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The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

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Publication Lopes-Lima et al. (2017): attributed nominal species
publication summary | cladograms

Complete Citation

Lopes-Lima, M., E. Froufe, V.T. Do, M. Ghamizi, K.E. Mock et al. 2017. Phylogeny of the most species-rich freshwater bivalve family (Bivalvia: Unionida: Unionidae): Defining modern subfamilies and tribes. Molecular Phylogenetics and Evolution 106: 174-191. Available electronically.

Attributed Nominal Species

Anodontites elongata Whelan et al., 2011
‘Swainson’ of Authors.
Whelan et al., 2011. MPE 61: 506.
Graf et al., 2015. MPE 85: 3.
Bolotov et al., 2016. MPE 103: 108.
Lopes-Lima et al., 2017. MPE 106: 178.
Bolotov et al., 2017. Sci. Repts. 7: Table S1.
Bolotov et al., 2017. Sci. Repts. 7: Table S1.
Konopleva et al., 2017. Systemat. Biodiv. 15(3): 208.
Combosch et al., 2017. MPE 107: 195, Table 1.
Bolotov et al., 2018. Sci. Repts. 8: Table S1.
Huang et al., 2018. MPE 120: 302.
Lopes-Lima et al., 2018. MPE 127: 103.
Huang et al., 2019. MPE 130: Table S4.
Bolotov et al., 2019. Sci. Repts. 9: Table S1.
Pfeiffer et al., 2019. MPE 137: 116.
Bolotov et al., 2020. Sci. Repts. 10: Table S1.
Sano et al., 2020. Plankton Benthos Res. 15(2): 101.
Wu et al., 2020. Invert. Syst. 34: 195.
Wu et al., 2021. J. Lake Sci. 33(6): attached Table 1.
Chen et al., 2025. Zoosyst. Evol. 101(4): Table 1.
(misidentified reference to Anodon elongatus Swainson, 1823)

Chamberlainia hainesiana Kongim et al., 2015
‘Lea’ of Authors.
Kongim et al., 2015. ZooKeys 514: 98, fig. 2A.
Lopes-Lima et al., 2017. MPE 106: 177.
Bolotov et al., 2017. Sci. Repts. 7: Table S1.
Bolotov et al., 2017. Sci. Repts. 7: Table S1.
Froufe et al., 2019. Heredity 124: 185.
Huang et al., 2019. MPE 130: Table S4.
Chen et al., 2020. mtDNA B 5(3): 2800.
Teiga-Teixeira et al., 2020. mtDNA B 5(3): 2978.
Zieritz et al., 2020. Biodivers. Conserv. 29: 2244.
Bolotov et al., 2020. Sci. Repts. 10: Table S1.
Lopes-Lima et al., 2020. MPE 146: Table 1.
Zieritz et al., 2021. Zool. J. Linn. Soc. 193(3): Table 1.
Zieritz et al., 2021. Aquatic Conserv.: Mar. Freshw. Ecosyst. 31: 3176.
Jeratthitikul et al., 2021. Eur. J. Taxonomy 775: 125.
Wu et al., 2021. ZooKeys 1054: 87.
Wu et al., 2021. J. Lake Sci. 33(6): attached Table 1.
Wu et al., 2022. Zool. Scripta 51: Table S2.
Pavan-Kumar et al., 2022. Mol. Biol. Rep. 49: Table S1.
Jeratthitikul et al., 2022. Sci. Repts. 12: Table S1.
Dai et al., 2023. Invert. Syst. 37(2): Tables 1-2.
Konopleva et al., 2023. Zoosyst. Evol. 99(1): Table S1.
Wu et al., 2023. Org Divers Evol (2023): Table 2.
Dai et al., 2024. Zool. Scripta 53: Table 1.
Dai et al., 2024. Invert. Syst. 38(6): Table 3.
Dai et al., 2024. Zoosyst. Evol. 100(4): Table 2.
Chen et al., 2025. Zoosyst. Evol. 101(4): Table S3.
Dai et al., 2025. Zoosyst. Evol. 101(2): Table 1.
Dai et al., 2025. Zoosyst. Evol. 101(2): Table 2.
Chen et al., 2025. Cathaica 1(7): Table 1.
Klymus et al., 2025. mtDNA B 10(6): Table 1.
Campbell & Lydeard, 2026. Malacologia 68(1/2): 289. [in part]
(misidentified reference to Unio hainesianus Lea, 1856)

Contradens semmelinki fultoni Lopes-Lima et al., 2017
‘Haas’ Lopes-Lima et al., 2017. MPE 106: 177.
(misidentified reference to Contradens semmelincki [sic] fultoni Haas, 1930)

