The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)
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Bivalvia | Unionoida | Unionoidea | Unionidae | Gonideinae | Gonideini genus Microcondylaea Vest, 1866: cladomics 1 Recent valid species (1 total species) Cladomics Summarynumber of species = 1 (1 total) analyzed in 61 cladograms in 37 publications maximum number of terminals in an analysis =
1 (mean = 1, median = 1)
Dai et al., 2025 [Publ 4441], fig. 1 ** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram ** iT | tT | bi | riS | oT | roT | roS | iD | 1 | 1 | 0 | 1 | 47 | 16 | 11 | -- |
+-------------------- Sinosolenaia
|
| +----- Gonidea
--+ +-----------+
| | | +-- Leguminaia
| | +--+
+--+ +-- Microcondylaea_bonellii **
|
| +-------------- Pseudopostolata
| |
+--+ +-- Koreosolenaia
| +--------+
| | +-- Parvasolenaia
+--+
| +-------- Postolata
| |
+--+ +-- Obovalis
| +--+
| | +-- Ptychorhynchus
+--+
| +-- Guiunio
+--+
+-- Cosmopseudodon
Wu et al., 2021 [Publ 4366], fig. 2_I iT | tT | bi | riS | oT | roT | roS | iD | 1 | 1 | 0 | 1 | 90 | 7 | 6 | -- |
+----- Gonidea
+-----+
| | +-- Leguminaia
| +--+
--+ +-- Microcondylaea_bonellii **
|
| +-------- Inversidens
+--+
| +----- Sinosolenaia
+--+
| +-- Parvasolenaia
+--+
+-- Ptychorhynchus
Zieritz et al., 2021 [Publ 4218], fig. 5 iT | tT | bi | riS | oT | roT | roS | iD | 1 | 1 | 0 | 1 | 31 | 1 | 1 | -- |
+-- Gonidea
--+
+-- Microcondylaea_bonellii **
Wu et al., 2021 [Publ 4366], fig. 3-BI iT | tT | bi | riS | oT | roT | roS | iD | 1 | 1 | 0 | 1 | 72 | 4 | 3 | -- |
+-- Parvasolenaia
+--+
| +-- Ptychorhynchus
--+
| +-- Microcondylaea_bonellii **
+--+
+-- Sinosolenaia
Huang et al., 2019 [Publ 3843], fig. S4 iT | tT | bi | riS | oT | roT | roS | iD | 1 | 1 | 0 | 1 | 198 | 9 | 6 | -- |
+-------- Sinosolenaia_carinata
|
| +----- Gonidea
+--+
| | +-- Leguminaia
| +--+
| +-- Microcondylaea_bonellii **
--+
| +----- Ptychorhynchus
+--+
| | +-- Parvasolenaia
| +--+
| +-- Koreosolenaia
|
| +----- Sinosolenaia_recognita
+--+
| +-- Sinosolenaia_iridinea
+--+
+-- Sinosolenaia_oleivora
Zieritz et al., 2021 [Publ 4231], fig. 3-M_4 iT | tT | bi | riS | oT | roT | roS | iD | 1 | 1 | 0 | 1 | 22 | 1 | 1 | -- |
+-- Microcondylaea_bonellii **
--+
+-- Sinosolenaia
Bolotov et al., 2020 [Publ 4100], fig. S1 iT | tT | bi | riS | oT | roT | roS | iD | 1 | 1 | 0 | 1 | 111 | 8 | 5 | -- |
+-- Sinosolenaia_iridinea
|
+-- Sinosolenaia_recognita__1
|
+-- Sinosolenaia_carinata
|
+-- Leguminaia
|
+-- Microcondylaea_bonellii **
--+
+-- Gonidea
|
+-- Ptychorhynchus
|
+-- Parvasolenaia
|
