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The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

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Publication Bolotov et al. (2018): attributed nominal species
publication summary | cladograms

Complete Citation

Bolotov, I.N., J.M. Pfeiffer, E.S. Konopleva, I.V. Vikhrev, A.V. Kondakov et al. 2018. A new genus and tribe of freshwater mussel (Unionidae) from Southeast Asia. Scientific Reports 8. 10030 (12 pp.). Available electronically.

Attributed Nominal Species

Anodontites elongata Whelan et al., 2011
‘Swainson’ of Authors.
Whelan et al., 2011. MPE 61: 506.
Graf et al., 2015. MPE 85: 3.
Bolotov et al., 2016. MPE 103: 108.
Lopes-Lima et al., 2017. MPE 106: 178.
Bolotov et al., 2017. Sci. Repts. 7: Table S1.
Bolotov et al., 2017. Sci. Repts. 7: Table S1.
Konopleva et al., 2017. Systemat. Biodiv. 15(3): 208.
Combosch et al., 2017. MPE 107: 195, Table 1.
Bolotov et al., 2018. Sci. Repts. 8: Table S1.
Huang et al., 2018. MPE 120: 302.
Lopes-Lima et al., 2018. MPE 127: 103.
Huang et al., 2019. MPE 130: Table S4.
Bolotov et al., 2019. Sci. Repts. 9: Table S1.
Pfeiffer et al., 2019. MPE 137: 116.
Guerra et al., 2019. BMC Evol. Biol. 19 (229): 3.
Bolotov et al., 2020. Sci. Repts. 10: Table S1.
Sano et al., 2020. Plankton Benthos Res. 15(2): 101.
Wu et al., 2020. Invert. Syst. 34: 195.
Wu et al., 2021. J. Lake Sci. 33(6): attached Table 1.
Chen et al., 2025. Zoosyst. Evol. 101(4): Table 1.
(misidentified reference to Anodon elongatus Swainson, 1823)

Contradens contradens Pfeiffer & Graf, 2015
‘Lea’ of Authors.
Pfeiffer & Graf, 2015. Zool. J. Linn. Soc. 175(2): 310.
Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘Contradens aff. contradens sp. 1’].
Konopleva et al., 2017. Systemat. Biodiv. 15(3): 207.
Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘Contradens aff. contradens sp. 1’].
Bolotov et al., 2018. Sci. Repts. 8: Table S1 [as ‘Contradens aff. contradens sp. 1’].
Konopleva et al., 2019. Ecologica Montenegrina 24: Appendix 1 [as ‘Contradens aff. contradens sp. 1’].
Konopleva et al., 2019. Sci. Repts. 9: Table S1 [as ‘Contradens aff. contradens sp. 1’].
Huang et al., 2019. MPE 130: Table S4 [as ‘Contradens aff. contradens sp. 1’].
Bolotov et al., 2020. Sci. Repts. 10: Table S1 [as ‘Contradens aff. contradens sp. 1’].
Konopleva et al., 2023. Diversity 15(1), 10: Table S2 [as ‘Lens aff. contradens’].
(misidentified reference to Unio contradens Lea, 1838)

Ensidens sagittarius Bolotov et al., 2017
‘Lea’ of Authors.
Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘Ensidens aff. sagittarius sp. 2’].
Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘Ensidens aff. sagittarius sp. 2’].
Bolotov et al., 2018. Sci. Repts. 8: Table S1 [as ‘E. aff. sagittarius sp. 2’].
Muanta et al., 2019. J. Moll. Studies 85(2): 227 [as ‘Clade B4’].
Konopleva et al., 2019. Sci. Repts. 9: Table S1 [as ‘E. aff. sagittarius sp. 2’].
Konopleva et al., 2019. Ecologica Montenegrina 24: Appendix 1 [as ‘E. aff. sagittarius sp. 2’].
Huang et al., 2019. MPE 130: Table S4 [as ‘Ensidens aff. sagittarius sp. 2’].
Zieritz et al., 2020. Biodivers. Conserv. 29: 2242 [as ‘Ensidens aff. sagittarius sp. 2’].
(misidentified reference to Unio sagittarius Lea, 1856)

