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Thu 17 July 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!


Publication Bolotov et al. (2020): cladograms
     publication summary | cladograms

Complete Citation

Bolotov, I.N., E.S. Konopleva, I.V. Vikhrev, M.Yu. Gofarov, M. Lopes-Lima et al. 2020. New freshwater mussel taxa discoveries clarify biogeographic division of Southeast Asia. Scientific Reports 10. 6616 (22 pp.). Available electronically.

Cladogram Figures

(click the figure number to see the cladogram)

Cladogram Figure 1
                                            +-- Gibbosula_laosensis          
                                            |                                
                                            +-- Margaritifera_dahurica       
                                            |                                
  +-----------------------------------------+-- Margaritifera_margaritifera  
  |                                         |                                
  |                                         +-- Margaritifera_laevis         
  |                                         |                                
  |                                         +-- Margaritifera_middendorffi   
  |                                                                          
  |                                   +-------- Trapezidens_exolescens       
  |                                +--+                                      
  |                                |  |  +----- Trapezidens_scutum           
  |                                |  +--+                                   
  |                                |     |  +-- Trapezidens_dolichorhynchus  
  |     +--------------------------+     +--+                                
  |     |                          |        +-- Trapezidens_angustior        
  |     |                          |                                         
  |     |                          |  +-------- Lamellidens_savadiensis      
  |     |                          +--+                                      
  |     |                             |  +----- Lamellidens_generosus        
  |     |                             +--+                                   
  |     |                                |  +-- Lamellidens_ferrugineus      
  |     |                                +--+                                
  |     |                                   +-- Lamellidens_mainwaringi      
  |     |                                                                    
  |     |                          +----------- Parreysia_keralaensis        
  |     |                       +--+                                         
  |  +--+                       |  |  +-------- Parreysia_plagiosoma         
  |  |  |                       |  +--+                                      
  |  |  |                       |     |  +----- Parreysia_corrugata          
  |  |  |                       |     +--+                                   
  |  |  |                       |        |  +-- Parreysia_nagpoorensis       
  |  |  |  +--------------------+        +--+                                
  |  |  |  |                    |           +-- Parreysia_rakhinensis        
  |  |  |  |                    |                                            
  |  |  |  |                    |           +-- Leoparreysia_burmana         
  |  |  |  |                    |     +-----+                                
  |  |  |  |                    |     |     +-- Leoparreysia_whitteni        
  |  |  |  |                    +-----+                                      
  |  |  |  |                          |  +----- Leoparreysia_tavoyensis      
  |  |  |  |                          +--+                                   
  |  |  +--+                             |  +-- Leoparreysia_olivacea        
  |  |     |                             +--+                                
  |  |     |                                +-- Leoparreysia_canefrii        
  |  |     |                                                                 
  |  |     |                                +-- Coelatura_aegyptiaca         
--+  |     |  +-----------------------------+                                
  |  |     |  |                             +-- Nitia_teretiuscula           
  |  |     |  |                                                              
  |  |     |  |                    +----------- Indonaia_shurtleffiana       
  |  |     |  |  +-----------------+                                         
  |  |     +--+  |                 |  +-------- Indonaia_rugosa              
  |  |        |  |                 +--+                                      
  |  |        |  |                    |  +----- Indonaia_andersoniana        
  |  |        |  |                    +--+                                   
  |  |        |  |                       |  +-- Indonaia_subclathrata        
  |  |        +--+                       +--+                                
  |  |           |                          +-- Indonaia_caerulea            
  |  |           |                                                           
  |  |           |  +-------------------------- Indochinella_pugio           
  |  |           |  |                                                        
  |  |           |  |                 +-------- Radiatula_crispisulcata      
  |  |           |  |  +--------------+                                      
  |  |           +--+  |              |  +----- Radiatula_mouhoti            
  |  |              |  |              +--+                                   
  |  |              |  |                 |  +-- Radiatula_chaudhurii         
  |  |              |  |                 +--+                                
  |  |              +--+                    +-- Radiatula_myitkyinae         
  |  |                 |                                                     
  |  |                 |  +-------------------- Scabiellus_songkramensis     
  |  |                 |  |                                                  
  |  |                 |  |                 +-- Unionetta_fabagina           
  |  |                 +--+  +--------------+                                
  |  |                    |  |              +-- Harmandia_somboriensis       
  |  |                    |  |                                               
  |  |                    +--+  +-------------- Scabies_nucleus              
  |  |                       |  |                                            
  |  |                       |  |           +-- Scabies_crispata             
  |  |                       +--+  +--------+                                
  |  |                          |  |        +-- Scabies_mandarinus           
  |  |                          |  |                                         
  |  |                          +--+        +-- Scabies_humilis              
  |  |                             |  +-----+                                
  |  |                             |  |     +-- Scabies_pilata               
  |  |                             +--+                                      
  |  |                                |  +----- Scabies_anceps               
  |  |                                +--+                                   
  |  |                                   |  +-- Scabies_phaselus             
