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Page last updated
Thu 3 September 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Search the database pages:

Bivalvia | Unionida | Unionoidea | Unionidae | Gonideinae | Rectidentini | Rectidentina

genus Hyriopsis Conrad, 1853: cladomics
     10 Recent valid species (10 total species)

Cladomics Summary

number of species = 10 (10 total)
analyzed in 60 cladograms in 39 publications
maximum number of terminals in an analysis = 9 (mean = 2.65, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 28
maximum number of included terminals = 9, (mean = 4.54 median = 5)
proportion of cladograms that support basic monophyly = 0.32

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.


PCoA Ordination


NJ Tree

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
91199622820
  +----------------- Ensidens               
  |                                         
  |           +----- Rectidens              
--+  +--------+                             
  |  |        |  +-- Hyriopsis_delaportei **   
  |  |        +--+                          
  |  |           +-- Hyriopsis_desowitzi **    
  +--+                                      
     |  +----------- Hyriopsis_panhai **       
     |  |                                   
     |  |        +-- Hyriopsis_bialata **      
     +--+  +-----+                          
        |  |     +-- Hyriopsis_khoratensis **  
        |  |                                
        +--+     +-- Hyriopsis_kratiensis **   
           |  +--+                          
           |  |  +-- Hyriopsis_phuphaniensis **
           +--+                             
              |  +-- Hyriopsis_myersiana **    
              +--+                          
                 +-- Hyriopsis_sakhonensis **

Konopleva et al., 2023 [Publ 4262], fig. 8

iTtTbiriSoTroTroSiD
9108973727
  +----------------------- Ensidens               
  |                                               
--+                    +-- Hyriopsis_desowitzi **    
  |  +-----------------+                          
  |  |                 +-- Hyriopsis_delaportei **   
  +--+                                            
     |  +----------------- Rectidens              
     |  |                                         
     +--+  +-------------- Hyriopsis_panhai **       
        |  |                                      
        +--+  +----------- Hyriopsis_myersiana **    
           |  |                                   
           +--+        +-- Hyriopsis_khoratensis **  
              |  +-----+                          
              |  |     +-- Hyriopsis_bialata **      
              +--+                                
                 |  +----- Hyriopsis_sakhonensis **  
                 +--+                             
                    |  +-- Hyriopsis_phuphaniensis **
                    +--+                          
                       +-- Hyriopsis_kratiensis **

Pfeiffer et al., 2021 [Publ 4126], fig. 27

iTtTbiriSoTroTroSiD
99592118
  +-------------- Rectidens              
  |                                      
  |           +-- Hyriopsis_delaportei **   
--+  +--------+                          
  |  |        +-- Hyriopsis_desowitzi **    
  |  |                                   
  +--+  +-------- Hyriopsis_khoratensis **  
     |  |                                
     |  |     +-- Hyriopsis_kratiensis **   
     |  +-----+                          
     +--+     +-- Hyriopsis_phuphaniensis **
        |                                
        |  +----- Hyriopsis_sakhonensis **  
        |  |                             
        +--+  +-- Hyriopsis_myersiana **    
           |  |                          
           +--+-- Hyriopsis_panhai **       
              |                          
              +-- Hyriopsis_bialata **

Jeratthitikul et al., 2021 [Publ 4212], fig. 4

iTtTbiriSoTroTroSiD
8978133212
  +----------------- Ensidens               
  |                                         
  |              +-- Hyriopsis_desowitzi **    
--+  +-----------+                          
  |  |           +-- Hyriopsis_delaportei **   
  |  |                                      
  +--+  +----------- Rectidens              
     |  |                                   
     |  |     +----- Hyriopsis_myersiana **    
     +--+  +--+                             
        |  |  |  +-- Hyriopsis_bialata **      
        |  |  +--+                          
        +--+     +-- Hyriopsis_panhai **       
           |                                
           |  +----- Hyriopsis_khoratensis **  
           +--+                             
              |  +-- Hyriopsis_phuphaniensis **
              +--+                          
                 +-- Hyriopsis_sakhonensis **

Pfeiffer et al., 2021 [Publ 4126], fig. 9

iTtTbiriSoTroTroSiD
6646237211
  +-------------------- Ensidens             
--+                                          
  |  +----------------- Rectidens            
  +--+                                       
     |  +-------------- Hyriopsis_delaportei ** 
     +--+                                    
        |  +----------- Hyriopsis_myersiana **  
        +--+                                 
           |  +-------- Hyriopsis_bialata **    
           +--+                              
              |  +----- Hyriopsis_khoratensis **
              +--+                           
                 |  +-- Hyriopsis_kratiensis ** 
                 +--+                        
                    +-- Hyriopsis_sakhonensis **

