The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)
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Publication Huang et al. (2019): cladograms
publication summary | cladograms Complete CitationHuang, X.-C., J.-H. Su, J.-X. Ouyang, S. Ouyang, C.-H. Zhou & X.-P. Wu. 2019. Towards a global phylogeny of freshwater mussels (Bivalivia: Unionida): Species delimitation of Chinese taxa, mitochondrial phylogenomics, and diversification patterns. Molecular Phylogenetics and Evolution 130: 45-59. Available electronically.
Cladogram Figures(click the figure number to see the cladogram) Cladogram Figure 1
+-- Margaritifera_dahurica
+-----------------------------------------+
| +-- Gibbosula_rochechouartii
|
| +----- Lamprotula_leaii
| |
| +-----------+ +-- Lamprotula_caveata__1
| | | |
| | +--+-- Lamprotula_cornuumlunae
| | |
--+ +-----------------------+ +-- Lamprotula_caveata__2
| | |
| | | +-- Parvasolenaia_rivularis
| | | +-----------+
| | | | +-- Parvasolenaia_triangularis
| | +--+
| | | +----------- Ptychorhynchus_pfisteri
| | +--+
| | | +-------- Sinosolenaia_carinata
| | +--+
+--+ | +----- Sinosolenaia_recognita
| +--+
| | +-- Sinosolenaia_oleivora
| +--+
| +-- Sinosolenaia_iridinea
|
| +----- Sinohyriopsis_cumingii__1
| +--------------------------------+
| | | +-- Sinohyriopsis_cumingii__2
| | +--+
| | +-- Sinohyriopsis_schlegelii
+--+
| +----------------------------------- Anemina_harlandi
| |
| | +-------------------------------- Acuticosta_chinensis
| | |
+--+ | +-- Sinanodonta_angula
| | +--------+
| | | +-- Sinanodonta_lucida
| | +-----------------+
| | | | +-------- Sinanodonta_jourdyi
+--+ | +--+
| | | +----- Sinanodonta_woodiana
| | +--+
| | | +-- Sinanodonta_elliptica
| | +--+
| | +-- Sinanodonta_pacifica
| |
+--+ +----------- Scriptolamprotula_tortuosa
| +--------------+
| | | +-------- Scriptolamprotula_scripta
| | +--+
| | | +----- Scriptolamprotula_coreana
| | +--+
| | | +-- Scriptolamprotula_fibrosa
| | +--+
| | +-- Scriptolamprotula_tientsinensis
+--+
| +----------- Cristaria_plicata
| +-----------+
| | | +-------- Lanceolaria_gladiola
| | +--+
| | | +----- Lanceolaria_grayii
| | +--+
| | | +-- Lanceolaria_lanceolata
+--+ +--+
| +-- Lanceolaria_triformis
|
| +-------------------- Lepidodesma_languilati
| |
| | +-------- Schistodesmus_spinosus
| | +--------+
+--+ | | +----- Schistodesmus_tongpenensis
| | +--+
| | | +-- Schistodesmus_xinyuensis
| | +--+
+--+ +-- Schistodesmus_lampreyanus
|
| +----- Nodularia_douglasiae
| +--------+
| | | +-- Nodularia_huana
| | +--+
+--+ +-- Nodularia_nuxpersicae
|
| +----------- Cuneopsis_rufescens
+--+
| +-------- Tchangsinaia_piscicula
+--+
| +----- Cuneopsis_celtiformis
+--+
| +-- Cuneopsis_szechenyii
+--+
+-- Cuneopsis_heudei
Cladogram Figure 2
+-- Neotrigonia_margaritacea
+--------------------------------------------------+
| +-- Neotrigonia_lamarckii
|
| +----- Alathyria_jacksoni
| +--+
| | | +-- Alathyria_profuga
| | +--+
| +--------+ +-- Velesunio_ambiguus
| | |
| | | +----- Alathyria_pertexta
| | +--+
| | | +-- Lortiella_froggatti
| +--------------------------------+ +--+
| | | +-- Microdontia_anodontaeformis
| | |
| | | +-------------- Westralunio_carteri
| | | |
| | +--+ +----------- Echyridella_menziesii
| | | |
| | | | +----- Diplodon_suavidicus
| | +--+ +--+
--+ | | | | +-- Castalia_ambigua
| | | | +--+
| | +--+ +-- Triplodon_corrugatus
| | |
| | | +----- Hyridella_australis
| | +--+
| | | +-- Hyridella_depressa
| | +--+
| | +-- Hyridella_drapeta
| |
| | +-------- Anodontites_trapesialis
| | +--+
| | | | +----- Mutela_rostrata
| | | +--+
| | | | +-- Mutela_dubia
| | | +--+
| | +-----------------------------------+ +-- Mutela_hargeri
| | | |
| | | | +----- Mycetopoda_siliquosa
| | | | +--+
| | | | | | +-- Tamsiella_schroeteriana
+--+ | | | +--+
| | +--+ +-- Lamproscapha_ensiformis
| | |
| | | +----- Etheria_elliptica
| | +--+
| | | +-- Aspatharia_pfeifferiana
| | +--+
| | +-- Chambardia_wahlbergi
| |
| | +----------------- Gibbosula_rochechouartii
| | |
| | | +-------------- Gibbosula_laosensis
| | +--------------------------+ |
| | | | | +----------- Cumberlandia_monodonta
| | | | | |
| | | +--+ | +----- Pseudunio_marocanus
| | | | +-----+
| | | | | | +-- Pseudunio_auricularius
| | | | | +--+
| | | +--+ +-- Pseudunio_homsensis
| | | |
| | | | +-- Margaritifera_dahurica
| | | | +-----+
| | | | | +-- Margaritifera_margaritifera
| | | | |
| | | +--+ +----- Margaritifera_laevis
| | | | |
| | | | | +-- Margaritifera_falcata
| | | +--+--+
| | | | +-- Margaritifera_middendorffi
+--+ | |
| | | +-- Margaritifera_hembeli
| | +--+
| | +-- Margaritifera_marrianae
| |
| | +----- Trapezidens_exolescens
| | +-----------------+
| | | | +-- Trapezidens_dolichorhynchus