+-- Sinosolenaia_recognita__2
|
+-- Sinosolenaia_oleivora
Bolotov et al., 2017 [Publ 3761], fig. S1 iT | tT | bi | riS | oT | roT | roS | iD | 1 | 1 | 0 | 1 | 138 | 6 | 3 | -- |
+-------- Sinosolenaia
--+
| +----- Gonidea
+--+
| +-- Microcondylaea_bonellii **
+--+
+-- Leguminaia
Dai et al., 2025 [Publ 4440], fig. 3 iT | tT | bi | riS | oT | roT | roS | iD | 1 | 1 | 0 | 1 | 87 | 5 | 4 | -- |
+-- Microcondylaea_bonellii **
+-----+
| +-- Sinosolenaia
--+
| +----- Parvasolenaia
+--+
| +-- Postolata
+--+
+-- Ptychorhynchus
Froufe et al., 2019 [Publ 3936], fig. 3 iT | tT | bi | riS | oT | roT | roS | iD | 1 | 1 | 0 | 1 | 43 | 3 | 2 | -- |
+----- Ptychorhynchus
--+
| +-- Microcondylaea_bonellii **
+--+
+-- Sinosolenaia
Dai et al., 2024 [Publ 4358], fig. 4 iT | tT | bi | riS | oT | roT | roS | iD | 1 | 1 | 0 | 1 | 51 | 3 | 3 | -- |
+-------- Microcondylaea_bonellii **
--+
| +----- Sinosolenaia
+--+
| +-- Ptychorhynchus
+--+
+-- Parvasolenaia
Huang et al., 2019 [Publ 3843], fig. 2 iT | tT | bi | riS | oT | roT | roS | iD | 1 | 1 | 0 | 1 | 198 | 9 | 6 | -- |
+----------- Sinosolenaia
|
| +----- Gonidea
--+ +--+
| | | +-- Leguminaia
| | +--+
+--+ +-- Microcondylaea_bonellii **
|
| +-- Ptychorhynchus
| |
+-----+-- Parvasolenaia
|
+-- Koreosolenaia
Wu et al., 2021 [Publ 4366], fig. 6 iT | tT | bi | riS | oT | roT | roS | iD | 1 | 1 | 0 | 1 | 90 | 7 | 6 | -- |
+----- Sinosolenaia
+-----+
| | +-- Ptychorhynchus
| +--+
--+ +-- Parvasolenaia
|
| +-------- Inversidens
+--+
| +----- Gonidea
+--+
| +-- Microcondylaea_bonellii **
+--+
+-- Leguminaia
Nagel et al., 1998 [Publ 1946], fig. 9 iT | tT | bi | riS | oT | roT | roS | iD | 1 | 1 | 0 | 1 | 5 | 0 | 0 | -- |
Konopleva et al., 2023 [Publ 4269], fig. S1_4 iT | tT | bi | riS | oT | roT | roS | iD | 1 | 1 | 0 | 1 | 55 | 5 | 5 | -- |
+----------- Sinosolenaia
|
--+ +-- Ptychorhynchus
| +-----+
| | +-- Parvasolenaia
+--+
| +----- Gonidea
+--+
| +-- Microcondylaea_bonellii **
+--+
+-- Leguminaia
Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2 iT | tT | bi | riS | oT | roT | roS | iD | 1 | 1 | 0 | 1 | 79 | 5 | 4 | -- |
+-------- Sinosolenaia
--+
| +----- Gonidea
+--+
| +-- Microcondylaea_bonellii **
+--+
+-- Leguminaia
Zieritz et al., 2021 [Publ 4231], fig. 3-F_5 iT | tT | bi | riS | oT | roT | roS | iD | 1 | 1 | 0 | 1 | 47 | 3 | 2 | -- |
+----- Ptychorhynchus
--+
| +-- Microcondylaea_bonellii **
+--+
+-- Sinosolenaia
Froufe et al., 2017 [Publ 3802], fig. 7 iT | tT | bi | riS | oT | roT | roS | iD | 1 | 1 | 0 | 1 | 9 | 0 | 0 | -- |
-- Microcondylaea_bonellii **