Ensidens sagittarius Pfeiffer & Graf, 2015
‘Lea’ of Authors.
Pfeiffer & Graf, 2015. Zool. J. Linn. Soc. 175(2): 310.
Konopleva et al., 2017. Systemat. Biodiv. 15(3): 207.
Lopes-Lima et al., 2017. MPE 106: 177.
Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘Ensidens aff. sagittarius sp. 1].
Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘Ensidens aff. sagittarius sp. 1].
Bolotov et al., 2018. Sci. Repts. 8: Table S1 [as ‘Ensidens aff. sagittarius sp. 1].
Konopleva et al., 2019. Sci. Repts. 9: Table S1 [as ‘E. aff. sagittarius sp. 1].
Huang et al., 2019. MPE 130: Table S4 [as ‘Ensidens aff. sagittarius sp. 1].
Zieritz et al., 2020. Biodivers. Conserv. 29: 2242.
Lopes-Lima et al., 2020. MPE 146: Table 1.
Wu et al., 2021. J. Lake Sci. 33(6): attached Table 1.
(misidentified reference to Unio sagittarius Lea, 1856)

Indonaia lima Bolotov et al., 2017
‘Simpson’ of Authors.
Bolotov et al., 2017. Sci. Repts. 7: Table S1.
Bolotov et al., 2018. Sci. Repts. 8: Table S1.
Huang et al., 2019. MPE 130: Table S4.
Konopleva et al., 2019. Sci. Repts. 9: Table S1.
Jeratthitikul et al., 2019. Tropical Nat. Hist. 19(1): 25.
Bolotov et al., 2020. Sci. Repts. 10: Table S1.
Bolotov et al., 2022. Hindawi 6600359: Table S1.
(misidentified reference to Nodularia (sec. Radiatula) lima Simpson, 1900)

Lamellidens generosus Bolotov et al., 2017
‘Gould’ of Authors.
Bolotov et al., 2017. Sci. Repts. 7: Table S1.
Bolotov et al., 2017. Sci. Repts. 7: Table S1.
Bolotov et al., 2018. Sci. Repts. 8: Table S1.
Huang et al., 2019. MPE 130: Table S4.
(misidentified reference to Unio generosus Gould, 1847)

Lamellidens marginalis Bolotov et al., 2017
‘Lamarck’ of Authors.
Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘Lamellidens marginalis sp. 2’ and ‘sp. 3’].
Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘Lamellidens marginalis sp. 2’ and ‘sp. 3’].
Konopleva et al., 2017. Systemat. Biodiv. 15(3): 207. [in part]
Bolotov et al., 2018. Sci. Repts. 8: Table S1 [as ‘Lamellidens marginalis sp. 3’].
Huang et al., 2019. MPE 130: Table S4 [as ‘Lamellidens marginalis sp. 2’ and ‘sp. 3’].
Jeratthitikul et al., 2019. Tropical Nat. Hist. 19(1): 25.
Konopleva et al., 2020. Ecologica Montenegrina 27: 47.
Bolotov et al., 2020. Sci. Repts. 10: Table S1.
(misidentified reference to Unio marginalis Lamarck, 1819)

Lasmigona compressa Graf & Ó Foighil, 2000
‘Lea’ of Authors.
Graf & Ó Foighil, 2000. J. Moll. Studies 66: 160.
Graf & Cummings, 2006. Zool. J. Linn. Soc. 148: 348.
Hoeh et al., 2009. Malacologia 51(2): 310.
Jadhav & Jamkhedkar, 2009. Res. J. BioTech. 4(1): 43.
Whelan et al., 2011. MPE 61: 506.
Lopes-Lima et al., 2017. MPE 106: 177.
Bolotov et al., 2017. Sci. Repts. 7: Table S1.
Bolotov et al., 2017. Sci. Repts. 7: Table S1.
Bolotov et al., 2018. Sci. Repts. 8: Table S1.
Huang et al., 2019. MPE 130: Table S4.
Lopes-Lima et al., 2020. MPE 146: Table 1.
Wu et al., 2021. J. Lake Sci. 33(6): attached Table 1.
Jeratthitikul et al., 2022. Sci. Repts. 12: Table S1.
Wu et al., 2023. ZooKeys 1166: 263, Table 1.
Liu et al., 2023. ZooKeys 1179: Table 1.
Dai et al., 2025. Zoosyst. Evol. 101(2): Table 1. [in part]
Chen et al., 2025. Zoosyst. Evol. 101(4): Table 1.
(misidentified reference to Symphynota compressa Lea, 1829)

Nodularia jourdyi Bolotov et al., 2018
Bolotov et al., 2018. Sci. Repts. 8: 4.
Pfeiffer et al., 2019. MPE 137: 117.
(reference to Unio jourdyi Morlet, 1886)