Cuneopsis heudei Ouyang et al., 2015
‘Heude’ of Authors.
Ouyang et al., 2015. J. Agri. Biotech. 23(6): Table 1.
Liu et al., 2017. PLoS One 12(12 ), e0189737: Table S4.
Lopes-Lima et al., 2017. MPE 106: Table 2.
Bolotov et al., 2017. Sci. Repts. 7: Table S1.
Bolotov et al., 2017. Sci. Repts. 7: Table S1.
Wu et al., 2018. PLoS One 13(8), e0200956: 5.
Huang et al., 2019. MPE 130: Tables S1, S4. [in part]
Lopes-Lima et al., 2020. MPE 146: Table 1. [in part]
Wu et al., 2021. J. Lake Sci. 33(6): attached Table 1. [in part]
Wu et al., 2022. Zool. Scripta 51: Table S1. [in part]
Jeratthitikul et al., 2022. Sci. Repts. 12: Table S1.
Konopleva et al., 2023. Zoosyst. Evol. 99(1): Table S1.
Chen et al., 2023. ZooKeys 1173: Table 1.
Wu et al., 2023. ZooKeys 1166: Table 1.
Liu et al., 2023. ZooKeys 1179: Table 1.
Chen et al., 2025. Zoosyst. Evol. 101(4): Table S2. [in part]
Dai et al., 2025. Zoosyst. Evol. 101(2): Table 1. [in part]
Chen et al., 2025. Zoosyst. Evol. 101(4): Table 1.
(misidentified reference to Unio heudei Heude, 1874)

Ensidens sagittarius Pfeiffer & Graf, 2015
‘Lea’ of Authors.
Pfeiffer & Graf, 2015. Zool. J. Linn. Soc. 175(2): 310.
Konopleva et al., 2017. Systemat. Biodiv. 15(3): 207.
Lopes-Lima et al., 2017. MPE 106: 177.
Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘Ensidens aff. sagittarius sp. 1].
Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘Ensidens aff. sagittarius sp. 1].
Bolotov et al., 2018. Sci. Repts. 8: Table S1 [as ‘Ensidens aff. sagittarius sp. 1].
Konopleva et al., 2019. Sci. Repts. 9: Table S1 [as ‘E. aff. sagittarius sp. 1].
Huang et al., 2019. MPE 130: Table S4 [as ‘Ensidens aff. sagittarius sp. 1].
Zieritz et al., 2020. Biodivers. Conserv. 29: 2242.
Lopes-Lima et al., 2020. MPE 146: Table 1.
Wu et al., 2021. J. Lake Sci. 33(6): attached Table 1.
(misidentified reference to Unio sagittarius Lea, 1856)

Hyriopsis bialata Lopes-Lima et al., 2017
‘Simpson’ of Authors.
Lopes-Lima et al., 2017. MPE 106: 177.
Muanta et al., 2019. J. Moll. Studies 85(2): 225.
Lopes-Lima et al., 2020. MPE 146: Table 1.
(misidentified reference to Hyriopsis bialatus Simpson, 1900)

Lasmigona compressa Graf & Ó Foighil, 2000
‘Lea’ of Authors.
Graf & Ó Foighil, 2000. J. Moll. Studies 66: 160.
Graf & Cummings, 2006. Zool. J. Linn. Soc. 148: 348.
Hoeh et al., 2009. Malacologia 51(2): 310.
Jadhav & Jamkhedkar, 2009. Res. J. BioTech. 4(1): 43.
Whelan et al., 2011. MPE 61: 506.
Lopes-Lima et al., 2017. MPE 106: 177.
Bolotov et al., 2017. Sci. Repts. 7: Table S1.
Bolotov et al., 2017. Sci. Repts. 7: Table S1.
Bolotov et al., 2018. Sci. Repts. 8: Table S1.
Huang et al., 2019. MPE 130: Table S4.
Lopes-Lima et al., 2020. MPE 146: Table 1.
Wu et al., 2021. J. Lake Sci. 33(6): attached Table 1.
Jeratthitikul et al., 2022. Sci. Repts. 12: Table S1.
Wu et al., 2023. ZooKeys 1166: 263, Table 1.
Liu et al., 2023. ZooKeys 1179: Table 1.
Dai et al., 2025. Zoosyst. Evol. 101(2): Table 1. [in part]
Chen et al., 2025. Zoosyst. Evol. 101(4): Table 1.
(misidentified reference to Symphynota compressa Lea, 1829)

Pilsbryoconcha exilis Lopes-Lima et al., 2017
‘Lea’ Lopes-Lima et al., 2017. MPE 106: 177. [in part]
(misidentified reference to Anodonta exilis Lea, 1838)