| | | +--+
| | +--+ +-- Trapezidens_angustior
| | | |
| | | | +-------------------- Lamellidens_corrianus
| | | | |
| | | +--+ +----------------- Lamellidens_savadiensis
| | | | |
| | | +--+ +-- Lamellidens_lamellatus
| | | | +-----------+
| | | | | +-- Lamellidens_jenkinsianus
| | | +--+
| | | | +----------- Lamellidens_generosus
| | | +--+
| | | | +-------- Lamellidens_mainwaringi__2
| | | +--+
| | +--------------+ | +----- Lamellidens_mainwaringi__1
| | | | +--+
| | | | | +-- Lamellidens_ferrugineus
| | | | +--+
| | | | +-- Lamellidens_marginalis
| | | |
| | | | +----- Parreysia_keralaensis
| | | | +-----------------+
| | | | | | +-- Parreysia_nagpoorensis
| | | | | +--+
| | | | | +-- Parreysia_plagiosoma
| | | | |
+--+ | | | +-- Leoparreysia_tavoyensis
| | | | |
| | +--+ +-----------------+-- Leoparreysia_canefrii
| | | | |
| | | | +-- Leoparreysia_olivacea
| | | |
| | | | +-------- Coelatura_aegyptiaca__2
| | | | |
| | | | +--------+ +----- Coelatura_aegyptiaca__1
| | +--+ | | |
| | | | | +----- Nitia_teretiuscula
| | | | +--+
| | | | | +-- Coelatura_gabonensis
| | | | | |
| | | | +--+-- Coelatura_kunenensis
| | | | |
| | +--+ +-- Prisodontopsis_mweruensis
| | |
| | | +-------- Indonaia_shurtleffiana
| | | |
| | | +-----+ +----- Indonaia_bonneaudii
| | | | | |
| | | | +--+----- Indonaia_rugosa
| | | | |
| | | | | +-- Indonaia_andersoniana
| | +--+ +--+
| | | +-- Indonaia_caerulea
| | |
| | | +----------- Indochinella_pugio
| | | |
| | | | +----- Scabies_humilis
| | +--+ +--+
| | | | | +-- Scabies_phaselus
| | | | +--+
| | +--+ +-- Scabies_mandarinus
| | |
| | | +----- Radiatula_myitkyinae
| | +--+
| | | +-- Radiatula_chaudhurii
| | +--+
| | +-- Radiatula_mouhoti
| |
| | +-------------------------------- Acuticosta_chinensis
| | |
| | | +----------------------------- Lepidodesma_languilati
| | | |
| | | | +-------- Scriptolamprotula_tortuosa
| | | | |
| | | | +--------------+ +----- Scriptolamprotula_coreana
| | | | | | |
| | | | | +--+----- Scriptolamprotula_scripta
| | | | | |
| | | | | | +-- Scriptolamprotula_fibrosa
| | | | | +--+
+--+ | | | +-- Scriptolamprotula_tientsinensis
| | | |
| | +-----+ +-------- Unio_tumidus
| +-----+ | | +-----------+
| | | | | | | +----- Unio_crassus
| | | | | | +--+
| | | | | | | +-- Unio_delphinus
| | | | | | +--+
| | | | | | +-- Unio_pictorum
| | | | | |
| | | | +--+ +----- Schistodesmus_spinosus
| | | | | |
| | | | | +-----------+ +-- Schistodesmus_xinyuensis
| | | | | | | |
| | | | | | +--+-- Schistodesmus_tongpenensis
| | | | | | |
| | | | | | +-- Schistodesmus_lampreyanus
| | | | +--+
| | +--+ | +----- Nodularia_douglasiae
| | | | +--------+
| | | | | | +-- Nodularia_huana
| | | | | +--+
| | | +--+ +-- Nodularia_nuxpersicae
| | | |
| | | | +----------- Tchangsinaia_piscicula
| | | +--+
| | | | +-------- Cuneopsis_rufescens
| | | +--+
| | | | +----- Cuneopsis_celtiformis
| | | +--+
| | | | +-- Cuneopsis_szechenyii
| | | +--+
| | | +-- Cuneopsis_heudei
| | |
| | | +-------- Lanceolaria_gladiola
| | | |
| | | +-----------------+ +----- Lanceolaria_grayii__1
| | | | | |
| | | | +--+----- Lanceolaria_grayii__2
| | | | |
| | | | | +-- Lanceolaria_lanceolata
| | | | +--+
| | | | +-- Lanceolaria_triformis
| | | |
| | | | +-------------------- Anemina_harlandi
| | | | |
| | | | +--+ +----------------- Cristaria_plicata
| | +--+ | | |
| | | | +--+ +-------------- Pletholophus_tenuis
| | | | | |
| | | | +--+ +-- Sinanodonta_angula
| | | | | +--------+
| | | | | | +-- Sinanodonta_lucida
| | | | +--+
| | | | | +-------- Sinanodonta_woodiana
| | | | +--+
| | | | | +----- Sinanodonta_jourdyi
| | | | +--+
| | +--+ | +-- Sinanodonta_elliptica
| | | +--+
| | | +-- Sinanodonta_pacifica
| | |
| | | +-------- Alasmidonta_marginata
| | | |
| | | +-------- Lasmigona_costata
| | | |
| | | +-------- Pseudanodonta_complanata
+--+ | |
| | +-------- Simpsonaias_ambigua
| | |
| +--------------+-------- Strophitus_undulatus
| |
| | +----- Anodonta_anatina
| +--+
| | | +-- Anodonta_cygnea
| | +--+
| | +-- Brachyanodon_nuttalliana
| |
| | +----- Pyganodon_grandis
| +--+
| | +-- Utterbackia_imbecillis
| +--+
| +-- Utterbackia_peninsularis
|
| +----------- Hyriopsis_desowitzi
| |
| +----------- Rectidens_sumatrensis
| |
| | +-- Hyriopsis_bialata
| | |
| | +-- Hyriopsis_myersiana
| +-----+--------+
| | | +-- Hyriopsis_panhai
| | | |
| | | +-- Hyriopsis_khoratensis
| | |
| | | +-- Ensidens_ingallsianus
| | | +-----+
| | | | +-- Ensidens_telus
| | +--+
| | | +----- Ensidens_sagittarius
| +-----------------+ +--+
| | | | +-- Ensidens_spiculus
| | | +--+
| | | +-- Ensidens_jaculus
| | |
| | | +----- Physunio
| | | +--------+
| | | | | +-- Physunio_modelli
| | | | +--+
