  
        +--+                       
           +-- Bineurus_anodontinum **

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	56	3	3	--

  +-------- Bineurus_anodontinum **
--+                             
  |  +----- Pilsbryoconcha      
  +--+                          
     |  +-- Pseudodon           
     +--+                       
        +-- Sundadontina

Pfeiffer et al., 2019 [Publ 3937], fig. 6

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	56	3	3	--

  +-------- Pilsbryoconcha      
--+                             
  |  +----- Bineurus_anodontinum **
  +--+                          
     |  +-- Pseudodon           
     +--+                       
        +-- Sundadontina

Konopleva et al., 2017 [Publ 3807], fig. 6

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	30	5	4	--

  +-------- Sundadontina   
  |                        
--+     +-- Nyeinchanconcha
  |  +--+                  
  |  |  +-- Bineurus_exilis **
  +--+                     
     |  +-- Pilsbryoconcha 
     +--+                  
        +-- Pseudodon

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	79	4	3	--

  +-------- Pilsbryoconcha 
--+                        
  |  +----- Pseudodon      
  +--+                     
     |  +-- Bineurus_exilis **
     +--+                  
        +-- Nyeinchanconcha

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	39	5	4	--

  +----------- Pilsbryoconcha 
--+                           
  |  +-------- Pseudodon      
  +--+                        
     |  +----- Sundadontina   
     +--+                     
        |  +-- Nyeinchanconcha
        +--+                  
           +-- Bineurus_exilis **

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	120	5	4	--

  +-------- Pilsbryoconcha   
  |                          
--+  +----- Pseudodon        
  |  |                       
  +--+  +-- Nyeinchanconcha  
     |  |                    
     +--+-- Bineurus_mouhotii **
        |                    
        +-- Sundadontina

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