  |  |                                   +--+                                
  +--+                                      +-- Scabies_scobinatus           
     |                                                                       
     |                             +----------- Hyriopsis_desowitzi          
     |                             |                                         
     |                          +--+  +-------- Rectidens_sumatrensis        
     |                          |  |  |                                      
     |                          |  +--+     +-- Hyriopsis_khoratensis        
     |                          |     |  +--+                                
     |                          |     |  |  +-- Hyriopsis_bialata            
     |                          |     +--+                                   
     |                          |        |  +-- Hyriopsis_myersiana          
     |                 +--------+        +--+                                
     |                 |        |           +-- Hyriopsis_panhai             
     |                 |        |                                            
     |                 |        |           +-- Ensidens_telus               
     |                 |        |     +-----+                                
     |                 |        |     |     +-- Ensidens_ingallsianus        
     |                 |        |     |                                      
     |                 |        +-----+     +-- Ensidens_jaculus             
     |                 |              |  +--+                                
     |                 |              |  |  +-- Ensidens_spiculus            
     |                 |              +--+                                   
     |                 |                 |  +-- Ensidens_dugasti             
     |                 |                 +--+                                
     |                 |                    +-- Ensidens_sagittarius         
     |           +-----+                                                     
     |           |     |                    +-- Trapezoideus_lenya           
     |           |     |              +-----+                                
     |           |     |              |     +-- Trapezoideus_foliaceus       
     |           |     |  +-----------+                                      
     |           |     |  |           |  +----- Physunio_superbus            
     |           |     |  |           +--+                                   
     |           |     |  |              |  +-- Physunio_modelli             
     |           |     |  |              +--+                                
     |           |     |  |                 +-- Physunio                     
     |           |     |  |                                                  
     |           |     |  |                 +-- Yaukthwa_elongatula          
     |           |     |  |     +-----------+                                
     |           |     |  |     |           +-- Yaukthwa_baniensis           
     |           |     +--+     |                                            
     |           |        |  +--+        +----- Yaukthwa_inlenensis          
     |           |        |  |  |  +-----+                                   
     |           |        |  |  |  |     |  +-- Yaukthwa_avaensis            
     |           |        |  |  |  |     +--+                                
     |           |        |  |  +--+        +-- Yaukthwa_paiensis            
     |           |        |  |     |                                         
     |           |        |  |     |  +-------- Yaukthwa_panhai              
     |           |        |  |     +--+                                      
     |           |        |  |        |  +----- Yaukthwa_rectangularis       
     |           |        +--+        +--+                                   
     |           |           |           |  +-- Yaukthwa_zayleymanensis      
     |           |           |           +--+                                
     |           |           |              +-- Yaukthwa_nesemanni           
     |           |           |                                               
     |           |           |           +----- Lens_misellus                
     |           |           |     +-----+                                   
     |           |           |     |     |  +-- Lens_contradens              
     |           |           |     |     +--+                                
     |           |           |     |        +-- Lens_micropterus             
     |           |           +-----+                                         
     |           |                 |     +----- Lens_pallegoixi              
     |           |                 |  +--+                                   
     +-----------+                 |  |  |  +-- Lens_comptus                 
                 |                 |  |  +--+                                
                 |                 +--+     +-- Lens_novoselovi              
                 |                    |                                      
                 |                    |  +----- Lens_maenamensis             
                 |                    +--+                                   
                 |                       |  +-- Lens_eximius                 
                 |                       +--+                                
                 |                          +-- Lens_rolfbrandti             
                 |                                                           
                 |                          +-- Sinosolenaia_iridinea        
                 |                          |                                
                 |                          +-- Sinosolenaia_recognita__1    
                 |                          |                                
                 |                          +-- Sinosolenaia_carinata        
                 |                          |                                
                 |                          +-- Leguminaia_wheatleyi         
                 |                          |                                
                 |                          +-- Microcondylaea_bonellii      
                 |                          |                                
                 |                          +-- Gonidea_angulata             
                 |                          |                                
                 |                          +-- Ptychorhynchus_pfisteri      
                 |                          |                                
                 |                          +-- Parvasolenaia_rivularis      
                 |                          |                                
                 |  +-----------------------+-- Lamprotula_leaii             
                 |  |                       |                                
                 |  |                       +-- Lamprotula                   
                 |  |                       |                                
                 |  |                       +-- Lamprotula_caveata           
                 |  |                       |                                
                 |  |                       +-- Potomida_semirugata          
                 |  |                       |                                