| | | | +-- Physunio_superbus
| | | |
| | +--+ +-------- Yaukthwa_inlenensis
| | | +--+
| | | | | +----- Yaukthwa_panhai
| | | | +--+
| | | | | +-- Yaukthwa_nesemanni
| | | | +--+
| | +--+ +-- Yaukthwa_rectangularis
| | |
| | | +-- Lens_contradens
| | | +-----+
| | | | +-- Lens_micropterus
| | +--+
| | | +----- Lens_eximius
| | +--+
| | | +-- Lens_comptus
| | +--+
| | +-- Lens_novoselovi
| |
| | +-- Megalonaias_nervosa
| | +--+
+--+ | +-- Uniomerus_carolinianus
| +-----------------------+
| | | +-- Quadrula_quadrula
| | +--+
| | +-- Quadrula_verrucosa
| |
| | +-------- Elliptoideus_sloatianus
| +--+ |
| | | +-----------------+ +-- Elliptio_complanata
| | | | | +--+
| | | | | | +-- Eurynia_dilatata
| | | | +--+
| | | | | +-- Fusconaia_flava
| | +--+ +--+
| | | +-- Pleurobema_sintoxia
| | |
| | | +----------------------- Amblema_plicata
| | | |
| | | | +----- Toxolasma_parvum
| | +--+ +--------------+
| | | | | +-- Obliquaria_reflexa
| | | | +--+
| | | | +-- Plectomerus_dombeyanus
| | +--+
| | | +----------------- Potamilus_leptodon
| | | |
| | | | +-------------- Potamilus_alatus
| | +--+ |
| | | | +----------- Venustaconcha_ellipsiformis
| | | | |
| | +--+ +----------- Leaunio_lienosus
| | | |
+--+ | | +-- Ellipsaria_lineolata
| | +--------+
| +--+ +-- Truncilla_truncata
| |
| | +-- Medionidus_conradicus
| +--------+
| | +-- Ptychobranchus_subtentus
| |
| | +-------- Cambarunio_iris
| +--+
| | +----- Ortmanniana_ligamentina
| +--+
| | +-- Lampsilis_cardium
| +--+
| +-- Lampsilis_ornata
|
| +----- Chamberlainia_somsakpanhai
| +-----------------------+
| | | +-- Sinohyriopsis_cumingii
| | +--+
| | +-- Sinohyriopsis_schlegelii
| |
| | +-- Potomida_littoralis
| | +-----+
| | | +-- Pronodularia_japanensis
| | +-----------------+
+--+ | | +----- Lamprotula_leaii
| | +--+
| | | +-- Lamprotula_caveata
| | +--+
| | +-- Lamprotula_cornuumlunae
| |
| | +-------- Sinosolenaia_carinata
| | +--+
| | | | +----- Sinosolenaia_recognita
| | | +--+
+--+ | | +-- Sinosolenaia_oleivora
| | +--+
| +-----------+ +-- Sinosolenaia_iridinea
| | |
| | | +----- Gonidea_angulata
| | | +--+
| | | | | +-- Leguminaia_wheatleyi
| | | | +--+
| | +--+ +-- Microcondylaea_bonellii
| | |
| | | +-- Ptychorhynchus_pfisteri
| | | |
+--+ +-----+-- Parvasolenaia_rivularis
| |
| +-- Koreosolenaia_sitgyensis
|
| +----- Indopseudodon_salwenianus
| +--------------+
| | | +-- Indopseudodon_indawgyiensis
| | +--+
| | +-- Indopseudodon_bogani
| |
| | +-------- Pilsbryoconcha_exilis__3
+--+ +--------+
| | | +----- Pilsbryoconcha_exilis__2
| | +--+
| | | +-- Pilsbryoconcha_exilis__1
| | +--+
| | +-- Pilsbryoconcha_carinifera
| |
+--+ +----------- Nyeinchanconcha_nyeinchani
| +--+
| | | +-------- Thaiconcha_callifera
| | +--+
| | | +----- Bineurus_loeiensis
| | +--+
| | | +-- Bineurus_mouhotii
+--+ +--+
| +-- Bineurus_exilis
|
| +-- Pseudodon_mekongi
| +--+
| | +-- Pseudodon_vondembuschianus
+--------+
| +-- Sundadontina_cumingii
| |
+--+-- Sundadontina_tumida__1
|
+-- Sundadontina_tumida__2
Cladogram Figure 3
+-------------------------------------------- Neotrigonia_margaritacea
|
| +-- Anodontites_trapesialis
| +--------------------------------------+
| | +-- Mutela_dubia
--+ |
| | +-------------------------------------- Echyridella_menziesii
| | |
| | | +-------------- Gibbosula_rochechouartii
| | | +--------------------+
+--+ | | | +----------- Cumberlandia_monodonta
| | | +--+
| | | | +-------- Pseudunio_marocanus
| | | +--+
| | | | +----- Margaritifera_falcata
| | | +--+
| | | | +-- Margaritifera_margaritifera
| | | +--+
+--+ | +-- Margaritifera_dahurica
| |
| | +-- Sinohyriopsis_cumingii
| | +-----------+
| | | +-- Sinohyriopsis_schlegelii
| | |
| | | +-------- Potomida_littoralis
| | +--------+ +--+
| | | | | | +----- Pronodularia_japanensis
| | | | | +--+
| | | | | | +-- Lamprotula_caveata
| | | | | +--+
+--+ | +--+ +-- Lamprotula_leaii
| | |
| | | +-- Ptychorhynchus_pfisteri
| | | +--+
| +--------+ | | +-- Parvasolenaia_rivularis
| | | +-----+
| | | | +-- Sinosolenaia_oleivora
| | | +--+
| | | +-- Sinosolenaia_carinata
| | |
| | | +-- Quadrula_quadrula
| | | +-----------------+
| | | | +-- Uniomerus_tetralasmus
| | +--+
| | | +----------------- Elliptio_complanata
| | | |
| | +--+ +-------------- Toxolasma_parvum
| | | |
| | +--+ +-- Potamilus_leptodon
| | | +--------+
| | | | +-- Potamilus_alatus
+--+ +--+
| | +-------- Leaunio_lienosus
| +--+
| | +----- Venustaconcha_ellipsiformis
| +--+
| | +-- Lampsilis_cardium
| +--+
| +-- Lampsilis_ornata
|
| +----------------------------- Lepidodesma_languilati
| |
| | +-------- Scriptolamprotula_tortuosa
| | +-----+
| | | | +----- Scriptolamprotula_scripta
| | | +--+
| | | | +-- Scriptolamprotula_tientsinensis
| | | +--+