Bolotov et al., 2017 [Publ 3764], fig. S1

iTtTbiriSoTroTroSiD
5625119627
  +----------- Ensidens             
  |                                 
--+  +-------- Hyriopsis_desowitzi **  
  |  |                              
  |  +-------- Rectidens            
  +--+                              
     |  +----- Hyriopsis_panhai **     
     |  |                           
     +--+  +-- Hyriopsis_myersiana **  
        |  |                        
        +--+-- Hyriopsis_bialata **    
           |                        
           +-- Hyriopsis_khoratensis **

Konopleva et al., 2019 [Publ 3900], fig. 1

iTtTbiriSoTroTroSiD
563546626
  +-------------- Ensidens             
  |                                    
--+  +----------- Hyriopsis_desowitzi **  
  |  |                                 
  +--+----------- Rectidens            
     |                                 
     |  +-------- Hyriopsis_panhai **     
     +--+                              
        |  +----- Hyriopsis_myersiana **  
        +--+                           
           |  +-- Hyriopsis_khoratensis **
           +--+                        
              +-- Hyriopsis_bialata **

Bolotov et al., 2020 [Publ 4100], fig. 1

iTtTbiriSoTroTroSiD
5645112729
  +-------------- Ensidens             
  |                                    
--+  +----------- Hyriopsis_desowitzi **  
  |  |                                 
  +--+  +-------- Rectidens            
     |  |                              
     +--+     +-- Hyriopsis_khoratensis **
        |  +--+                        
        |  |  +-- Hyriopsis_bialata **    
        +--+                           
           |  +-- Hyriopsis_myersiana **  
           +--+                        
              +-- Hyriopsis_panhai **

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
511651956213
  +----- Hyriopsis_desowitzi **  
  |                           
  +----- Hyriopsis_myersiana **  
  |                           
  +----- Hyriopsis_panhai **     
--+                           
  |  +-- Hyriopsis_bialata **    
  +--+                        
  |  +-- Hyriopsis_khoratensis **
  |                           
  |  +-- Rectidens            
  +--+                        
     +-- Ensidens

Zieritz et al., 2016 [Publ 3804], fig. S2-1

iTtTbiriSoTroTroSiD
55251119
  +----------- Rectidens            
  |                                 
--+  +-------- Hyriopsis_desowitzi **  
  |  |                              
  +--+  +----- Hyriopsis_khoratensis **
     |  |                           
     +--+  +-- Hyriopsis_myersiana **  
        |  |                        
        +--+-- Hyriopsis_panhai **     
           |                        
           +-- Hyriopsis_bialata **

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
56451936211
  +-------------- Ensidens             
  |                                    
--+  +----------- Hyriopsis_desowitzi **  
  |  |                                 
  +--+  +-------- Rectidens            
     |  |                              
     +--+     +-- Hyriopsis_panhai **     
        |  +--+                        
        |  |  +-- Hyriopsis_khoratensis **
        +--+                           
           |  +-- Hyriopsis_bialata **    
           +--+                        
              +-- Hyriopsis_myersiana **

Konopleva et al., 2019 [Publ 4041], fig. 3

iTtTbiriSoTroTroSiD
51205297215
  +-- Ensidens_ingallsianus
  |                        
  +-- Ensidens_telus       
  |                        
  +-- Ensidens_dugasti     
  |                        
  +-- Hyriopsis_khoratensis **
  |                        
  +-- Hyriopsis_bialata **    
  |                        
  +-- Hyriopsis_myersiana **  
  |                        
--+-- Hyriopsis_desowitzi **  
  |                        
  +-- Rectidens            
  |                        
  +-- Hyriopsis_panhai **     
  |                        
  +-- Ensidens_jaculus__1  
  |                        
  +-- Ensidens_jaculus__2  
  |                        
  +-- Ensidens_spiculus    
  |                        
  +-- Ensidens_sagittarius

Bolotov et al., 2020 [Publ 4100], fig. S1

iTtTbiriSoTroTroSiD
5645106727
  +----------------- Ensidens             
--+                                       
  |  +-------------- Hyriopsis_desowitzi **  
  +--+                                    
     |  +----------- Rectidens            
     +--+                                 
        |  +-------- Hyriopsis_panhai **     
        +--+                              
           |  +----- Hyriopsis_myersiana **  
           +--+                           
              |  +-- Hyriopsis_khoratensis **
              +--+                        
                 +-- Hyriopsis_bialata **