Parreysia corrugata Bolotov et al., 2017
‘Müller’ of Authors.
Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘Parreysia corrugata sp. 2’]. [in part]
Bolotov et al., 2017. Sci. Repts. 7: Table S1. [in part]
Bolotov et al., 2018. Sci. Repts. 8: Table S1 [as ‘Parreysia corrygata sp. 2’].
Pfeiffer et al., 2018. J. Moll. Studies 84(4): Table 1. [in part]
Huang et al., 2019. MPE 130: Table S4 [as ‘Parreysia corrugata sp. 2’].
Bolotov et al., 2020. Sci. Repts. 10: Table S1 [as ‘Parreysia corrugata sp. 3’].
Bolotov et al., 2020. Sci. Repts. 10: Table S1 [as ‘Parreysia corrugata sp. 2’].
Imsong & Murali, 2023. Uttar Pradesh J. Zool. 44(20): Table 1.
(misidentified reference to Mya corrugata Müller, 1774)

Parreysia corrugata Bolotov et al., 2017
‘Müller’ of Authors.
Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘Parreysia corrugata sp. 1’].
Bolotov et al., 2018. Sci. Repts. 8: Table S1 [as ‘Parreysia corrugata sp. 1’].
Pfeiffer et al., 2018. J. Moll. Studies 84(4): Table 1. [in part]
Huang et al., 2019. MPE 130: Table S4 [as ‘Parreysia corrugata sp. 1’].
Jeratthitikul et al., 2019. Tropical Nat. Hist. 19(1): 25.
Bolotov et al., 2020. Sci. Repts. 10: Table S1 [as ‘Parreysia corrugata sp. 1’].
(misidentified reference to Mya corrugata Müller, 1774)

Pilsbryoconcha exilis Pfeiffer & Graf, 2015
‘Lea’ of Authors.
Pfeiffer & Graf, 2015. Zool. J. Linn. Soc. 175(2): 310.
Ng et al., 2016. PLoS One 11(8), e0161130: [5]. [in part]
Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘P. aff. exilis sp. 1’].
Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘P. aff. exilis sp. 1’].
Konopleva et al., 2017. Systemat. Biodiv. 15(3): 207.
Bolotov et al., 2018. Sci. Repts. 8: Table S1 [as ‘P. aff. exilis sp. 1’].
Huang et al., 2019. MPE 130: Table S4 [as ‘P. aff. exilis sp. 1’].
Konopleva et al., 2019. Sci. Repts. 9: Table S1 [as ‘P. aff. exilis sp. 1’].
Lopes-Lima et al., 2020. MPE 146: Table 1 [as ‘P. aff. exilis sp. 1’].
Zieritz et al., 2020. Biodivers. Conserv. 29: 2244.
Wu et al., 2021. ZooKeys 1054: 87.
Wu et al., 2021. J. Lake Sci. 33(6): attached Table 1.
Wu et al., 2021. J. Lake Sci. 33(6): attached Table 1.
Konopleva et al., 2023. Diversity 15(1), 10: Table S2 [as ‘P. aff. exilis’].
Dai et al., 2023. Invert. Syst. 37(2): Tables 1-2. [in part]
Dai et al., 2024. Zoosyst. Evol. 100(4): Table 2.
Chen et al., 2025. Cathaica 1(7): Table 1.
Dai et al., 2025. Zoosyst. Evol. 101(2): Table 1.
(misidentified reference to Anodonta exilis Lea, 1838)
Type(s): SEQUENCED SPECIMEN UMMZ_304647, Beung Dang Teuk Leach Community Fishery Pursat Province, Cambodia.

Pseudodon avae Bolotov et al., 2017
‘Theobald’ of Authors.
Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘Pseudodon cf. avae’].
Bolotov et al., 2017. Sci. Repts. 7: Table S1.
Bolotov et al., 2018. Sci. Repts. 8: Table S1.
Konopleva et al., 2019. Ecologica Montenegrina 24: Appendix 1 [as ‘Pseudodon cf. avae’].
Huang et al., 2019. MPE 130: Table S4.
Konopleva et al., 2019. Sci. Repts. 9: Table S1.
Zieritz et al., 2020. Biodivers. Conserv. 29: 2244.
Lopes-Lima et al., 2020. MPE 146: Table 1.
Dai et al., 2023. Invert. Syst. 37(2): 155, Table 1.
(misidentified reference to Monocondylaea avae Theobald, 1873)