| | +-----------+ +-- Scriptolamprotula_coreana
+--+ | |
| | | +----- Nodularia_douglasiae
| | | +-----+
| | | | | +-- Tchangsinaia_piscicula
| | | | +--+
| | +--+ +-- Schistodesmus_lampreyanus
| | |
| | | +-------- Unio_tumidus
| | +--+
+--+ | +----- Unio_nanus
| +--+
| | +-- Unio_delphinus
| +--+
| +-- Unio_pictorum
|
| +----- Lanceolaria_gladiola
| +-----------------+
| | | +-- Lanceolaria_lanceolata
| | +--+
| | +-- Lanceolaria_grayii
+--+
| +-------------------- Acuticosta_chinensis
| |
| | +-------- Cristaria_plicata
+--+ +--------+
| | | +----- Anemina_harlandi
| | +--+
| | | +-- Sinanodonta_lucida
+--+ +--+
| +-- Sinanodonta_woodiana
|
| +-------------- Anodonta_cygnea
+--+
| +----------- Anodonta_anatina
+--+
| +-------- Lasmigona_costata
+--+
| +----- Pyganodon_grandis
+--+
| +-- Utterbackia_imbecillis
+--+
+-- Utterbackia_peninsularis
Cladogram Figure 3-M
+----------------------------- Echyridella_menziesii
|
| +-- Pseudunio_marocanus
| +-----------------------+
--+ | +-- Margaritifera_margaritifera
| |
| | +----- Quadrula_quadrula
| | +-----------------+
| | | | +-- Venustaconcha_ellipsiformis
+--+ | +--+
| | +-- Potamilus_alatus
| |
| | +----------- Sinohyriopsis_cumingii
| | |
| | | +----- Potomida_littoralis
+--+ +--------+ +--+
| | | | | +-- Lamprotula_leaii
| | | | +--+
| | +--+ +-- Pronodularia_japanensis
| | |
| | | +----- Parvasolenaia_rivularis
| | +--+
| | | +-- Sinosolenaia_oleivora
+--+ +--+
| +-- Sinosolenaia_carinata
|
| +-------- Sinanodonta_woodiana
| +--------+
| | | +----- Anodonta_anatina
| | +--+
| | | +-- Pyganodon_grandis
+--+ +--+
| +-- Utterbackia_peninsularis
|
| +-------------- Lanceolaria_lanceolata
+--+
| +----------- Schistodesmus_lampreyanus
+--+
| +-------- Scriptolamprotula_tortuosa
+--+
| +----- Unio_tumidus
+--+
| +-- Unio_nanus
+--+
+-- Unio_delphinus
Cladogram Figure S1
+-- Gibbosula_rochechouartii
+--------------------------------------+
| +-- Margaritifera_dahurica
|
| +----- Sinohyriopsis_cumingii__2
| +--------------+
| | | +-- Sinohyriopsis_cumingii__1
| | +--+
| | +-- Sinohyriopsis_schlegelii
| |
| | +----- Parvasolenaia_rivularis
| +-----------------+ +-----------+
| | | | | +-- Parvasolenaia_triangularis
| | | | +--+
| | | | +-- Ptychorhynchus_pfisteri
--+ | | |
| | | | +-------- Sinosolenaia_recognita
| | +--+ |
| | | +-----+ +----- Sinosolenaia_oleivora__2
| | | | | |
| | | | +--+ +-- Sinosolenaia_oleivora__1
| | | | | |
| | | | +--+-- Sinosolenaia_iridinea
| | +--+ |
| | | +-- Sinosolenaia_oleivora__3
| | |
| | | +----------- Sinosolenaia_carinata
| | | |
| | +--+ +-------- Lamprotula_leaii
| | | |
+--+ +--+ +----- Lamprotula_cornuumlunae__1
| | |
| | +----- Lamprotula_cornuumlunae__3
| +--+
| +----- Lamprotula_cornuumlunae__4
| |
| | +-- Lamprotula_cornuumlunae__2
| +--+
| +-- Lamprotula_caveata
|
| +----------------------------------- Anemina_harlandi
| |
| | +-- Sinanodonta_angula
| | +-----------+
| | | +-- Sinanodonta_lucida
| | +-----------------+
| | | | +----------- Sinanodonta_woodiana__2
| | | +--+
| | | | +-------- Sinanodonta_jourdyi
+--+ | +--+
| | | +----- Sinanodonta_elliptica
| | +--+
| | | +-- Sinanodonta_woodiana__1
| | +--+
| | +-- Sinanodonta_lauta
| |
| | +-------------- Acuticosta_ovata__1
| | |
| | | +----------- Acuticosta_chinensis__3
+--+ +--------------+ |
| | | +----------- Acuticosta_chinensis__4
| | | |
| | +--+----------- Acuticosta_chinensis__6
| | |
| | | +-- Acuticosta_chinensis__1
| | | +-----+
| | | | +-- Acuticosta_ovata__2
| | +--+
| | | +----- Acuticosta_chinensis__5
| | | |
| | +--+----- Acuticosta_chinensis__7
| | |
| | | +-- Acuticosta_chinensis__2
+--+ +--+
| +-- Acuticosta_ovata__3
|
| +-- Lepidodesma_languilati
| +-----------------------+
| | +-- Cristaria_plicata
| |
| | +----------- Lanceolaria_gladiola
| | |
| | | +-------- Lanceolaria_grayii__5
| | +-----------+ |
| | | | | +-- Lanceolaria_lanceolata
| | | | +-----+
| | | | | +-- Lanceolaria_triformis
+--+ | +--+
| | | +----- Lanceolaria_grayii__4
| | | |
| | | | +-- Lanceolaria_eucylindrica__1
| | | | |
| | +--+ +-- Lanceolaria_eucylindrica__2
| | | |
| | | +-- Lanceolaria_eucylindrica__3
| | | |
| | +--+-- Lanceolaria_eucylindrica__4
+--+ |
| +-- Lanceolaria_grayii__1
| |
| +-- Lanceolaria_grayii__2
| |
| +-- Lanceolaria_grayii__3
|
| +----------- Scriptolamprotula_tortuosa
| +--------+
| | | +-------- Scriptolamprotula_tientsinensis
| | +--+
| | | +----- Scriptolamprotula_fibrosa
| | +--+
| | | +-- Scriptolamprotula_coreana
| | +--+
+--+ +-- Scriptolamprotula_scripta
|
| +----- Nodularia_huana
| +-----------+
| | | +-- Nodularia_nuxpersicae
| | +--+
| | +-- Nodularia_douglasiae
+--+
| +-------------- Tchangsinaia_piscicula
| |