Wu et al., 2024 [Publ 4388], fig. 6 iT | tT | bi | riS | oT | roT | roS | iD | 1 | 1 | 0 | 1 | 53 | 8 | 5 | -- |
+-- Microcondylaea_bonellii **
+--------+
| +-- Sinosolenaia
--+
| +-------- Parvasolenaia
+--+
| +----- Ptychorhynchus
+--+
| +-- Postolata
+--+
+-- Cosmopseudodon
Liu et al., 2024 [Publ 4380], fig. 5 iT | tT | bi | riS | oT | roT | roS | iD | 1 | 1 | 0 | 1 | 36 | 8 | 7 | -- |
+-------------- Sinosolenaia
|
| +-- Gonidea
--+ +--------+
| | +-- Microcondylaea_bonellii **
| |
+--+ +-- Koreosolenaia
| +-----+
| | +-- Parvasolenaia
+--+
| +----- Postolata
+--+
| +-- Obovalis
+--+
+-- Ptychorhynchus
Dai et al., 2024 [Publ 4389], fig. 3 iT | tT | bi | riS | oT | roT | roS | iD | 1 | 1 | 0 | 1 | 46 | 15 | 10 | -- |
+-------------------- Sinosolenaia
|
| +----- Gonidea
--+ +-----------+
| | | +-- Leguminaia
| | +--+
+--+ +-- Microcondylaea_bonellii **
|
| +-------------- Pseudopostolata
| |
+--+ +-- Parvasolenaia
| +--------+
| | +-- Koreosolenaia
+--+
| +-------- Postolata
+--+
| +----- Ptychorhynchus
+--+
| +-- Obovalis
+--+
+-- Cosmopseudodon
Pavan-Kumar et al., 2022 [Publ 4263], fig. 4 iT | tT | bi | riS | oT | roT | roS | iD | 1 | 1 | 0 | 1 | 35 | 3 | 2 | -- |
+----- Parvasolenaia
--+
| +-- Microcondylaea_bonellii **
+--+
+-- Sinosolenaia
Wu et al., 2023 [Publ 4339], fig. 4_S5-aa iT | tT | bi | riS | oT | roT | roS | iD | 1 | 1 | 0 | 1 | 94 | 5 | 3 | -- |
+-------- Microcondylaea_bonellii **
--+
| +----- Sinosolenaia
+--+
| +-- Parvasolenaia
+--+
+-- Ptychorhynchus
Wu et al., 2021 [Publ 4215], fig. 3 iT | tT | bi | riS | oT | roT | roS | iD | 1 | 1 | 0 | 1 | 23 | 7 | 6 | -- |
+-- Ptychorhynchus
+--------+
| +-- Parvasolenaia
--+
| +-------- Inversidens
+--+
| +----- Gonidea
+--+
| +-- Microcondylaea_bonellii **
+--+
+-- Leguminaia
Araujo et al., 2017 [Publ 3801], fig. 2 iT | tT | bi | riS | oT | roT | roS | iD | 1 | 1 | 0 | 1 | 13 | 1 | 1 | -- |
+-- Gonidea
--+
+-- Microcondylaea_bonellii **
Wu et al., 2021 [Publ 4366], fig. 4 iT | tT | bi | riS | oT | roT | roS | iD | 1 | 1 | 0 | 1 | 67 | 4 | 3 | -- |
+-- Parvasolenaia
+--+
| +-- Ptychorhynchus
--+
| +-- Microcondylaea_bonellii **
+--+
+-- Sinosolenaia
Dai et al., 2023 [Publ 4285], fig. 3-ML iT | tT | bi | riS | oT | roT | roS | iD | 1 | 1 | 0 | 1 | 44 | 13 | 9 | -- |
+-------------- Sinosolenaia
|
| +----- Gonidea
--+ +-----+
| | | +-- Leguminaia
| | +--+
| | +-- Microcondylaea_bonellii **
+--+
| +-- Koreosolenaia
| +-----+
| | +-- Parvasolenaia
+--+
| +----- Postolata
+--+
| +-- Ptychorhynchus
+--+
+-- Obovalis
Wu et al., 2024 [Publ 4350], fig. 8b_S2 iT | tT | bi | riS | oT | roT | roS | iD | 1 | 1 | 0 | 1 | 90 | 6 | 4 | -- |
