The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)
FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!
Publication Bolotov et al. (2017): cladograms
publication summary | cladograms Complete CitationBolotov, I.N., A.V. Kondakov, I.V. Vikhrev, O.V. Aksenova, Y.V. Bespalaya et al. 2017. Ancient river inference explains exceptional Oriental freshwater mussel radiations. Scientific Reports 7. 2135 (14 pp.). Available electronically.
Cladogram Figures(click the figure number to see the cladogram) Cladogram Figure S1
+-- Neotrigonia_margaritacea
+-----------------------------------------+
| +-- Neotrigonia_lamarckii
|
| +----------------- Cumberlandia_monodonta
| |
| | +----- Gibbosula_laosensis
| +-----------------------+ +--------+
| | | | | +-- Pseudunio_auricularius
| | | | +--+
| | | | +-- Pseudunio_marocanus
| | +--+
| | | +-- Margaritifera_dahurica
| | | +--------+
| | | | +-- Margaritifera_margaritifera
| | +--+
| | | +-------- Margaritifera_falcata
| | +--+
| | | +----- Margaritifera_middendorffi
| | +--+
--+ | | +-- Margaritifera_laevis
| | +--+
| | +-- Margaritifera_marrianae
| |
| | +-------- Tamsiella_schroeteriana
| | +-----+
| | | | +----- Etheria_elliptica
| | | +--+
| | | | +-- Chambardia_wahlbergi
| | | +--+
| | | +-- Aspatharia_pfeifferiana
| | +-----------------------+
| | | | +----- Diplodon_suavidicus
| | | | +-----+
| | | | | | +-- Castalia_ambigua
| | | | | +--+
| | | | | +-- Triplodon_corrugatus
| | | +--+
| | | | +----- Alathyria_pertexta
+--+ | | +--+
| | | | | +-- Microdontia_anodontaeformis
| | | | +--+
| | +--+ +-- Lortiella_froggatti
| | |
| | | +----- Velesunio_ambiguus
| | +--+
| | | +-- Alathyria_jacksoni
| | +--+
| | +-- Alathyria_profuga
| |
| | +-- Trapezidens_dolichorhynchus
| | +-----------+
| | | +-- Trapezidens_exolescens
| | +-----------+
| | | | +----------- Lamellidens_corrianus
| | | | |
| | | +--+ +-------- Lamellidens_savadiensis
| | | | |
| | | +--+ +-- Lamellidens_jenkinsianus
| | | | +--+
| | | | | +-- Lamellidens_mainwaringi
| | | +--+
| | | | +-- Lamellidens_ferrugineus
| | +--------+ +--+
| | | | +-- Lamellidens_marginalis
| | | |
| | | | +-- Parreysia_nagpoorensis
| | | | |
| | | | +--------------------+-- Parreysia_plagiosoma
| | | | | |
| | | | | +-- Parreysia_corrugata
+--+ | | |
| | | | +-- Leoparreysia_tavoyensis
| | +--+ +-----------------+
| | | | +-- Leoparreysia_olivacea
| | | |
| | | | +-------------- Nitia_teretiuscula
| | | | +--+
| | | | | | +----------- Coelatura_aegyptiaca__2
| | +--+ | +--+
| | | | | +-------- Coelatura_aegyptiaca__1
| | | | +--+
| | | | | +----- Prisodontopsis_mweruensis
| | | | +--+
| | | | | +-- Coelatura_gabonensis
| | +--+ +--+
| | | +-- Coelatura_kunenensis
| | |
| | | +-------- Indonaia_shurtleffiana
| | | +-----+
| | | | | +----- Indonaia_andersoniana
| | | | +--+
| | | | | +-- Indonaia_rugosa
| | | | +--+
| | +--+ +-- Indonaia_bonneaudii
| | |
| | | +-- Indochinella_pugio
| | | +--------+
| | | | +-- Scabies_crispata
| | | |
| | +--+ +-- Radiatula_myitkyinae
| | | +-----+
| | | | +-- Radiatula_chaudhurii
| | +--+
| | | +----- Scabies_mandarinus
| | +--+
| | | +-- Scabies_humilis
| | +--+
| | +-- Scabies_phaselus
| |
| | +-- Ensidens_telus
+--+ +-----+
| | +-- Ensidens_ingallsianus