| | +-------- Cuneopsis_rufescens
+--+ +--+
| | | +----- Cuneopsis_celtiformis
| | +--+
| | | +-- Cuneopsis_heudei
+--+ +--+
| +-- Cuneopsis_szechenyii
|
| +-------- Schistodesmus_spinosus
+--+
| +----- Schistodesmus_tongpenensis
+--+
| +-- Schistodesmus_lampreyanus
+--+
+-- Schistodesmus_xinyuensis
Cladogram Figure S3
+-- Neotrigonia_margaritacea
+-----------------------------------------------------------------+
| +-- Neotrigonia_lamarckii
|
| +----- Velesunio_ambiguus
| +--+
| | | +-- Alathyria_jacksoni
| | +--+
| +--------+ +-- Alathyria_profuga
| | |
| | | +----- Alathyria_pertexta
| | +--+
| | | +-- Lortiella_froggatti
| +-----------------------------------------------+ +--+
| | | +-- Microdontia_anodontaeformis
| | |
| | | +-------------- Westralunio_carteri
| | | |
| | +--+ +----------- Echyridella_menziesii
| | | |
| | | | +----- Diplodon_suavidicus
--+ | +--+ +--+
| | | | | +-- Castalia_ambigua
| | | | +--+
| | +--+ +-- Triplodon_corrugatus
| | |
| | | +----- Hyridella_australis
| | +--+
| | | +-- Hyridella_depressa
| | +--+
| | +-- Hyridella_drapeta
| |
| | +-------- Anodontites_trapesialis
| | +--+
| | | | +----- Mutela_rostrata
| | | +--+
| | | | +-- Mutela_dubia
| | | +--+
| | +--------------------------------------------------+ +-- Mutela_hargeri
| | | |
| | | | +----- Mycetopoda_siliquosa
+--+ | | +--+
| | | | | +-- Tamsiella_schroeteriana
| | | | +--+
| | +--+ +-- Lamproscapha_ensiformis
| | |
| | | +----- Etheria_elliptica
| | +--+
| | | +-- Aspatharia_pfeifferiana
| | +--+
| | +-- Chambardia_wahlbergi
| |
| | +-------------------- Gibbosula_rochechouartii
| | |
| | | +----------------- Gibbosula_laosensis
| | +--------------------------------------+ |
| | | | | +-------- Cumberlandia_monodonta
| | | | | +-----+
| | | +--+ | | +----- Pseudunio_marocanus
| | | | | +--+
| | | | | | +-- Pseudunio_auricularius
| | | | | +--+
| | | +--+ +-- Pseudunio_homsensis
| | | |
| | | | +-- Margaritifera_dahurica
| | | | +--------+
| | | | | +-- Margaritifera_margaritifera
| | | +--+
| | | | +-------- Margaritifera_falcata
+--+ | | |
| | +--+ +-- Margaritifera_hembeli
| | | +--+
| | | | +-- Margaritifera_marrianae
| | +--+
| | | +-- Margaritifera_laevis
| | +--+
| | +-- Margaritifera_middendorffi
| |
| | +----- Trapezidens_exolescens
| | +-----+
| | | | +-- Trapezidens_angustior
| | | +--+
| | | +-- Trapezidens_dolichorhynchus
| | |
| | +--------------+ +-- Lamellidens_corrianus
| | | | +-----+
| | | | | +-- Lamellidens_savadiensis
| | | | |
| | | | | +----- Lamellidens_lamellatus
| | | +--+ |
| | | | +----- Lamellidens_jenkinsianus
| | | | |
| | | | | +-- Lamellidens_marginalis
| | | +--+--+
| | | | +-- Lamellidens_ferrugineus
| | | |
| | +-----------------------------+ | +-- Lamellidens_mainwaringi__1
| | | | | |
| | | | +--+-- Lamellidens_mainwaringi__2
| | | | |
| | | | +-- Lamellidens_generosus
+--+ | |
| | | +----- Parreysia_keralaensis
| | | +-----------------+
| | | | | +-- Parreysia_nagpoorensis
| | | | +--+
| | | | +-- Parreysia_plagiosoma
| | | |
| | | | +----- Leoparreysia_canefrii
| | +--+ +--------------+
| | | | | +-- Leoparreysia_tavoyensis
| | | | +--+
| | | | +-- Leoparreysia_olivacea
| | | |
| | | | +----------- Coelatura_aegyptiaca__2
| | | | |
| | +--+ +-----+ +-- Coelatura_aegyptiaca__1
| | | | | +-----+
| | | | | | +-- Nitia_teretiuscula
| | | | +--+
| | | | | +----- Coelatura_kunenensis
| | | | +--+
| | | | | +-- Coelatura_gabonensis
| | | | +--+
| | +--+ +-- Prisodontopsis_mweruensis
| | |
| | | +-------- Indonaia_shurtleffiana
| | | |
| | | +-----+ +----- Indonaia_bonneaudii
| | | | | |
| | | | +--+----- Indonaia_rugosa
| | | | |
| | | | | +-- Indonaia_andersoniana
| | +--+ +--+
| | | +-- Indonaia_caerulea
| | |
| | | +----------- Indochinella_pugio
| | | |
| | | | +----- Scabies_phaselus
| | +--+ +--+
| | | | | +-- Scabies_humilis
| | | | +--+
| | +--+ +-- Scabies_mandarinus
| | |
| | | +----- Radiatula_mouhoti
| | +--+
| | | +-- Radiatula_myitkyinae
+--+ +--+
| +-- Radiatula_chaudhurii
|
| +----- Indopseudodon_indawgyiensis
| +--------------+
| | | +-- Indopseudodon_bogani
| | +--+
| | +-- Indopseudodon_salwenianus
| |
| | +-------- Pilsbryoconcha_exilis__3
| +--------------------------------+ +--------+
| | | | | +----- Pilsbryoconcha_exilis__2
| | | | +--+
| | | | | +-- Pilsbryoconcha_carinifera
| | | | +--+
| | | | +-- Pilsbryoconcha_exilis__1
| | +--+
| | | +----- Sundadontina_tumida
| | | +--------+
| | | | | +-- Pseudodon_mekongi
| | | | +--+
| | | | +-- Pseudodon_vondembuschianus
| | +--+
| | | +-- Nyeinchanconcha_nyeinchani
| | | +--------+
| | | | +-- Sundadontina_cumingii
| | +--+
| | | +-------- Thaiconcha_callifera
| | +--+
| | | +----- Bineurus_loeiensis
| | +--+