+-------- Inversidens
|
--+ +-- Parvasolenaia
| +--+
| | +-- Ptychorhynchus
+--+
| +-- Microcondylaea_bonellii **
+--+
+-- Sinosolenaia
Nagel et al., 1998 [Publ 1946], fig. 1 iT | tT | bi | riS | oT | roT | roS | iD | 1 | 1 | 0 | 1 | 8 | 0 | 0 | -- |
-- Microcondylaea_bonellii **
Wu et al., 2024 [Publ 4350], fig. 5c iT | tT | bi | riS | oT | roT | roS | iD | 1 | 1 | 0 | 1 | 36 | 8 | 7 | -- |
+-------------- Sinosolenaia
|
--+ +----------- Gonidea
| |
+--+----------- Microcondylaea_bonellii **
|
| +-------- Koreosolenaia
+--+
| +----- Parvasolenaia
+--+
| +-- Postolata
+--+
+-- Ptychorhynchus
Lyubas et al., 2022 [Publ 4255], fig. 7 iT | tT | bi | riS | oT | roT | roS | iD | 1 | 1 | 0 | 1 | 30 | 0 | 0 | -- |
-- Microcondylaea_bonellii **
Dai et al., 2023 [Publ 4285], fig. 5 iT | tT | bi | riS | oT | roT | roS | iD | 1 | 1 | 0 | 1 | 52 | 5 | 4 | -- |
+-- Microcondylaea_bonellii **
+-----+
| +-- Sinosolenaia
--+
| +----- Parvasolenaia
+--+
| +-- Postolata
+--+
+-- Ptychorhynchus
Wu et al., 2021 [Publ 4366], fig. 3-ML iT | tT | bi | riS | oT | roT | roS | iD | 1 | 1 | 0 | 1 | 72 | 4 | 3 | -- |
+-- Parvasolenaia
+--+
| +-- Ptychorhynchus
--+
| +-- Microcondylaea_bonellii **
+--+
+-- Sinosolenaia
Froufe et al., 2016 [Publ 3727], fig. 3_4 iT | tT | bi | riS | oT | roT | roS | iD | 1 | 1 | 0 | 1 | 6 | 1 | 1 | -- |
-- Microcondylaea_bonellii **
Pfarrer et al., 2022 [Publ 4305], fig. 2-5 iT | tT | bi | riS | oT | roT | roS | iD | 1 | 1 | 0 | 1 | 11 | 0 | 0 | -- |
-- Microcondylaea_bonellii **
Wu et al., 2024 [Publ 4350], fig. 8a_S1 iT | tT | bi | riS | oT | roT | roS | iD | 1 | 1 | 0 | 1 | 90 | 6 | 4 | -- |
+-------- Inversidens
|
--+ +-- Parvasolenaia
| +--+
| | +-- Ptychorhynchus
+--+
| +-- Microcondylaea_bonellii **
+--+
+-- Sinosolenaia
Matos et al., 2024 [Publ 4384], fig. 3 iT | tT | bi | riS | oT | roT | roS | iD | 1 | 1 | 0 | 1 | 19 | 3 | 2 | -- |
+----- Parvasolenaia
--+
| +-- Microcondylaea_bonellii **
+--+
+-- Sinosolenaia
Zhang et al., 2024 [Publ 4397], fig. 5 iT | tT | bi | riS | oT | roT | roS | iD | 1 | 1 | 0 | 1 | 27 | 11 | 7 | -- |
+----------------- Inversidens
|
--+ +-- Microcondylaea_bonellii **
| +-----------+
| | +-- Sinosolenaia
+--+
| +----------- Parvasolenaia
+--+
| +-------- Koreosolenaia
+--+
| +----- Ptychorhynchus
+--+
| +-- Postolata
+--+
+-- Cosmopseudodon
Lyubas et al., 2022 [Publ 4255], fig. S1 iT | tT | bi | riS | oT | roT | roS | iD | 1 | 1 | 0 | 1 | 30 | 0 | 0 | -- |
-- Microcondylaea_bonellii **
Froufe et al., 2019 [Publ 3936], fig. 6 iT | tT | bi | riS | oT | roT | roS | iD | 1 | 1 | 0 | 1 | 43 | 3 | 2 | -- |