Bolotov et al., 2017 [Publ 3761], fig. S1

iTtTbiriSoTroTroSiD
5645133629
  +-------------- Ensidens             
  |                                    
--+           +-- Hyriopsis_bialata **    
  |  +--------+                        
  |  |        +-- Hyriopsis_khoratensis **
  +--+                                 
     |  +-------- Hyriopsis_panhai **     
     +--+                              
        |  +----- Hyriopsis_myersiana **  
        +--+                           
           |  +-- Rectidens            
           +--+                        
              +-- Hyriopsis_desowitzi **

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
511551956213
  +----- Hyriopsis_desowitzi **  
  |                           
  +----- Rectidens            
  |                           
--+----- Ensidens             
  |                           
  |  +-- Hyriopsis_bialata **    
  |  |                        
  |  +-- Hyriopsis_myersiana **  
  +--+                        
     +-- Hyriopsis_panhai **     
     |                        
     +-- Hyriopsis_khoratensis **

Bolotov et al., 2017 [Publ 3761], fig. S2

iTtTbiriSoTroTroSiD
511751336213
        +-- Hyriopsis_desowitzi **  
        |                        
  +-----+-- Rectidens            
  |     |                        
  |     +-- Ensidens             
--+                              
  |  +----- Hyriopsis_panhai **     
  |  |                           
  +--+----- Hyriopsis_khoratensis **
     |                           
     |  +-- Hyriopsis_myersiana **  
     +--+                        
        +-- Hyriopsis_bialata **

Zieritz et al., 2020 [Publ 4102], fig. 4

iTtTbiriSoTroTroSiD
562526627
  +----------- Ensidens             
  |                                 
--+  +-------- Hyriopsis_desowitzi **  
  |  |                              
  |  +-------- Rectidens            
  +--+                              
     |  +----- Hyriopsis_panhai **     
     |  |                           
     +--+  +-- Hyriopsis_khoratensis **
        |  |                        
        +--+-- Hyriopsis_bialata **    
           |                        
           +-- Hyriopsis_myersiana **

Muanta et al., 2019 [Publ 3921], fig. 2

iTtTbiriSoTroTroSiD
44245519
  +----------- Ensidens             
--+                                 
  |  +-------- Hyriopsis_desowitzi **  
  +--+                              
     |  +----- Hyriopsis_myersiana **  
     +--+                           
        |  +-- Hyriopsis_khoratensis **
        +--+                        
           +-- Hyriopsis_panhai **

Bolotov et al., 2018 [Publ 3826], fig. 2

iTtTbiriSoTroTroSiD
3803795114
  +-- Ensidens_telus       
  |                        
  +-- Ensidens_ingallsianus
  |                        
  +-- Ensidens_jaculus     
  |                        
  +-- Hyriopsis_desowitzi **  
--+                        
  +-- Hyriopsis_myersiana **  
  |                        
  +-- Hyriopsis_khoratensis **
  |                        
  +-- Ensidens_spiculus    
  |                        
  +-- Ensidens_sagittarius

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iTtTbiriSoTroTroSiD
3723774211
  +----- Hyriopsis_desowitzi **  
  |                           
  +----- Rectidens            
--+                           
  +----- Ensidens             
  |                           
  |  +-- Hyriopsis_khoratensis **
  +--+                        
     +-- Hyriopsis_myersiana **

Zieritz et al., 2021 [Publ 4218], fig. 5

iTtTbiriSoTroTroSiD
22022922--
  +-------- Ensidens           
--+                            
  |  +----- Rectidens          
  +--+                         
     |  +-- Hyriopsis_bialata **  
     +--+                      
        +-- Hyriopsis_desowitzi **

Liu et al., 2024 [Publ 4380], fig. 5

iTtTbiriSoTroTroSiD
230234228
  +-------- Ensidens           
--+                            
  |  +----- Hyriopsis_bialata **  
  +--+                         
     |  +-- Hyriopsis_desowitzi **
     +--+                      
        +-- Rectidens

Wu et al., 2024 [Publ 4350], fig. 5a

iTtTbiriSoTroTroSiD
22023422--
  +-------- Ensidens           
--+                            
  |  +----- Rectidens          
  +--+                         
     |  +-- Hyriopsis_bialata **  
     +--+                      
        +-- Hyriopsis_desowitzi **