| | | +-- Bineurus_mouhotii
| | +--+
| | +-- Bineurus_exilis
| |
| | +-- Ensidens_telus
| | +-----+
| | | +-- Ensidens_ingallsianus
| | +-----+
| | | | +----- Ensidens_sagittarius
| | | +--+
| | | | +-- Ensidens_spiculus
| | +--+ +--+
| | | | +-- Ensidens_jaculus
| | | |
| | | | +----------- Hyriopsis_desowitzi
| | | | |
| | | +--+ +-------- Rectidens_sumatrensis
| | | | |
| | | +--+ +-- Hyriopsis_panhai
| | | | +--+
| | | | | +-- Hyriopsis_khoratensis
+--+ +--------------------------------+ +--+
| | | | +-- Hyriopsis_bialata
| | | +--+
| | | +-- Hyriopsis_myersiana
| | |
| | | +----- Physunio
| | | +--------+
| | | | | +-- Physunio_superbus
| | | | +--+
| | | | +-- Physunio_modelli
| | | |
| | +--+ +-------- Yaukthwa_inlenensis
| | | +--+
| | | | | +----- Yaukthwa_panhai
| | | | +--+
| | | | | +-- Yaukthwa_nesemanni
| | | | +--+
| | +--+ +-- Yaukthwa_rectangularis
| | |
| | | +-- Lens_contradens
| | | +-----+
| | | | +-- Lens_micropterus
| | +--+
| | | +----- Lens_eximius
| | +--+
| | | +-- Lens_novoselovi
| | +--+
| | +-- Lens_comptus
| |
| | +----- Chamberlainia_somsakpanhai
| | +--------------------------------------+
| | | | +-- Sinohyriopsis_cumingii
| | | +--+
| | | +-- Sinohyriopsis_schlegelii
| | |
| | | +-- Potomida_littoralis
| | | +-----+
| | | | +-- Pronodularia_japanensis
| | | +-----+
| | | | | +----- Lamprotula_leaii
| | | | +--+
| | | | | +-- Lamprotula_caveata
+--+ | | +--+
| | | +-- Lamprotula_cornuumlunae
| +--+ +--------------------------+
| | | | | +----- Sinosolenaia_recognita
| | | | | +-----+
| | | | | | | +-- Sinosolenaia_oleivora
| | | | | | +--+
| | | | | | +-- Sinosolenaia_iridinea
| | | | +--+
| | | | | +----- Gonidea_angulata
| | | | | +--+
| | | | | | | +-- Leguminaia_wheatleyi
| | | | | | +--+
| | | | +--+ +-- Microcondylaea_bonellii
| | | | |
| | | | | +----- Ptychorhynchus_pfisteri
| | +--+ +--+
| | | | +-- Parvasolenaia_rivularis
| | | +--+
| | | +-- Koreosolenaia_sitgyensis
| | |
| | | +-- Megalonaias_nervosa
| | | +--+
| | | | +-- Uniomerus_carolinianus
| | | +--------------------------------+
| | | | | +-- Quadrula_quadrula
| | | | +--+
| | | | +-- Quadrula_verrucosa
| | | |
| | | | +-------- Elliptoideus_sloatianus
| | +--+ |
| | | +--------------------------+ +-- Elliptio_complanata
| | | | | +--+
| | | | | | +-- Eurynia_dilatata
| | | | +--+
| | | | | +-- Fusconaia_flava
| | +--+ +--+
| | | +-- Pleurobema_sintoxia
| | |
| | | +-------------------------------- Amblema_plicata
| | | |
| | +--+ +----------------------------- Toxolasma_parvum
| | | |
+--+ +--+ +-- Obliquaria_reflexa
| | +-----------------------+
| | | +-- Plectomerus_dombeyanus
| +--+
| | +----------------------- Potamilus_leptodon
| | |
| +--+ +-------------------- Potamilus_alatus
| | |
| | | +-- Medionidus_conradicus
| +--+ +--------------+
| | | +-- Ptychobranchus_subtentus
| | |
| +--+ +-- Ellipsaria_lineolata
| | +-----------+
| | | +-- Truncilla_truncata
| +--+
| | +----------- Leaunio_lienosus
| | |
| +--+ +-- Venustaconcha_ellipsiformis
| | +-----+
| | | +-- Cambarunio_iris
| +--+
| | +----- Ortmanniana_ligamentina
| +--+
| | +-- Lampsilis_cardium
| +--+
| +-- Lampsilis_ornata
|
| +----------------------------------- Acuticosta_chinensis
| |
| | +-------- Scriptolamprotula_tortuosa
| | |
| | +--------------+ +-- Scriptolamprotula_coreana
| | | | +--+
| | | | | +-- Scriptolamprotula_scripta
| | | +--+
| | | | +-- Scriptolamprotula_fibrosa
| | | +--+
| | | +-- Scriptolamprotula_tientsinensis
| | +--------+
| | | | +-------- Unio_tumidus
| | | | +-----------+
+-----------+ | | | | +----- Unio_crassus
| | | | +--+
| | | | | +-- Unio_delphinus
| | | | +--+
| | | | +-- Unio_pictorum
| | +--+
| | | +----- Nodularia_douglasiae
| | | +-----------+
| | | | | +-- Nodularia_huana
| | | | +--+
| | | | +-- Nodularia_nuxpersicae
| | +--+
| | | +----- Schistodesmus_tongpenensis
| | | +--------+
| | | | | +-- Schistodesmus_xinyuensis
+--+ | | +--+
| +--+ +-- Schistodesmus_lampreyanus
| |
| | +----------- Tchangsinaia_piscicula
| +--+
| | +-------- Cuneopsis_rufescens
| +--+
| | +----- Cuneopsis_celtiformis
| +--+
| | +-- Cuneopsis_szechenyii
| +--+
| +-- Cuneopsis_heudei
|
| +----------------------------- Lepidodesma_languilati
| |
| | +-------- Lanceolaria_gladiola
| | +-----------------+
| | | | +----- Lanceolaria_grayii
| | | +--+
+--+ | | +-- Lanceolaria_lanceolata
| | +--+
| | +-- Lanceolaria_triformis
| |
| | +-------------------- Anemina_harlandi
| | |
| | +--+ +----------------- Cristaria_plicata
+--+ | | |
| | +--+ +-------------- Pletholophus_tenuis
| | | |
| | +--+ +-- Sinanodonta_angula
| | | +--------+
| | | | +-- Sinanodonta_lucida
| | +--+
| | | +-------- Sinanodonta_woodiana
| | +--+
