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Page last updated
Thu 17 July 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

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Publication Pfeiffer et al. (2018): cladograms
     publication summary | cladograms

Complete Citation

Pfeiffer, J.M., D.L. Graf, K.S. Cummings & L.M. Page. 2018. Molecular phylogeny and taxonomic revision of two enigmatic freshwater mussel genera (Bivalvia: Unionidae incertae sedis: Harmandia and Unionetta) reveals a diverse clade of southeast Asia Parreysiinae. Journal of Molluscan Studies 84(4): 404-416. Available electronically.

Cladogram Figures

(click the figure number to see the cladogram)

Cladogram Figure 1
     +----- Physunio       
     |                     
  +--+----- Trapezoideus   
  |  |                     
  |  |  +-- Lens_comptus   
  |  +--+                  
--+     +-- Lens_contradens
  |                        
  |     +-- Ensidens       
  |     |                  
  |     +-- Hyriopsis__1   
  +-----+                  
        +-- Hyriopsis__2   
        |                  
        +-- Rectidens      
Cladogram Figure 2
  +----------------------------------------------- Margaritifera_margaritifera  
  |                                                                             
  |                                            +-- Trapezidens_dolichorhynchus  
--+  +-----------------------------------------+                                
  |  |                                         +-- Lamellidens_savadiensis      
  |  |                                                                          
  |  |                                      +----- Parreysia_keralaensis        
  +--+  +-----------------------------------+                                   
     |  |                                   |  +-- Parreysia_nagpoorensis       
     |  |                                   +--+                                
     |  |                                      +-- Parreysia_plagiosoma         
     |  |                                                                       
     +--+                                      +-- Leoparreysia_tavoyensis      
        |  +-----------------------------------+                                
        |  |                                   +-- Leoparreysia_olivacea        
        |  |                                                                    
        |  |                                +----- Nitia_teretiuscula           
        +--+  +-----------------------------+                                   
           |  |                             |  +-- Coelatura_aegyptiaca         
           |  |                             +--+                                
           |  |                                +-- Prisodontopsis_aviculaeformis
           +--+                                                                 
              |                             +----- Indonaia_bonneaudii          
              |  +--------------------------+                                   
              |  |                          |  +-- Indonaia_andersoniana        
              |  |                          +--+                                
              +--+                             +-- Indonaia_caerulea            
                 |                                                              
                 |  +----------------------------- Indochinella_pugio           
                 |  |                                                           
                 +--+                          +-- Radiatula_chaudhurii         
                    |  +-----------------------+                                
                    |  |                       +-- Radiatula_myitkyinae         
                    +--+                                                        
                       |  +----------------------- Scabiellus_songkramensis     
                       |  |                                                     
                       +--+  +-------------------- Unionetta_fabagina__2        
                          |  |                                                  
                          +--+  +----------------- Harmandia_somboriensis       
                             |  |                                               
                             +--+  +-------------- Unionetta_fabagina__1        
                                |  |                                            
                                +--+           +-- Scabies_crispata             
                                   |  +--------+                                
                                   |  |        +-- Scabies_mandarinus           
                                   +--+                                         
                                      |  +-------- Scabies_nucleus              
                                      |  |                                      
                                      +--+     +-- Scabies_pilata               
                                         |  +--+                                
                                         |  |  +-- Scabies_humilis              
                                         +--+                                   
                                            |  +-- Scabies_phaselus             
                                            |  |                                
                                            +--+-- Scabies_anceps               
                                               |                                
                                               +-- Scabies_scobinatus           
 
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