+----- Ptychorhynchus
--+
| +-- Microcondylaea_bonellii **
+--+
+-- Sinosolenaia
Bolotov et al., 2020 [Publ 4100], fig. 1 iT | tT | bi | riS | oT | roT | roS | iD | 1 | 1 | 0 | 1 | 117 | 8 | 5 | -- |
-- Microcondylaea_bonellii **
Bolotov et al., 2017 [Publ 3764], fig. S1 iT | tT | bi | riS | oT | roT | roS | iD | 1 | 1 | 0 | 1 | 124 | 6 | 3 | -- |
-- Microcondylaea_bonellii **
Wu et al., 2024 [Publ 4350], fig. 6-7 iT | tT | bi | riS | oT | roT | roS | iD | 1 | 1 | 0 | 1 | 36 | 8 | 7 | -- |
+----------------- Inversidens
|
--+ +-------------- Sinosolenaia
| |
+--+ +-- Gonidea
| +--------+
| | +-- Microcondylaea_bonellii **
+--+
| +-------- Koreosolenaia
+--+
| +----- Parvasolenaia
+--+
| +-- Postolata
+--+
+-- Ptychorhynchus
Wu et al., 2023 [Publ 4339], fig. 4_S5-nt iT | tT | bi | riS | oT | roT | roS | iD | 1 | 1 | 0 | 1 | 94 | 5 | 3 | -- |
+-- Ptychorhynchus
+--+
| +-- Microcondylaea_bonellii **
--+
| +-- Parvasolenaia
+--+
+-- Sinosolenaia
Wu et al., 2024 [Publ 4350], fig. 5a iT | tT | bi | riS | oT | roT | roS | iD | 1 | 1 | 0 | 1 | 36 | 8 | 7 | -- |
+-------------- Sinosolenaia
|
--+ +-- Gonidea
| +--------+
| | +-- Microcondylaea_bonellii **
+--+
| +-------- Parvasolenaia
+--+
| +----- Koreosolenaia
+--+
| +-- Postolata
+--+
+-- Ptychorhynchus
Bolotov et al., 2017 [Publ 3761], fig. S2 iT | tT | bi | riS | oT | roT | roS | iD | 1 | 1 | 0 | 1 | 138 | 6 | 3 | -- |
+-------- Sinosolenaia_carinata
|
| +-- Sinosolenaia_oleivora
--+-----+
| +-- Sinosolenaia_recognita
|
| +----- Gonidea
+--+
| +-- Microcondylaea_bonellii **
+--+
+-- Leguminaia
Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4 iT | tT | bi | riS | oT | roT | roS | iD | 1 | 1 | 0 | 1 | 120 | 11 | 8 | -- |
+-------- Sinosolenaia
+--+
| | +----- Gonidea
| +--+
| | +-- Leguminaia
--+ +--+
| +-- Microcondylaea_bonellii **
|
| +-------- Inversidens
| |
+--+ +-- Obovalis
| +--+
| | +-- Ptychorhynchus
+--+
| +-- Parvasolenaia
+--+
+-- Koreosolenaia
Wu et al., 2024 [Publ 4388], fig. 5 iT | tT | bi | riS | oT | roT | roS | iD | 1 | 1 | 0 | 1 | 37 | 9 | 8 | -- |
+-------------------- Sinosolenaia
|
--+ +-- Gonidea
| +--------------+
| | +-- Microcondylaea_bonellii **
+--+
| +-------------- Koreosolenaia
+--+
| +----------- Parvasolenaia
+--+
| +-------- Postolata
+--+
| +----- Cosmopseudodon
+--+
| +-- Ptychorhynchus
+--+
+-- Obovalis
Wu et al., 2024 [Publ 4350], fig. 5b iT | tT | bi | riS | oT | roT | roS | iD | 1 | 1 | 0 | 1 | 36 | 8 | 7 | -- |
+-------------- Sinosolenaia
|
--+ +-- Gonidea
| +--------+