Wu et al., 2024 [Publ 4350], fig. 6-7

iTtTbiriSoTroTroSiD
22023422--
  +-------- Ensidens           
--+                            
  |  +----- Rectidens          
  +--+                         
     |  +-- Hyriopsis_bialata **  
     +--+                      
        +-- Hyriopsis_desowitzi **

Zieritz et al., 2024 [Publ 4381], fig. 2

iTtTbiriSoTroTroSiD
22024332--
  +-------- Ensidens           
--+                            
  |  +----- Rectidens          
  +--+                         
     |  +-- Hyriopsis_bialata **  
     +--+                      
        +-- Hyriopsis_desowitzi **

Wu et al., 2024 [Publ 4350], fig. 5b

iTtTbiriSoTroTroSiD
21302342218
  +----------------------- Hyriopsis_bialata **  
  |                                           
--+                    +-- Rectidens          
  |  +-----------------+                      
  |  |                 +-- Khairuloconcha     
  +--+                                        
     |  +----------------- Ensidens           
     |  |                                     
     +--+  +-------------- Hyriopsis_desowitzi **
        |  |                                  
        +--+  +----------- Yaukthwa_elongatula
           |  |                               
           +--+  +-------- Trapezoideus       
              |  |                            
              +--+     +-- Physunio           
                 |  +--+                      
                 |  |  +-- Yaukthwa_inlenensis
                 +--+                         
                    |  +-- Yaukthwa_paiensis  
                    +--+                      
                       +-- Lens

Wu et al., 2024 [Publ 4388], fig. 5

iTtTbiriSoTroTroSiD
22023422--
  +-------- Ensidens           
--+                            
  |  +----- Rectidens          
  +--+                         
     |  +-- Hyriopsis_bialata **  
     +--+                      
        +-- Hyriopsis_desowitzi **

Wu et al., 2024 [Publ 4350], fig. 5c

iTtTbiriSoTroTroSiD
21302342219
  +----- Ensidens           
  |                         
  +----- Hyriopsis_desowitzi **
  |                         
  +----- Rectidens          
  |                         
  +----- Hyriopsis_bialata **  
--+                         
  +----- Khairuloconcha     
  |                         
  |  +-- Physunio           
  |  |                      
  |  +-- Yaukthwa_paiensis  
  |  |                      
  |  +-- Yaukthwa_inlenensis
  +--+                      
     +-- Trapezoideus       
     |                      
     +-- Yaukthwa_elongatula
     |                      
     +-- Lens

Zieritz et al., 2026 [Publ 4812], fig. 7a

iTtTbiriSoTroTroSiD
1101331--
  +-- Hyriopsis_bialata **
--+                    
  +-- Rectidens

Jeratthitikul et al., 2021 [Publ 4219], fig. 2

iTtTbiriSoTroTroSiD
11013422--
  +----- Ensidens         
--+                       
  |  +-- Hyriopsis_bialata **
  +--+                    
     +-- Rectidens

Zieritz et al., 2021 [Publ 4231], fig. 3-M_4

iTtTbiriSoTroTroSiD
11012211--
  +-- Hyriopsis_bialata **
--+                    
  +-- Rectidens

Dai et al., 2023 [Publ 4285], fig. 3-ML

iTtTbiriSoTroTroSiD
11014411--
  +-- Hyriopsis_bialata **
--+                    
  +-- Ensidens

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

iTtTbiriSoTroTroSiD
11017522--
  +----- Ensidens         
--+                       
  |  +-- Rectidens        
  +--+                    
     +-- Hyriopsis_bialata **

Dai et al., 2025 [Publ 4440], fig. 3

iTtTbiriSoTroTroSiD
11018711--
  +-- Hyriopsis_bialata **
--+                    
  +-- Rectidens

Pfeiffer et al., 2018 [Publ 3827], fig. 1

iTtTbiriSoTroTroSiD
130262212
  +-- Ensidens    
  |               
  +-- Hyriopsis__1 **
--+               
  +-- Hyriopsis__2 **
  |               
  +-- Rectidens

Dai et al., 2025 [Publ 4441], fig. 1

iTtTbiriSoTroTroSiD
11014611--
  +-- Ensidens         
--+                    
  +-- Hyriopsis_bialata **

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

iTtTbiriSoTroTroSiD
110114011--
  +-- Hyriopsis_bialata **
--+                    
  +-- Rectidens