                 |  |                       +-- Pronodularia_japanensis      
                 |  |                       |                                
                 |  |                       +-- Sinohyriopsis_cumingii       
                 |  |                       |                                
                 +--+                       +-- Chamberlainia_somsakpanhai   
                    |                       |                                
                    |                       +-- Sinosolenaia_recognita__2    
                    |                       |                                
                    |                       +-- Sinosolenaia_oleivora        
                    |                                                        
                    |                 +-------- Indopseudodon_kayinensis     
                    |  +--------------+                                      
                    |  |              |  +----- Indopseudodon_indawgyiensis  
                    |  |              +--+                                   
                    |  |                 |  +-- Indopseudodon_salwenianus    
                    |  |                 +--+                                
                    |  |                    +-- Indopseudodon_bogani         
                    +--+                                                     
                       |                 +----- Bineurus_loeiensis           
                       |  +--------------+                                   
                       |  |              |  +-- Bineurus_exilis              
                       |  |              +--+                                
                       |  |                 +-- Bineurus_mouhotii            
                       |  |                                                  
                       +--+  +----------------- Nyeinchanconcha_nyeinchani   
                          |  |                                               
                          |  |              +-- Thaiconcha_callifera         
                          |  |        +-----+                                
                          |  |        |     +-- Thaiconcha_munelliptica      
                          +--+  +-----+                                      
                             |  |     |  +----- Pilsbryoconcha_exilis__2     
                             |  |     +--+                                   
                             |  |        |  +-- Pilsbryoconcha_exilis__1     
                             |  |        +--+                                
                             |  |           +-- Pilsbryoconcha               
                             +--+                                            
                                |     +-------- Pseudodon_lenyanensis        
                                |  +--+                                      
                                |  |  |  +----- Pseudodon_vagulus            
                                |  |  +--+                                   
                                |  |     |  +-- Pseudodon_vondembuschianus   
                                |  |     +--+                                
                                +--+        +-- Pseudodon_mekongi            
                                   |                                         
                                   |        +-- Sundadontina_taskaevi        
                                   |  +-----+                                
                                   |  |     +-- Sundadontina_brandti         
                                   +--+                                      
                                      |  +----- Sundadontina_tumida          
                                      +--+                                   
                                         |  +-- Sundadontina_cumingii        
                                         +--+                                
                                            +-- Sundadontina_tanintharyiensis
Cladogram Figure S1
                                         +-------- Trapezidens_exolescens       
                                      +--+                                      
                                      |  |  +----- Trapezidens_scutum           
                                      |  +--+                                   
                                      |     |  +-- Trapezidens_dolichorhynchus  
     +--------------------------------+     +--+                                
     |                                |        +-- Trapezidens_angustior        
     |                                |                                         
     |                                |  +-------- Lamellidens_savadiensis      
     |                                +--+                                      
     |                                   |  +----- Lamellidens_generosus        
     |                                   +--+                                   
     |                                      |  +-- Lamellidens_ferrugineus      
     |                                      +--+                                
     |                                         +-- Lamellidens_mainwaringi      
     |                                                                          
     |                                +----------- Parreysia_keralaensis        
     |                             +--+                                         
  +--+                             |  |  +-------- Parreysia_corrugata          
  |  |                             |  +--+                                      
  |  |                             |     |  +----- Parreysia_nagpoorensis       
  |  |                             |     +--+                                   
  |  |                             |        |  +-- Parreysia_rakhinensis        
  |  |  +--------------------------+        +--+                                
  |  |  |                          |           +-- Parreysia_plagiosoma         
  |  |  |                          |                                            
  |  |  |                          |           +-- Leoparreysia_burmana         
  |  |  |                          |     +-----+                                
  |  |  |                          |     |     +-- Leoparreysia_whitteni        
  |  |  |                          +-----+                                      
  |  |  |                                |  +----- Leoparreysia_olivacea        
  |  |  |                                +--+                                   
  |  +--+                                   |  +-- Leoparreysia_tavoyensis      
  |     |                                   +--+                                
  |     |                                      +-- Leoparreysia_canefrii        
  |     |                                                                       
  |     |                                      +-- Coelatura_aegyptiaca         
  |     |  +-----------------------------------+                                
  |     |  |                                   +-- Nitia_teretiuscula           
  |     |  |                                                                    
  |     |  |                             +-------- Indonaia_shurtleffiana       
  |     +--+  +--------------------------+                                      