Pseudodon cambodjensis Pfeiffer & Graf, 2013
‘Petit’ of Authors.
Pfeiffer & Graf, 2013. J. Moll. Studies 79: 251.
Pfeiffer & Graf, 2015. Zool. J. Linn. Soc. 175(2): 310.
Zieritz et al., 2016. Sci. Total Environ. 571: 1071.
Bolotov et al., 2017. Sci. Repts. 7: Table S1.
Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘Monodontina cambodjensis’].
Lopes-Lima et al., 2017. MPE 106: 177.
Konopleva et al., 2017. Systemat. Biodiv. 15(3): 207.
Bolotov et al., 2018. Sci. Repts. 8: Table S1 [as ‘Monodontina cambodjensis’].
Razak et al., 2018. Conserv. Genet. Resour. 11: 459.
Huang et al., 2019. MPE 130: Table S4 [as ‘Monodontina cambodjensis’].
Konopleva et al., 2019. Sci. Repts. 9: Table S1 [as ‘Monodontina cambodjensis’].
Bolotov et al., 2020. Sci. Repts. 10: Table S1 [as ‘Monodontina cambodiensis’ (sic)].
Konopleva et al., 2021. Sci. Repts. 11: Table S1 [as ‘Monodontina cambodiensis’ (sic)].
Jeratthitikul et al., 2021. Eur. J. Taxonomy 775: 123 [as ‘Monodontina cambodiensis’ (sic)].
Pfeiffer et al., 2021. Invert. Syst. 35: 453 [as ‘Monodontina cambodjensis’].
Wu et al., 2021. ZooKeys 1054: 87 [as ‘Monodontina cambodjensis’].
Wu et al., 2021. J. Lake Sci. 33(6): attached Table 1.
Konopleva et al., 2023. Diversity 15(1), 10: Table S2 [as ‘Monodontina cambodjensis’].
Dai et al., 2023. Invert. Syst. 37(2): Table 1 [as ‘Monodontina cambodjensis’].
Dai et al., 2024. Zoosyst. Evol. 100(4): Table 2.
Dai et al., 2025. Zoosyst. Evol. 101(2): Table 1.
(reference to Monocondylea cambodjensis Petit de la Saussaye, 1865)
Type(s): SEQUENCED SPECIMEN UMMZ_304350, Pursat River, just downstream (north) of Highway 5 bridge, Pursat City, Cambodia.

Pseudodon mouhotii Pfeiffer & Graf, 2015
‘Lea’ of Authors.
Pfeiffer & Graf, 2015. Zool. J. Linn. Soc. 175(2): 310.
Lopes-Lima et al., 2017. MPE 106: 177.
Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘Pseudodon aff. mouhotii sp. 3’].
Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘Bineurus aff. mouhotii sp. 3’].
Konopleva et al., 2017. Systemat. Biodiv. 15(3): 207.
Bolotov et al., 2018. Sci. Repts. 8: Table S1 [as ‘Bineurus aff. mouhotii sp. 3’].
Konopleva et al., 2019. Sci. Repts. 9: Table S1 [as ‘Bineurus aff. mouhotii sp. 3’].
Huang et al., 2019. MPE 130: Table S4 [as ‘Bineurus aff. mouhotii sp. 3’].
(misidentified reference to Monocondyloea mouhotii Lea, 1863)

Pseudodon mouhotii Bolotov et al., 2017
‘Lea’ of Authors.
Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘Pseudodon aff. mouhotii sp. 2’].
Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘Bineurus aff. mouhotii sp. 2’].
Bolotov et al., 2018. Sci. Repts. 8: Table S1 [as ‘Bineurus aff. mouhotii sp. 2’].
Konopleva et al., 2019. Sci. Repts. 9: Table S1 [as ‘Bineurus aff. mouhotii sp. 2’].
Huang et al., 2019. MPE 130: Table S4 [as ‘Bineurus aff. mouhotii sp. 2’].
(misidentified reference to Monocondyloea mouhotii Lea, 1863)

Pseudodon vondembuschianus Pfeiffer & Graf, 2015
‘Lea’ of Authors.
Pfeiffer & Graf, 2015. Zool. J. Linn. Soc. 175(2): 310. [in part]
Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘Pseudodon aff. vondembuschianus sp. 2’]. [in part]
Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘Monodontina aff. vondembuschiana sp. 2’]. [in part]
Konopleva et al., 2017. Systemat. Biodiv. 15(3): 207. [in part]
Bolotov et al., 2018. Sci. Repts. 8: Table S1 [as ‘Monodontina aff. vondembuschiana sp. 2’]. [in part]
Huang et al., 2019. MPE 130: Table S4 [as ‘Monodontina aff. vondembuschiana sp. 2’]. [in part]
Konopleva et al., 2019. Sci. Repts. 9: Table S1 [as ‘Monodontina aff. vondembuschiana sp. 2’]. [in part]
(reference to Margaritana vondembuschiana Lea, 1840)