| +-----+
| | | +----- Ensidens_sagittarius
| | +--+
| | | +-- Ensidens_spiculus
| | +--+
| +--+ +-- Ensidens_jaculus
| | |
| | | +-- Hyriopsis_bialata
| | | +--------+
| | | | +-- Hyriopsis_khoratensis
| | +--+
| | | +-------- Hyriopsis_panhai
| | +--+
| | | +----- Hyriopsis_myersiana
| | +--+
| +--------------+ | +-- Rectidens_sumatrensis
| | | +--+
| | | +-- Hyriopsis_desowitzi
| | |
| | | +----- Physunio
| | | +--------+
| | | | | +-- Physunio_modelli
| | | | +--+
| | | | +-- Physunio_superbus
| | | |
| | +--+ +-------- Yaukthwa_inlenensis
| | | +--+
| | | | | +----- Yaukthwa_panhai
| | | | +--+
| | | | | +-- Yaukthwa_rectangularis
| | | | +--+
| | +--+ +-- Yaukthwa_nesemanni
| | |
| | | +-- Lens_contradens
| | | +-----+
| | | | +-- Lens_micropterus
| | +--+
| | | +----- Lens_eximius
| | +--+
| | | +-- Lens_comptus
| | +--+
| | +-- Lens_novoselovi
| |
| | +-------------------- Indopseudodon_indawgyiensis
| | |
| | | +-------- Pilsbryoconcha_exilis__1
| | | +--------+
+--+ +--------+ | | +----- Pilsbryoconcha_exilis__3
| | | | +--+
| | | | | +-- Pilsbryoconcha_exilis__2
| | | | +--+
| | | | +-- Pilsbryoconcha_carinifera
| | +--+
| | | +-------- Sundadontina_tumida
| | | +-----+
| | | | | +----- Pseudodon_vagulus
| | | | +--+
| | | | | +-- Pseudodon_mekongi
| | | | +--+
| | +--+ +-- Pseudodon_vondembuschianus
| | |
| | | +-- Nyeinchanconcha_nyeinchani
| | | +--------+
| | | | +-- Sundadontina_cumingii
| | +--+
| | | +-------- Thaiconcha_callifera
| | +--+
| | | +----- Bineurus_exilis
| | +--+
| | | +-- Bineurus_mouhotii
| | +--+
| | +-- Bineurus_loeiensis
| |
| | +-------- Amblema_plicata
| | +--+
| | | | +----- Ortmanniana_ligamentina
| | | +--+
| | | | +-- Lampsilis_cardium
| | | +--+
+--+ +-----+ +-- Cambarunio_iris
| | |
| | | +-- Quadrula_quadrula
| | | +-----+
| | | | +-- Quadrula_verrucosa
| | +--+
| | | +----- Pleurobema_sintoxia
| | +--+
| | | +-- Eurynia_dilatata
| | +--+
| | +-- Elliptio_complanata
| +--------+
| | | +-- Sinohyriopsis_cumingii
| | | +-----+
| | | | +-- Chamberlainia_somsakpanhai
| | | |
| | | +-----+ +-- Potomida_littoralis
| | | | | +--+
| | | | | | +-- Pronodularia_japanensis
| | | | +--+
| | | | | +-- Lamprotula_caveata
| | | | +--+
| | +--+ +-- Lamprotula_leaii
| | |
| | | +----- Sinosolenaia_carinata
| | | +-----+
| | | | | +-- Sinosolenaia_oleivora
| | | | +--+
| | +--+ +-- Sinosolenaia_recognita
| | |
| | | +-------- Gonidea_angulata
| | +--+
+--+ | +----- Microcondylaea_bonellii
| +--+
| | +-- Leguminaia_wheatleyi
| +--+
| +-- Leguminaia
|
| +----- Unio_tumidus
| +--------+
| | | +-- Unio_pictorum
| | +--+
| | +-- Unio_crassus
| +--------+
| | | +----------- Aculamprotula_tortuosa
| | | |
| | | | +-- Nodularia_douglasiae
| | +--+ +-----+
| | | | +-- Nodularia_nuxpersicae
| | | |
| | +--+ +-- Cuneopsis_rufescens
| | | +--+
| | | | +-- Cuneopsis_szechenyii
| | +--+
+--+ | +-- Tchangsinaia_piscicula
| +--+
| +-- Schistodesmus_lampreyanus
|
| +-- Lanceolaria_grayii
| +-----------------+
| | +-- Lanceolaria_gladiola
| |
| | +-- Cristaria_plicata
| | +--------+
| | | +-- Cristaria_radiata
+--+ +-----+
| | | +-------- Pletholophus_tenuis
| | +--+
| | | +----- Sinanodonta_lucida
| | +--+
| | | +-- Sinanodonta_jourdyi
+--+ +--+