  |        |  |                          |  +----- Indonaia_andersoniana        
  |        |  |                          +--+                                   
  |        |  |                             |  +-- Indonaia_caerulea            
  |        |  |                             +--+                                
  |        +--+                                +-- Indonaia_subclathrata        
  |           |                                                                 
  |           |  +-------------------------------- Indochinella_pugio           
  |           |  |                                                              
  |           |  |                          +----- Radiatula_mouhoti            
  |           +--+  +-----------------------+                                   
  |              |  |                       |  +-- Radiatula_myitkyinae         
  |              |  |                       +--+                                
  |              +--+                          +-- Radiatula_chaudhurii         
  |                 |                                                           
  |                 |  +-------------------------- Radiatula_crispisulcata      
  |                 |  |                                                        
  |                 +--+  +----------------------- Scabiellus_songkramensis     
  |                    |  |                                                     
  |                    +--+  +-------------------- Harmandia_somboriensis       
  |                       |  |                                                  
  |                       +--+  +----------------- Unionetta_fabagina           
  |                          |  |                                               
  |                          +--+  +-------------- Scabies_mandarinus           
  |                             |  |                                            
  |                             +--+  +----------- Scabies_crispata             
  |                                |  |                                         
  |                                +--+  +-------- Scabies_nucleus              
  |                                   |  |                                      
  |                                   +--+     +-- Scabies_humilis              
  |                                      |  +--+                                
  |                                      |  |  +-- Scabies_pilata               
  |                                      +--+                                   
  |                                         |  +-- Scabies_phaselus             
  |                                         |  |                                
  |                                         +--+-- Scabies_scobinatus           
  |                                            |                                
--+                                            +-- Scabies_anceps               
  |                                                                             
  |                                +-------------- Hyriopsis_desowitzi          
  |                             +--+                                            
  |                             |  |  +----------- Rectidens_sumatrensis        
  |                             |  +--+                                         
  |                             |     |  +-------- Hyriopsis_panhai             
  |                             |     +--+                                      
  |                             |        |  +----- Hyriopsis_myersiana          
  |                             |        +--+                                   
  |                    +--------+           |  +-- Hyriopsis_khoratensis        
  |                    |        |           +--+                                
  |                    |        |              +-- Hyriopsis_bialata            
  |                    |        |                                               
  |                    |        |              +-- Ensidens_telus               
  |                    |        |        +-----+                                
  |                    |        |        |     +-- Ensidens_ingallsianus        
  |                    |        |        |                                      
  |                    |        +--------+     +-- Ensidens_dugasti             
  |                    |                 |  +--+                                
  |                    |                 |  |  +-- Ensidens_sagittarius         
  |                    |                 +--+                                   
  |                 +--+                    |  +-- Ensidens_spiculus            
  |                 |  |                    +--+                                
  |                 |  |                       +-- Ensidens_jaculus             
  |                 |  |                                                        
  |                 |  |                    +----- Physunio                     
  |                 |  |  +-----------------+                                   
  |                 |  |  |                 |  +-- Physunio_modelli             
  |                 |  |  |                 +--+                                
  |                 |  |  |                    +-- Physunio_superbus            
  |                 |  |  |                                                     
  |                 |  |  |                    +-- Trapezoideus_lenya           
  |                 |  |  |  +-----------------+                                
  |                 |  +--+  |                 +-- Trapezoideus_foliaceus       
  |                 |     |  |                                                  
  |                 |     |  |              +----- Lens_misellus                
  |                 |     |  |        +-----+                                   
  |                 |     |  |        |     |  +-- Lens_contradens              
  |                 |     |  |        |     +--+                                
  |                 |     |  |        |        +-- Lens_micropterus             
  |                 |     +--+  +-----+                                         
  |                 |        |  |     |     +----- Lens_maenamensis             
  |                 |        |  |     |  +--+                                   
  |                 |        |  |     |  |  |  +-- Lens_eximius                 
  |                 |        |  |     |  |  +--+                                
  |                 |        |  |     +--+     +-- Lens_rolfbrandti             
  |                 |        |  |        |                                      
  |                 |        |  |        |  +----- Lens_pallegoixi              
  |                 |        +--+        +--+                                   
  |                 |           |           |  +-- Lens_novoselovi              
  |                 |           |           +--+                                
  |                 |           |              +-- Lens_comptus                 
  |                 |           |                                               
  |                 |           |              +-- Yaukthwa_elongatula          
  |                 |           |  +-----------+                                
  |                 |           |  |           +-- Yaukthwa_baniensis           