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

iTtTbiriSoTroTroSiD
11017522--
  +-- Ensidens         
  |                    
--+-- Hyriopsis_bialata **
  |                    
  +-- Rectidens

Jeratthitikul et al., 2021 [Publ 4219], fig. 1

iTtTbiriSoTroTroSiD
11013322--
  +----- Ensidens         
--+                       
  |  +-- Hyriopsis_bialata **
  +--+                    
     +-- Rectidens

Campbell & Lydeard, 2026 [Publ 4810], fig. 4

iTtTbiriSoTroTroSiD
11014611--
  +-- Hyriopsis_bialata **
--+                    
  +-- Rectidens

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iTtTbiriSoTroTroSiD
11019511--
  +-- Hyriopsis_bialata **
--+                    
  +-- Rectidens

Dai et al., 2024 [Publ 4389], fig. 3

iTtTbiriSoTroTroSiD
11014511--
  +-- Ensidens         
--+                    
  +-- Hyriopsis_bialata **

Chen et al., 2025 [Publ 4663], fig. 1

iTtTbiriSoTroTroSiD
11014711--
  +-- Ensidens         
--+                    
  +-- Hyriopsis_bialata **

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iTtTbiriSoTroTroSiD
11019511--
  +-- Hyriopsis_bialata **
--+                    
  +-- Rectidens

Chen et al., 2025 [Publ 4448], fig. 3

iTtTbiriSoTroTroSiD
11018511--
  +-- Hyriopsis_bialata **
--+                    
  +-- Rectidens

Dai et al., 2023 [Publ 4285], fig. 3-BI

iTtTbiriSoTroTroSiD
11014411--
  +-- Hyriopsis_bialata **
--+                    
  +-- Ensidens

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iTtTbiriSoTroTroSiD
11019111--
  +-- Hyriopsis_bialata **
--+                    
  +-- Rectidens

Matos et al., 2024 [Publ 4384], fig. 3

iTtTbiriSoTroTroSiD
11011911--
  +-- Hyriopsis_bialata **
--+                    
  +-- Rectidens

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iTtTbiriSoTroTroSiD
11015622--
  +----- Ensidens         
--+                       
  |  +-- Hyriopsis_bialata **
  +--+                    
     +-- Rectidens

Dai et al., 2023 [Publ 4285], fig. 5

iTtTbiriSoTroTroSiD
11015211--
  +-- Hyriopsis_bialata **
--+                    
  +-- Rectidens

Zhang et al., 2024 [Publ 4397], fig. 5

iTtTbiriSoTroTroSiD
11012611--
  +-- Hyriopsis_bialata **
--+                    
  +-- Rectidens

Wu et al., 2021 [Publ 4366], fig. 6

iTtTbiriSoTroTroSiD
11019031--
  +-- Hyriopsis_desowitzi **
--+                      
  +-- Ensidens

Jeratthitikul et al., 2024 [Publ 4387], fig. 1

iTtTbiriSoTroTroSiD
11015822--
  +----- Ensidens         
--+                       
  |  +-- Hyriopsis_bialata **
  +--+                    
     +-- Rectidens

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iTtTbiriSoTroTroSiD
11019111--
  +-- Hyriopsis_bialata **
--+                    
  +-- Rectidens

Matos et al., 2024 [Publ 4383], fig. 3

iTtTbiriSoTroTroSiD
11011911--
  +-- Hyriopsis_bialata **
--+                    
  +-- Rectidens

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

iTtTbiriSoTroTroSiD
11014711--
  +-- Hyriopsis_bialata **
--+                    
  +-- Rectidens

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iTtTbiriSoTroTroSiD
11015622--
  +----- Ensidens         
--+                       
  |  +-- Hyriopsis_bialata **
  +--+                    
     +-- Rectidens

Wu et al., 2024 [Publ 4388], fig. 6

iTtTbiriSoTroTroSiD
11015211--
  +-- Hyriopsis_bialata **
--+                    
  +-- Rectidens

Wu et al., 2026 [Publ 4811], fig. 4

iTtTbiriSoTroTroSiD
11019411--
  +-- Hyriopsis_bialata **
--+                    
  +-- Rectidens

Wu et al., 2021 [Publ 4366], fig. 2_I

iTtTbiriSoTroTroSiD
11019031--
  +-- Hyriopsis_desowitzi **
--+                      
  +-- Ensidens
 
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