Pilsbryoconcha exilis Graf, 2002
‘Lea’ of Authors.
Graf, 2002. J. Moll. Studies 68: 66.
Graf & Cummings, 2006. Zool. J. Linn. Soc. 148: 372.
Hoeh et al., 2009. Malacologia 51(2): 310.
Lopes-Lima et al., 2017. MPE 106: 177. [in part]
(reference to Anodonta exilis Lea, 1838)

Pseudodon cambodjensis Pfeiffer & Graf, 2013
‘Petit’ of Authors.
Pfeiffer & Graf, 2013. J. Moll. Studies 79: 251.
Pfeiffer & Graf, 2015. Zool. J. Linn. Soc. 175(2): 310.
Zieritz et al., 2016. Sci. Total Environ. 571: 1071.
Bolotov et al., 2017. Sci. Repts. 7: Table S1.
Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘Monodontina cambodjensis’].
Lopes-Lima et al., 2017. MPE 106: 177.
Konopleva et al., 2017. Systemat. Biodiv. 15(3): 207.
Bolotov et al., 2018. Sci. Repts. 8: Table S1 [as ‘Monodontina cambodjensis’].
Razak et al., 2018. Conserv. Genet. Resour. 11: 459.
Huang et al., 2019. MPE 130: Table S4 [as ‘Monodontina cambodjensis’].
Konopleva et al., 2019. Sci. Repts. 9: Table S1 [as ‘Monodontina cambodjensis’].
Bolotov et al., 2020. Sci. Repts. 10: Table S1 [as ‘Monodontina cambodiensis’ (sic)].
Konopleva et al., 2021. Sci. Repts. 11: Table S1 [as ‘Monodontina cambodiensis’ (sic)].
Jeratthitikul et al., 2021. Eur. J. Taxonomy 775: 123 [as ‘Monodontina cambodiensis’ (sic)].
Pfeiffer et al., 2021. Invert. Syst. 35: 453 [as ‘Monodontina cambodjensis’].
Wu et al., 2021. ZooKeys 1054: 87 [as ‘Monodontina cambodjensis’].
Wu et al., 2021. J. Lake Sci. 33(6): attached Table 1.
Konopleva et al., 2023. Diversity 15(1), 10: Table S2 [as ‘Monodontina cambodjensis’].
Dai et al., 2023. Invert. Syst. 37(2): Table 1 [as ‘Monodontina cambodjensis’].
Dai et al., 2024. Zoosyst. Evol. 100(4): Table 2.
Dai et al., 2025. Zoosyst. Evol. 101(2): Table 1.
(reference to Monocondylea cambodjensis Petit de la Saussaye, 1865)
Type(s): SEQUENCED SPECIMEN UMMZ_304350, Pursat River, just downstream (north) of Highway 5 bridge, Pursat City, Cambodia.

Pseudodon cumingii Pfeiffer & Graf, 2015
‘Lea’ of Authors.
Pfeiffer & Graf, 2015. Zool. J. Linn. Soc. 175(2): 310.
Lopes-Lima et al., 2017. MPE 106: 177.
Bolotov et al., 2017. Sci. Repts. 7: Table S1.
Konopleva et al., 2017. Systemat. Biodiv. 15(3): 207.
Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘”Bineurus” cumingii’].
Huang et al., 2019. MPE 130: Table S4 [as ‘”Bineurus” cumingii’].
Lopes-Lima et al., 2020. MPE 146: Table 1.
(misidentified reference to Anodonta cumingii Lea, 1851)
Type(s): SPECIMEN UMMZ_304648, Nam Phiat, Phonethong, ca. 2 km from confluence, Bolikhamxai Province, Laos. 18.08389, 104.97806.

Pseudodon mouhotii Pfeiffer & Graf, 2015
‘Lea’ of Authors.
Pfeiffer & Graf, 2015. Zool. J. Linn. Soc. 175(2): 310.
Lopes-Lima et al., 2017. MPE 106: 177.
Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘Pseudodon aff. mouhotii sp. 3’].
Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘Bineurus aff. mouhotii sp. 3’].
Konopleva et al., 2017. Systemat. Biodiv. 15(3): 207.
Bolotov et al., 2018. Sci. Repts. 8: Table S1 [as ‘Bineurus aff. mouhotii sp. 3’].
Konopleva et al., 2019. Sci. Repts. 9: Table S1 [as ‘Bineurus aff. mouhotii sp. 3’].
Huang et al., 2019. MPE 130: Table S4 [as ‘Bineurus aff. mouhotii sp. 3’].
(misidentified reference to Monocondyloea mouhotii Lea, 1863)