  |                 |           |  |                                            
  |                 |           +--+        +----- Yaukthwa_inlenensis          
  |                 |              |  +-----+                                   
  +-----------------+              |  |     |  +-- Yaukthwa_avaensis            
                    |              |  |     +--+                                
                    |              +--+        +-- Yaukthwa_paiensis            
                    |                 |                                         
                    |                 |  +-------- Yaukthwa_panhai              
                    |                 +--+                                      
                    |                    |  +----- Yaukthwa_rectangularis       
                    |                    +--+                                   
                    |                       |  +-- Yaukthwa_zayleymanensis      
                    |                       +--+                                
                    |                          +-- Yaukthwa_nesemanni           
                    |                                                           
                    |                          +-- Sinohyriopsis_cumingii       
                    |                       +--+                                
                    |                       |  +-- Chamberlainia_somsakpanhai   
                    |                       |                                   
                    |                    +--+  +-- Lamprotula_leaii             
                    |                    |  |  |                                
                    |                    |  |  +-- Lamprotula                   
                    |                    |  |  |                                
                    |                    |  +--+-- Lamprotula_caveata           
                    |                    |     |                                
                    |                    |     +-- Potomida_semirugata          
                    |                    |     |                                
                    |                    |     +-- Pronodularia_japanensis      
                    |  +-----------------+                                      
                    |  |                 |     +-- Sinosolenaia_iridinea        
                    |  |                 |     |                                
                    |  |                 |     +-- Sinosolenaia_recognita__1    
                    |  |                 |     |                                
                    |  |                 |     +-- Sinosolenaia_carinata        
                    |  |                 |     |                                
                    |  |                 |     +-- Leguminaia_wheatleyi         
                    |  |                 |     |                                
                    |  |                 |     +-- Microcondylaea_bonellii      
                    |  |                 +-----+                                
                    |  |                       +-- Gonidea_angulata             
                    |  |                       |                                
                    |  |                       +-- Ptychorhynchus_pfisteri      
                    +--+                       |                                
                       |                       +-- Parvasolenaia_rivularis      
                       |                       |                                
                       |                       +-- Sinosolenaia_recognita__2    
                       |                       |                                
                       |                       +-- Sinosolenaia_oleivora        
                       |                                                        
                       |                 +-------- Indopseudodon_kayinensis     
                       |  +--------------+                                      
                       |  |              |  +----- Indopseudodon_indawgyiensis  
                       |  |              +--+                                   
                       |  |                 |  +-- Indopseudodon_salwenianus    
                       |  |                 +--+                                
                       |  |                    +-- Indopseudodon_bogani         
                       +--+                                                     
                          |                 +----- Bineurus_loeiensis           
                          |  +--------------+                                   
                          |  |              |  +-- Bineurus_exilis              
                          |  |              +--+                                
                          |  |                 +-- Bineurus_mouhotii            
                          |  |                                                  
                          +--+  +----------------- Nyeinchanconcha_nyeinchani   
                             |  |                                               
                             |  |              +-- Thaiconcha_callifera         
                             |  |        +-----+                                
                             |  |        |     +-- Thaiconcha_munelliptica      
                             +--+  +-----+                                      
                                |  |     |  +----- Pilsbryoconcha_exilis__2     
                                |  |     +--+                                   
                                |  |        |  +-- Pilsbryoconcha_exilis__1     
                                |  |        +--+                                
                                |  |           +-- Pilsbryoconcha               
                                +--+                                            
                                   |     +-------- Pseudodon_lenyanensis        
                                   |  +--+                                      
                                   |  |  |  +----- Pseudodon_vagulus            
                                   |  |  +--+                                   
                                   |  |     |  +-- Pseudodon_vondembuschianus   
                                   |  |     +--+                                
                                   +--+        +-- Pseudodon_mekongi            
                                      |                                         
                                      |        +-- Sundadontina_tanintharyiensis
                                      |  +-----+                                
                                      |  |     +-- Sundadontina_cumingii        
                                      +--+                                      
                                         |  +----- Sundadontina_tumida          
                                         +--+                                   
                                            |  +-- Sundadontina_taskaevi        
                                            +--+                                
                                               +-- Sundadontina_brandti         
 
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