| | | +----- Sinanodonta_jourdyi
+--+ +--+
| | +-- Sinanodonta_elliptica
| +--+
| +-- Sinanodonta_pacifica
|
| +-- Alasmidonta_marginata
| +--------+
| | +-- Lasmigona_costata
| |
| | +----- Anodonta_anatina
| +-----+
| | | +-- Anodonta_cygnea
+-----------+ +--+
| +-- Brachyanodon_nuttalliana
|
| +----- Pseudanodonta_complanata
| +--+
| | | +-- Simpsonaias_ambigua
| | +--+
+--+ +-- Strophitus_undulatus
|
| +----- Pyganodon_grandis
+--+
| +-- Utterbackia_imbecillis
+--+
+-- Utterbackia_peninsularis
Cladogram Figure S4
+-- Neotrigonia_margaritacea
+-----------------------------------+
| +-- Neotrigonia_lamarckii
|
| +----------- Etheria_elliptica
| |
| | +-- Aspatharia_pfeifferiana
| +--------+
| | +-- Chambardia_wahlbergi
| |
| +-----------------------+ +----- Mycetopoda_siliquosa
| | +-----+
| | | | +-- Tamsiella_schroeteriana
| | | +--+
| | | +-- Lamproscapha_ensiformis
| | |
| | | +-------- Anodontites_trapesialis
| | +--+
| | | +----- Mutela_hargeri
| | +--+
| | | +-- Mutela_dubia
--+ | +--+
| | +-- Mutela_rostrata
| |
| | +----------- Echyridella_menziesii
| | |
| | +----------- Westralunio_carteri
| | |
| | | +----- Diplodon_suavidicus
| | +-----+
| | | | +-- Castalia_ambigua
| | | +--+
| | | +-- Triplodon_corrugatus
| | |
| | +--------------------+ +----- Hyridella_australis
| | | +-----+
| | | | | +-- Hyridella_depressa
| | | | +--+
| | | | +-- Hyridella_drapeta
| | | |
| | | | +----- Velesunio_ambiguus
| | | | +--+
+--+ | | | | +-- Alathyria_jacksoni
| | | | +--+
| | +--+ +-- Alathyria_profuga
| | |
| | | +----- Alathyria_pertexta
| | +--+
| | | +-- Lortiella_froggatti
| | +--+
| | +-- Microdontia_anodontaeformis
| |
| | +-------------------- Gibbosula_rochechouartii
| | |
| | +--------+ +----------------- Gibbosula_laosensis
| | | | |
| | | +--+ +-------------- Cumberlandia_monodonta
| | | | |
| | | | | +----- Pseudunio_marocanus
| | | +--+ +-----+
| | | | | | +-- Pseudunio_auricularius
| | | | | +--+
| | | | | +-- Pseudunio_homsensis
| | | +--+
| | | | +-------- Margaritifera_falcata
| | | | |
| | | | | +-- Margaritifera_dahurica
| | | | +-----+
| | | +--+ +-- Margaritifera_margaritifera
| | | |
| | | | +----- Margaritifera_laevis
| | | | |
| | | +--+----- Margaritifera_middendorffi
| | | |
+--+ | | +-- Margaritifera_hembeli
| | +--+
| | +-- Margaritifera_marrianae
| |
| | +-- Parreysia_keralaensis
| | |
| | +--------------------+-- Parreysia_nagpoorensis
| | | |
| | | +-- Parreysia_plagiosoma
| | |
| | | +----- Trapezidens_exolescens
| | | +--------+
| | | | | +-- Trapezidens_angustior
| | | | +--+
| | +--------+ +-- Trapezidens_dolichorhynchus
| | | |
| | | | +----------- Lamellidens_corrianus
| | | | |
| | | +--+ +-------- Lamellidens_savadiensis
| | | | |
| | | | | +----- Lamellidens_marginalis
| | | +--+ |
| | | | +----- Lamellidens_ferrugineus
| | | | |
| | +--+ +--+ +-- Lamellidens_mainwaringi__1
| | | | | |
| | | | | +-- Lamellidens_mainwaringi__2
| | | | | |
| | | | +--+-- Lamellidens_lamellatus
| | | | |
| | | | +-- Lamellidens_generosus
| | | | |
| | | | +-- Lamellidens_jenkinsianus
| | | |
| | | | +----- Leoparreysia_tavoyensis
| | | | +--------------+
| | | | | | +-- Leoparreysia_canefrii
| | | | | +--+
| | | | | +-- Leoparreysia_olivacea
| | | | |
+--+ | | | +-------- Coelatura_aegyptiaca__2
| | | | |
| | | | +-------- Nitia_teretiuscula
| | +--+ +--------+
| | | | | +----- Coelatura_aegyptiaca__1
| | | | | |
| | | | +--+ +-- Coelatura_gabonensis
| | | | | |
| | | | +--+-- Coelatura_kunenensis
| | | | |
| | | | +-- Prisodontopsis_mweruensis
| | +--+
| | | +----- Indonaia_shurtleffiana
| | | |
| | | | +-- Indonaia_bonneaudii
| | | +--------+--+
| | | | | +-- Indonaia_rugosa
| | | | |
| | | | | +-- Indonaia_andersoniana
| | | | +--+
| | +--+ +-- Indonaia_caerulea
| | |
| | | +----------- Indochinella_pugio
| | | |
| | | | +----- Scabies_humilis
| | +--+ +--+
| | | | | +-- Scabies_phaselus
| | | | +--+
| | +--+ +-- Scabies_mandarinus
| | |
| | | +----- Radiatula_mouhoti
| | +--+
| | | +-- Radiatula_myitkyinae
| | +--+
| | +-- Radiatula_chaudhurii
| |
| | +-------------------- Chamberlainia_somsakpanhai
| | |
| | +-------------------- Megalonaias_nervosa
| | |
| | +-------------------- Uniomerus_carolinianus
| | |
| | | +-- Quadrula_quadrula
| | +-----------------+
| | | +-- Quadrula_verrucosa
| | |
| | | +-- Sinohyriopsis_cumingii
| | +-----------------+
| | | +-- Sinohyriopsis_schlegelii
| | |
| | | +-- Potomida_littoralis
| | | +-----+
| | | | +-- Pronodularia_japanensis
| | +-----------+
| | | | +----- Lamprotula_leaii
| | | +--+
| | | | +-- Lamprotula_caveata
| | | +--+
| | | +-- Lamprotula_cornuumlunae
| | |
| | | +-------- Sinosolenaia_carinata
+--+ | |
| | | +----- Gonidea_angulata
| | +--+
