The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)
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Bivalvia | Unionida | Unionoidea | Unionidae | Gonideinae | Gonideini genus Obovalis Simpson, 1900: cladomics 1 Recent valid species (1 total species) Cladomics Summarynumber of species = 1 (1 total) analyzed in 19 cladograms in 12 publications maximum number of terminals in an analysis =
1 (mean = 1, median = 1)
Sano et al., 2020 [Publ 4138], fig. 1 ** Top Monophyly Cladogram ** | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 2 | 0 | 2 | 33 | 3 | 3 | -- |
+-------- Sinosolenaia
|
--+ +----- Inversidens
| |
+--+ +-- Obovalis_omiensis__1 **
| |
+--+-- Obovalis_omiensis__2 **
|
+-- Ptychorhynchus
Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14 ** Top Sister-Group Cladogram ** | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 631 | 20 | 12 | -- |
+----------------- Parvasolenaia
|
--+ +-------------- Pseudopostolata
| |
+--+ +----------- Koreosolenaia
| |
+--+ +-- Cosmopseudodon
| +-----+
| | +-- Guiunio
+--+
| +----- Ptychorhynchus
+--+
| +-- Obovalis_omiensis **
+--+
+-- Postolata
Wu et al., 2024 [Publ 4388], fig. 5 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 35 | 8 | 8 | -- |
+-------------------- Sinosolenaia
|
--+ +-- Gonidea
| +--------------+
| | +-- Microcondylaea
+--+
| +-------------- Koreosolenaia
+--+
| +----------- Parvasolenaia
+--+
| +-------- Postolata
+--+
| +----- Cosmopseudodon
+--+
| +-- Ptychorhynchus
+--+
+-- Obovalis_omiensis **
Kano et al., 2019 [Publ 4194], fig. 6 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 9 | 1 | 1 | -- |
Wu et al., 2024 [Publ 4388], fig. 4 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 8 | 3 | 3 | -- |
+-- Cosmopseudodon
+--+
| +-- Ptychorhynchus
--+
| +-- Postolata
+--+
+-- Obovalis_omiensis **
Sano et al., 2020 [Publ 4138], fig. 3 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 67 | 2 | 2 | -- |
+----- Inversidens
--+
| +-- Obovalis_omiensis **
+--+
+-- Ptychorhynchus
Dai et al., 2024 [Publ 4389], fig. 3 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 45 | 14 | 10 | -- |
+-------------------- Sinosolenaia
|
| +----- Gonidea
--+ +-----------+
| | | +-- Leguminaia
| | +--+
+--+ +-- Microcondylaea
|
| +-------------- Pseudopostolata
| |
+--+ +-- Parvasolenaia
| +--------+
| | +-- Koreosolenaia
+--+
| +-------- Postolata
+--+
| +----- Ptychorhynchus
+--+
| +-- Cosmopseudodon
+--+
+-- Obovalis_omiensis **
Sano et al., 2017 [Publ 3799], fig. 1 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 20 | 1 | 1 | -- |
Dai et al., 2023 [Publ 4285], fig. 3-ML | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 44 | 13 | 9 | -- |
+-------------- Sinosolenaia
|
| +----- Gonidea
--+ +-----+
| | | +-- Leguminaia
| | +--+
| | +-- Microcondylaea
+--+
| +-- Koreosolenaia
| +-----+
| | +-- Parvasolenaia
+--+
| +----- Postolata
+--+
| +-- Ptychorhynchus
+--+
+-- Obovalis_omiensis **
Dai et al., 2023 [Publ 4285], fig. 3-BI | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 44 | 13 | 9 | -- |
+-------------- Sinosolenaia
|
| +----- Gonidea
--+ +-----+
| | | +-- Leguminaia
| | +--+
| | +-- Microcondylaea
+--+
| +-- Koreosolenaia
| +-----+
| | +-- Parvasolenaia
+--+
| +----- Postolata
+--+
| +-- Ptychorhynchus
+--+
+-- Obovalis_omiensis **
Mabuchi & Nishida, 2020 [Publ 4120], fig. 1 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 11 | 1 | 1 | -- |
Liu et al., 2024 [Publ 4380], fig. 5 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 35 | 8 | 7 | -- |
+-------------- Sinosolenaia
|
| +-- Gonidea
--+ +--------+
| | +-- Microcondylaea
| |
+--+ +-- Koreosolenaia
| +-----+
| | +-- Parvasolenaia
+--+
| +----- Postolata
+--+
| +-- Obovalis_omiensis **
+--+
+-- Ptychorhynchus
Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 121 | 11 | 8 | -- |
+-------- Sinosolenaia
+--+
| | +----- Gonidea
| +--+
| | +-- Leguminaia
--+ +--+
| +-- Microcondylaea
|
| +-------- Inversidens
| |
+--+ +-- Obovalis_omiensis **
| +--+
| | +-- Ptychorhynchus
+--+
| +-- Parvasolenaia
+--+
+-- Koreosolenaia
Sano et al., 2020 [Publ 4138], fig. 2 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 67 | 2 | 2 | -- |
+-- Ptychorhynchus
--+
+-- Obovalis_omiensis **
Dai et al., 2025 [Publ 4441], fig. 1 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 46 | 15 | 11 | -- |
+-------------------- Sinosolenaia
|
| +----- Gonidea
--+ +-----------+
| | | +-- Leguminaia
| | +--+
+--+ +-- Microcondylaea
|
| +-------------- Pseudopostolata
| |
+--+ +-- Koreosolenaia
| +--------+
| | +-- Parvasolenaia
+--+
| +-------- Postolata
| |
+--+ +-- Guiunio
| +--+
| | +-- Cosmopseudodon
+--+
| +-- Obovalis_omiensis **
+--+
+-- Ptychorhynchus
Sano et al., 2017 [Publ 3799], fig. 2 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 32 | 3 | 3 | -- |
+-- Obovalis_omiensis **
--+
+-- Ptychorhynchus
Sano et al., 2020 [Publ 4138], fig. 4 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 67 | 2 | 2 | -- |
+-- Obovalis_omiensis **
--+
+-- Ptychorhynchus
Liu et al., 2024 [Publ 4380], fig. 4 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 11 | 9 | 6 | -- |
+-------------- Sinosolenaia_carinata
|
| +----- Inversidens
--+ +-----+
| | | +-- Gonidea
| | +--+
+--+ +-- Leguminaia
|
| +-------- Sinosolenaia_oleivora
+--+
| +----- Postolata
+--+
| +-- Obovalis_omiensis **
+--+
+-- Ptychorhynchus
Chen et al., 2025 [Publ 4663], fig. 1 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 47 | 16 | 12 | -- |
+----------------------- Sinosolenaia
|
| +----- Gonidea
--+ +--------------+
| | | +-- Leguminaia
| | +--+
+--+ +-- Microcondylaea
|
| +----------------- Pseudopostolata
| |
+--+ +-- Koreosolenaia
| +-----------+
| | +-- Parvasolenaia
+--+
| +----------- Postolata
| |
+--+ +-- Guiunio
| +-----+
| | +-- Cosmopseudodon
+--+
| +----- Obovalis_omiensis **
+--+
| +-- Tenuiunio
+--+
+-- Ptychorhynchus
|