| +-- Sinanodonta_lauta
|
| +-------------- Anodonta_anatina
| |
| | +----- Pseudanodonta_complanata
+--+ +-----+
| | | +-- Anodonta_cygnea
| | +--+
| | +-- Anodonta_nuttalliana
+--+
| +-- Lasmigona_costata
| +-----+
| | +-- Alasmidonta_marginata
+--+
| +----- Simpsonaias_ambigua
+--+
| +-- Pyganodon_grandis
+--+
+-- Strophitus_undulatus
Cladogram Figure S2
+-- Neotrigonia_margaritacea
+--------------------------------+
| +-- Neotrigonia_lamarckii
|
| +-------------------- Chamberlainia_somsakpanhai
| |
| +-----------+ +----------------- Cumberlandia_monodonta
| | | |
| | | | +-------------- Gibbosula_laosensis
| | +--+ |
| | | | +-- Pseudunio_auricularius
| | | +-----------+
| | +--+ +-- Pseudunio_marocanus
| | |
| | | +-- Margaritifera_dahurica
| | | +--------+
| | | | +-- Margaritifera_margaritifera
| | +--+
| | | +-------- Margaritifera_falcata
| | +--+
| | | +----- Margaritifera_middendorffi
| | +--+
| | | +-- Margaritifera_laevis
| | +--+
| | +-- Margaritifera_marrianae
| |
| | +-- Tamsiella_schroeteriana
| | +--+
| | | +-- Etheria_elliptica
| | +--------+
| | | | +-- Aspatharia_pfeifferiana
--+ | | +--+
| | | +-- Chambardia_wahlbergi
| | |
| +-----------------+ +----- Diplodon_suavidicus
| | | +-----+
| | | | | +-- Castalia_ambigua
| | | | +--+
| | | | +-- Triplodon_corrugatus
| | +--+
| | | +----- Velesunio_ambiguus
| | | +--+
| | | | | +-- Alathyria_jacksoni
| | | | +--+
| | +--+ +-- Alathyria_profuga
| | |
| | | +----- Microdontia_anodontaeformis
| | +--+
| | | +-- Alathyria_pertexta
| | +--+
| | +-- Lortiella_froggatti
| |
| | +-- Parreysia_nagpoorensis
| | |
| | +-----------+-- Parreysia_plagiosoma
| | | |
| | | +-- Parreysia_corrugata
| | |
| | +--------+ +-- Trapezidens_dolichorhynchus
| | | | +--------+
| | | | | +-- Trapezidens_exolescens
| | | | |
+--+ | +--+ +-------- Lamellidens_savadiensis
| | | |
| | | +-------- Lamellidens_corrianus
| | +--+
| | | +-- Lamellidens_marginalis
| | | +--+
| | | | +-- Lamellidens_ferrugineus
| +-----+ +--+
| | | | +-- Lamellidens_jenkinsianus
| | | +--+
| | | +-- Lamellidens_mainwaringi
| | |
| | | +-- Leoparreysia_olivacea
| | | +-----------------+
| | | | +-- Leoparreysia_tavoyensis
| | | |
| | | | +-------------- Nitia_teretiuscula
| | | | +--+
| | | | | | +----------- Coelatura_aegyptiaca__1
| | +--+ | +--+
| | | | | +-------- Coelatura_aegyptiaca__2
| | | | +--+
| | | | | +----- Coelatura_gabonensis
| | | | +--+
| | | | | +-- Coelatura_kunenensis
| | +--+ +--+
| | | +-- Prisodontopsis_mweruensis
| | |
| | | +-------- Indonaia_shurtleffiana
| | | +-----+
| | | | | +----- Indonaia_andersoniana
| | | | +--+
| | | | | +-- Indonaia_rugosa
| | | | +--+
| | +--+ +-- Indonaia_bonneaudii
| | |
| | | +-- Indochinella_pugio
| | | +--------+
| | | | +-- Scabies_crispata
| | | |
| | +--+ +-- Radiatula_myitkyinae
| | | +-----+
| | | | +-- Radiatula_chaudhurii
| | +--+
| | | +----- Scabies_mandarinus
| | +--+
| | | +-- Scabies_humilis
| | +--+
| | +-- Scabies_phaselus
| |
| | +----- Unio_tumidus
| | +-----+
| | | | +-- Unio_crassus
| | | +--+
| | | +-- Unio_pictorum
| | +-----+
+--+ | | +-------- Aculamprotula_tortuosa
| | | |
| | | +-------- Schistodesmus_lampreyanus
| | | |
| | +--+ +-- Nodularia_douglasiae
| | +-----+