Schistodesmus lampreyanus Huang et al., 2002
‘Baird & Adams’ of Authors.
Huang et al., 2002. J. Moll. Studies 68: Table 1.
Zhang et al., 2004. Acta Hydrobiol. Sinica 28(3): fig. 1.
Zhou et al., 2007. Acta Zool. Sinica 53(6): Table 1.
Szumowski et al., 2012. Amer. Malac. Bull. 30(1): Table S3.
Ouyang et al., 2015. J. Agri. Biotech. 23(6): Table 1.
Liu et al., 2017. PLoS One 12(12 ), e0189737: Table S4.
Lopes-Lima et al., 2017. MPE 106: Table 2.
Bolotov et al., 2017. Sci. Repts. 7: Table S1.
Bolotov et al., 2017. Sci. Repts. 7: Table S1.
Sano et al., 2017. J. Wat. Res. Prot. 9: Table 2.
Wu et al., 2018. PLoS One 13(8), e0200956: Table 1.
Wu et al., 2019. ZooKeys 812: 29.
Fukata & Iigo, 2019. mtDNA B 4(2): fig. 1.
Huang et al., 2019. MPE 130: Tables S1, S4. [in part]
Fukata & Iigo, 2020. mtDNA B 5(2): fig. 1.
Wu et al., 2020. Invert. Syst. 34: Table S2.
Wu et al., 2020. Malacologia 63(1): Table 1.
Lopes-Lima et al., 2020. MPE 146: Table 1.
Choi et al., 2020. Sci. Repts. 10: Table S5.
Sano et al., 2020. Plankton Benthos Res. 15(2): Table 1.
Kim et al., 2021. mtDNA B 6(1): fig. 1.
Wu et al., 2021. J. Lake Sci. 33(6): attached Tables 1-2.
Wu et al., 2022. Zool. Scripta 51: Tables S1-S2.
Pavan-Kumar et al., 2022. Mol. Biol. Rep. 49: Table S1.
Jeratthitikul et al., 2022. Sci. Repts. 12: Table S1.
Dai et al., 2023. Invert. Syst. 37(2): Table 2.
Wu et al., 2023. Org Divers Evol (2023): Table 2.
Wu et al., 2023. ZooKeys 1166: Table 1.
Liu et al., 2023. ZooKeys 1179: Table 1.
Bogan et al., 2023. Diversity 15(6), 710: Table 1.
Chen et al., 2023. ZooKeys 1173: Table 1.
Dai et al., 2024. Zool. Scripta 53: Tables 1, S2.
Wu et al., 2024. J. Zool. Syst. Evol. Res. 2024, 1499508: Table 2.
Wu et al., 2024. Zool. Scripta 53(5): Tables S2-S3.
Dai et al., 2024. Invert. Syst. 38(6): Tables 2-3.
Hou et al., 2024. Invert. Syst. 38, IS24071: Table 2.
Hou et al., 2025. Zoosyst. Evol. 101(1): Table S2.
Hou et al., 2025. Zoosyst. Evol. 101(1): Table S2.
Dai et al., 2025. Zoosyst. Evol. 101(2): Tables 1-2.
Chen et al., 2025. Zoosyst. Evol. 101(4): Table S3.
Chen et al., 2025. Zoosyst. Evol. 101(4): Table 1.
Campbell & Lydeard, 2026. Malacologia 68(1/2): 294. [in part]
(misidentified reference to Unio (Dysnomia) lampreyanus Baird & Adams, 1867)

Sinanodonta woodiana Lopes-Lima et al., 2017
‘Lea’ of Authors.
Vikhrev et al., 2017. Trop. Conserv. Sci. 10: Table 1 [as ‘lineage H’]. [in part]
Lopes-Lima et al., 2017. MPE 106: Table 2.
Bespalaya et al., 2018. Limnologica 69: Table 1 [as ‘lineage H’]. [in part]
Chen et al., 2023. ZooKeys 1173: 133, Table 1.
Chen et al., 2025. Zoosyst. Evol. 101(4): Table 1.
(misidentified reference to Symphynota woodiana Lea, 1834)

Solenaia triangularis Ouyang et al., 2015
‘Heude’ of Authors.
Ouyang et al., 2015. J. Agri. Biotech. 23(6): Table 1.
Lopes-Lima et al., 2017. MPE 106: 177.
Wu et al., 2018. PLoS One 13(8), e0200956: 7. [in part]
(misidentified reference to Mycetopus triangularis Heude, 1885)

Trapezoideus exolescens Pfeiffer & Graf, 2013
‘Gould’ of Authors.
Brandt, 1974. Arch. Moll. 105: 299.
Pfeiffer & Graf, 2013. J. Moll. Studies 79: 251.
Pfeiffer & Graf, 2015. Zool. J. Linn. Soc. 175(2): [4].
Lopes-Lima et al., 2017. MPE 106: 177.
(misidentified reference to Unio exolescens Gould, 1843)

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