Pseudodon vondembuschianus Graf, 2002
‘Lea’ of Authors.
Graf, 2002. J. Moll. Studies 68: 66.
Graf & Cummings, 2006. Zool. J. Linn. Soc. 148: 348.
Hoeh et al., 2009. Malacologia 51(2): 310.
Whelan et al., 2011. MPE 61: 506. [in part]
Pfeiffer & Graf, 2015. Zool. J. Linn. Soc. 175(2): 310. [in part]
Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘Monodontina aff. vondembuschiana sp. 2’]. [in part]
Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘Pseudodon aff. vondembuschianus sp. 2’]. [in part]
Konopleva et al., 2017. Systemat. Biodiv. 15(3): 207. [in part]
Bolotov et al., 2018. Sci. Repts. 8: Table S1 [as ‘Monodontina aff. vondembuschiana sp. 2’]. [in part]
Huang et al., 2019. MPE 130: Table S4 [as ‘Monodontina aff. vondembuschiana sp. 2’]. [in part]
Konopleva et al., 2019. Sci. Repts. 9: Table S1 [as ‘Monodontina aff. vondembuschiana sp. 2’]. [in part]
(reference to Margaritana vondembuschiana Lea, 1840)

Pseudodon vondembuschianus Bolotov et al., 2017
‘Lea’ of Authors.
Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘Pseudodon aff. vondembuschianus sp. 1’].
Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘Monodontina aff. vondembuschiana sp. 1’].
Bolotov et al., 2018. Sci. Repts. 8: Table S1 [as ‘Monodontina aff. vondembuschiana sp. 1’].
Huang et al., 2019. MPE 130: Table S4 [as ‘Monodontina aff. vondembuschiana sp. 1’].
Konopleva et al., 2019. Sci. Repts. 9: Table S1 [as ‘Monodontina aff. vondembuschiana sp. 1’].
Lopes-Lima et al., 2020. MPE 146: Table 1 [as ‘Monodontina aff. vondembuschiana sp. 1’].
(misidentified reference to Margaritana vondembuschiana Lea, 1840)

Radiatula bonneaudii Whelan et al., 2011
‘Eydoux’ of Authors.
Whelan et al., 2011. MPE 61: 506.
Pfeiffer & Graf, 2015. Zool. J. Linn. Soc. 175(2): [4].
Bolotov et al., 2017. Sci. Repts. 7: 10, Table 3 [as ‘sp. 1’].
Konopleva et al., 2017. Systemat. Biodiv. 15(3): 207.
Bolotov et al., 2017. Sci. Repts. 7: [5].
Bolotov et al., 2018. Sci. Repts. 8: Table S1.
Pfeiffer et al., 2018. J. Moll. Studies 84(4): [4]. [in part]
Jeratthitikul et al., 2019. Tropical Nat. Hist. 19(1): 25.
Konopleva et al., 2019. Sci. Repts. 9: Table S1 [as ‘Radiatula bonneaudii sp. 1’].
Huang et al., 2019. MPE 130: Table S4.
Ortiz-Sepulveda et al., 2020. MPE 148: Table S1.
(misidentified reference to Unio bonneaudii Eydoux, 1838)

Radiatula humilis Bolotov et al., 2017
‘Lea’ of Authors.
Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘Radiatula humilis sp. 3].
Bolotov et al., 2018. Sci. Repts. 8: Table S1 [as ‘Radiatula humilis sp. 3].
Huang et al., 2019. MPE 130: Table S4 [as ‘Radiatula humilis sp. 3’].
Konopleva et al., 2019. Sci. Repts. 9: Table S1 [as ‘Radiatula humilis sp. 3’].
Ortiz-Sepulveda et al., 2020. MPE 148: Table S1. [in part]
(misidentified reference to Unio humilis Lea, 1856)

Radiatula khadakvaslaensis Bolotov et al., 2017
‘Ray’ of Authors.
Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘Radiatula khadakvaslaensis sp. 1’].
Bolotov et al., 2017. Sci. Repts. 7: Table S1 [as ‘Indonaia khadakvaslaensis sp. 1’].
Bolotov et al., 2018. Sci. Repts. 8: Table S1 [as ‘Indonaia khadakvaslaensis sp. 1’].
Huang et al., 2019. MPE 130: Table S4 [as ‘Indonaia khadakvaslaensis sp. 1’].
Konopleva et al., 2019. Sci. Repts. 9: Table S1 [as ‘Indonaia khadakvaslaensis sp. 1’].
Bolotov et al., 2020. Sci. Repts. 10: Table S1 [as ‘Indonaia khadakvaslaensis’].
Bolotov et al., 2022. Hindawi 6600359: Table S1 [as ‘Indonaia khadakvaslaensis’].
(reference to Indonaia khadakvaslaensis Ray, 1966)

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