| | | | +-- Leguminaia_wheatleyi
| | | +--+
| | | +-- Microcondylaea_bonellii
| +-----------+
| | | +----- Ptychorhynchus_pfisteri
| | +--+
| | | | +-- Parvasolenaia_rivularis
| | | +--+
| | | +-- Koreosolenaia_sitgyensis
| | |
| | | +----- Sinosolenaia_recognita
| | +--+
| | | +-- Sinosolenaia_iridinea
| | +--+
| | +-- Sinosolenaia_oleivora
| |
| | +----- Indopseudodon_salwenianus
| | +-----+
| | | | +-- Indopseudodon_indawgyiensis
| | | +--+
| | | +-- Indopseudodon_bogani
| | |
| | | +-------- Nyeinchanconcha_nyeinchani
| | | |
| +--------+ +-------- Thaiconcha_callifera
| | | |
| | | +-------- Sundadontina_cumingii
| | | |
| | | +-------- Sundadontina_tumida__1
| | | |
| | | +-------- Sundadontina_tumida__2
| | | |
| | +--+ +-- Pseudodon_mekongi
| | +-----+
| | | +-- Pseudodon_vondembuschianus
| | |
| | | +----- Pilsbryoconcha_exilis__2
| | +--+
| | | | +-- Pilsbryoconcha_carinifera
| | | +--+
| | | +-- Pilsbryoconcha_exilis__1
| | |
| | | +----- Bineurus_loeiensis
| | +--+
| | | +-- Bineurus_mouhotii
| | +--+
| | +-- Bineurus_exilis
| |
| | +-------------- Amblema_plicata
| | |
| | +-------------- Toxolasma_parvum
| | |
| | | +-- Obliquaria_reflexa
| | +-----------+
| | | +-- Plectomerus_dombeyanus
| | |
| | | +----------- Elliptoideus_sloatianus
| | +--+
| | | | +-------- Elliptio_complanata
| | | +--+
| | | | +----- Eurynia_dilatata
| +-----+ +--+
| | | | +-- Fusconaia_flava
| | | +--+
| | | +-- Pleurobema_sintoxia
| | |
| | | +-------- Ellipsaria_lineolata
| | | |
| | | +-------- Potamilus_leptodon
| | | |
| | | +-------- Medionidus_conradicus
| | | |
| | | +-------- Potamilus_alatus
| | | |
+-----+ | +-------- Ptychobranchus_subtentus
| +-----+
| +-------- Truncilla_truncata
| |
| +-------- Venustaconcha_ellipsiformis
| |
| +-------- Cambarunio_iris
| |
| +-------- Leaunio_lienosus
| |
| | +----- Ortmanniana_ligamentina
| +--+
| | +-- Lampsilis_cardium
| +--+
| +-- Lampsilis_ornata
|
| +-------------- Hyriopsis_desowitzi
| |
| +-------------- Hyriopsis_myersiana
| |
| +-------------- Hyriopsis_panhai
| |
| +--+ +-- Hyriopsis_bialata
| | +-----------+
| | | +-- Hyriopsis_khoratensis
| | |
| | | +----------- Rectidens_sumatrensis
| | | |
| | +--+ +-- Ensidens_telus
| | | +-----+
| | | | +-- Ensidens_ingallsianus
| | +--+
| | | +----- Ensidens_sagittarius
| | +--+
+--+ | +-- Ensidens_spiculus
| | +--+
| | +-- Ensidens_jaculus
| |
| | +----------- Physunio_modelli
| | |
| | +----------- Physunio
| | |
| | +----------- Physunio_superbus
| | |
| | | +-------- Yaukthwa_inlenensis
| | +--+
| +-----+ | +----- Yaukthwa_panhai
| | +--+
| | | +-- Yaukthwa_nesemanni
| | +--+
| | +-- Yaukthwa_rectangularis
| |
| | +-- Lens_comptus
| | +-----+
| | | +-- Lens_novoselovi
| +--+
| | +----- Lens_eximius
| +--+
| | +-- Lens_contradens
| +--+
| +-- Lens_micropterus
|
| +----------------- Acuticosta_chinensis
| |
| +----------------- Alasmidonta_marginata
| |
| +----------------- Anodonta_anatina
| |
| +----------------- Anemina_harlandi
| |
| +----------------- Anodonta_cygnea
| |
| +----------------- Cristaria_plicata
| |
| +----------------- Brachyanodon_nuttalliana
| |
| +----------------- Lasmigona_costata
| |
| +----------------- Lepidodesma_languilati
| |
| +----------------- Pseudanodonta_complanata
| |
| +----------------- Pyganodon_grandis
| |
| +----------------- Simpsonaias_ambigua
| |
| +----------------- Strophitus_undulatus
| |
| | +-- Utterbackia_imbecillis
| +--------------+
| | +-- Utterbackia_peninsularis
| |
| | +-------- Lanceolaria_gladiola
| +--------+
| | | +----- Lanceolaria_grayii
| | +--+
| | | +-- Lanceolaria_lanceolata
| | +--+
| | +-- Lanceolaria_triformis
| |
| | +-------- Schistodesmus_spinosus
| +--------+
| | | +----- Schistodesmus_tongpenensis
+--+ +--+
| | +-- Schistodesmus_xinyuensis
| +--+
| +-- Schistodesmus_lampreyanus
|
| +-------- Unio_tumidus
+--------+
| | +----- Unio_crassus
| +--+
| | +-- Unio_delphinus
| +--+
| +-- Unio_pictorum
|
| +-------- Scriptolamprotula_tortuosa
| |
+--------+ +-- Scriptolamprotula_coreana
| | +--+
| | | +-- Scriptolamprotula_scripta
| +--+
| | +-- Scriptolamprotula_fibrosa
| +--+
| +-- Scriptolamprotula_tientsinensis
|
| +-------------- Pletholophus_tenuis
| |
+--+ +-- Sinanodonta_angula
| | +--------+
| | | +-- Sinanodonta_lucida
| +--+
| | +-------- Sinanodonta_woodiana
| +--+
| | +----- Sinanodonta_jourdyi
| +--+
| | +-- Sinanodonta_elliptica
| +--+
| +-- Sinanodonta_pacifica
|
| +-------- Cuneopsis_celtiformis
| |
| +-------- Tchangsinaia_piscicula
| |
| +-------- Cuneopsis_rufescens
+--------+
| +-- Cuneopsis_szechenyii
+-----+
| +-- Cuneopsis_heudei
|
| +----- Nodularia_huana
+--+
| +-- Nodularia_douglasiae
+--+
+-- Nodularia_nuxpersicae
|