| | +-- Microcondylaea_bonellii **
+--+
| +-------- Koreosolenaia
+--+
| +----- Parvasolenaia
+--+
| +-- Postolata
+--+
+-- Ptychorhynchus
Huang et al., 2019 [Publ 3843], fig. S3 iT | tT | bi | riS | oT | roT | roS | iD | 1 | 1 | 0 | 1 | 196 | 8 | 6 | -- |
+----------- Sinosolenaia
|
| +----- Gonidea
--+ +--+
| | | +-- Leguminaia
| | +--+
+--+ +-- Microcondylaea_bonellii **
|
| +----- Ptychorhynchus
+--+
| +-- Parvasolenaia
+--+
+-- Koreosolenaia
Froufe et al., 2019 [Publ 3936], fig. 4 iT | tT | bi | riS | oT | roT | roS | iD | 1 | 1 | 0 | 1 | 18 | 1 | 1 | -- |
+-- Microcondylaea_bonellii **
--+
+-- Sinosolenaia
Pavan-Kumar et al., 2022 [Publ 4263], fig. S5 iT | tT | bi | riS | oT | roT | roS | iD | 1 | 1 | 0 | 1 | 37 | 3 | 2 | -- |
+----- Sinosolenaia
--+
| +-- Parvasolenaia
+--+
+-- Microcondylaea_bonellii **
Wu et al., 2022 [Publ 4238], fig. 3 iT | tT | bi | riS | oT | roT | roS | iD | 1 | 1 | 0 | 1 | 72 | 4 | 3 | -- |
+-- Ptychorhynchus
+--+
| +-- Parvasolenaia
--+
| +-- Microcondylaea_bonellii **
+--+
+-- Sinosolenaia
Matos et al., 2024 [Publ 4383], fig. 3 iT | tT | bi | riS | oT | roT | roS | iD | 1 | 1 | 0 | 1 | 19 | 3 | 2 | -- |
+----- Parvasolenaia
--+
| +-- Microcondylaea_bonellii **
+--+
+-- Sinosolenaia
Dai et al., 2023 [Publ 4285], fig. 3-BI iT | tT | bi | riS | oT | roT | roS | iD | 1 | 1 | 0 | 1 | 44 | 13 | 9 | -- |
+-------------- Sinosolenaia
|
| +----- Gonidea
--+ +-----+
| | | +-- Leguminaia
| | +--+
| | +-- Microcondylaea_bonellii **
+--+
| +-- Koreosolenaia
| +-----+
| | +-- Parvasolenaia
+--+
| +----- Postolata
+--+
| +-- Ptychorhynchus
+--+
+-- Obovalis
Bolotov et al., 2020 [Publ 4084], fig. 1 iT | tT | bi | riS | oT | roT | roS | iD | 1 | 1 | 0 | 1 | 35 | 2 | 2 | -- |
-- Microcondylaea_bonellii **
Lopes-Lima et al., 2021 [Publ 4217], fig. 14 iT | tT | bi | riS | oT | roT | roS | iD | 1 | 1 | 0 | 1 | 4 | 4 | 2 | -- |
+----- Leguminaia
--+
| +-- Gonidea
+--+
+-- Microcondylaea_bonellii **
Bolotov et al., 2018 [Publ 3826], fig. 2 iT | tT | bi | riS | oT | roT | roS | iD | 1 | 1 | 0 | 1 | 82 | 2 | 2 | -- |
-- Microcondylaea_bonellii **
Lopes-Lima et al., 2017 [Publ 3734], fig. 1 iT | tT | bi | riS | oT | roT | roS | iD | 1 | 1 | 0 | 1 | 16 | 0 | 0 | -- |
-- Microcondylaea_bonellii **
Zieritz et al., 2024 [Publ 4381], fig. 2 iT | tT | bi | riS | oT | roT | roS | iD | 1 | 1 | 0 | 1 | 45 | 1 | 1 | -- |
+-- Gonidea
--+
+-- Microcondylaea_bonellii **
Froufe et al., 2019 [Publ 3936], fig. 3-M iT | tT | bi | riS | oT | roT | roS | iD | 1 | 1 | 0 | 1 | 18 | 1 | 1 | -- |
+-- Microcondylaea_bonellii **
--+
+-- Sinosolenaia
|