| | | +-- Nodularia_nuxpersicae
| | |
| | | +----- Cuneopsis_szechenyii
| | +--+
| | | +-- Tchangsinaia_piscicula
| | +--+
| +--------+ +-- Cuneopsis_rufescens
| | |
| | | +-- Lanceolaria_grayii
| | | +-----------+
| | | | +-- Lanceolaria_gladiola
| | | |
| | | | +-- Cristaria_plicata
| | | | +--------+
| | | | | +-- Cristaria_radiata
| | | +--+
| | | | | +-------- Pletholophus_tenuis
| | | | +--+
| | | | | +----- Sinanodonta_lucida
| | +--+ +--+
| | | | +-- Sinanodonta_lauta
| | | +--+
| | | +-- Sinanodonta_jourdyi
| | |
| | | +-------- Anodonta_anatina
| | | |
| | | +-------- Pseudanodonta_complanata
| | | |
| | | | +-- Anodonta_cygnea
| | +-----+-----+
| | | +-- Anodonta_nuttalliana
| | |
| | | +----- Pyganodon_grandis
| | | |
| | | +----- Simpsonaias_ambigua
| | +--+
| | +----- Strophitus_undulatus
| | |
| | | +-- Alasmidonta_marginata
| | +--+
| | +-- Lasmigona_costata
| |
| | +-------- Lens_contradens
| | +-----+
+--+ | | +----- Lens_eximius
| | +--+
| | | +-- Lens_comptus
| | +--+
| +--+ +-- Lens_novoselovi
| | |
| | | +----- Physunio
| | | +-----+
| | | | | +-- Physunio_modelli
| | | | +--+
| | +--+ +-- Physunio_superbus
| | |
| | | +-------- Yaukthwa_inlenensis
| | +--+
| | | +----- Yaukthwa_panhai
| +-----+ +--+
| | | | +-- Yaukthwa_rectangularis
| | | +--+
| | | +-- Yaukthwa_nesemanni
| | |
| | | +----- Hyriopsis_panhai
| | | |
| | | +--------+----- Hyriopsis_khoratensis
| | | | |
| | | | | +-- Hyriopsis_myersiana
| | | | +--+
| | +--+ +-- Hyriopsis_bialata
| | |
| | | +----------- Hyriopsis_desowitzi
| | | |
| | | +----------- Rectidens_sumatrensis
| | +--+
| | | +-- Ensidens_telus
| | | +-----+
| | | | +-- Ensidens_ingallsianus
| | +--+
| | | +----- Ensidens_sagittarius
| | +--+
| | | +-- Ensidens_spiculus
+--+ +--+
| +-- Ensidens_jaculus
|
| +-------- Amblema_plicata
| +--+
| | | +----- Cambarunio_iris
| | +--+
| | | +-- Ortmanniana_ligamentina
| | +--+
| +--------+ +-- Lampsilis_cardium
| | |
| | | +-- Quadrula_quadrula
| | | +-----+
| | | | +-- Quadrula_verrucosa
| | +--+
| | | +----- Pleurobema_sintoxia
| | +--+
| | | +-- Elliptio_complanata
| | +--+
| | +-- Eurynia_dilatata
| |
| | +-------- Sinohyriopsis_cumingii
| | |
| | +-----+ +-- Lamprotula_caveata
+--+ | | +--+
| | | | +-- Lamprotula_leaii
| | +--+
| | | +-- Potomida_littoralis
| +--+ +--+
| | | +-- Pronodularia_japanensis
| | |
| | | +----------- Sinosolenaia_carinata
| | | |
| | | | +-- Sinosolenaia_oleivora
| | +--+--------+
| | | +-- Sinosolenaia_recognita
| | |
| | | +-------- Gonidea_angulata
| | +--+
| | | +----- Microcondylaea_bonellii
+--+ +--+
| | +-- Leguminaia_wheatleyi
| +--+
| +-- Leguminaia
|
| +-------- Pilsbryoconcha_exilis__3
| +-----+
| | | +----- Pilsbryoconcha_exilis__2
| | +--+
| | | +-- Pilsbryoconcha_exilis__1
| | +--+
| | +-- Pilsbryoconcha_carinifera
| |
+--+ +----------- Indopseudodon_indawgyiensis
| |
| | +-------- Thaiconcha_callifera
| +--+
| | | +----- Bineurus_exilis
| | +--+
| | | +-- Bineurus_mouhotii
+--+ +--+
| +-- Bineurus_loeiensis
|
| +----- Pseudodon_vagulus
| +--+
| | | +-- Pseudodon_mekongi
| | +--+
| | +-- Pseudodon_vondembuschianus
+--+
| +-- Pseudodon_inoscularis
| +--+
| | +-- Sundadontina_tumida
+--+
| +-- Nyeinchanconcha_nyeinchani
+--+
+-- Sundadontina_cumingii
|
