The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)
FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!
Publication Wu et al. (2024): cladograms
publication summary | cladograms Complete CitationWu, R., L. Zhang, L. Liu, J. Jia & X. Liu. 2024. Unraveling the phylogenetic relationships and taxonomic status of a puzzling freshwater mussel genus Inversidens (Bivalvia, Unionidae) through multilocus phylogeny and mitochondrial phylogenomics. Journal of Zoological Systematics and Evolutionary Research 2024, 1499508: 1-18. Available electronically.
Cladogram Figures(click the figure number to see the cladogram) Cladogram Figure 5a
+-- Margaritifera_dahurica
+--------------------------+
| +-- Margaritifera_margaritifera
|
| +-------------------------- Chamberlainia_somsakpanhai
| |
| | +-- Khairuloconcha_lunbawangorum
| | +--------------+
--+ | | +-- Khairuloconcha_sahanae
| | |
| | | +-------- Ensidens_sagittarius
| | +-----+ +-----+
| | | | | | +----- Rectidens_sumatrensis
| | | | | +--+
| | | | | | +-- Hyriopsis_bialata
| | | | | +--+
+--+ | +--+ +-- Hyriopsis_desowitzi
| | |
| | | +-- Physunio_superbus
| | | +--------+
| | | | +-- Trapezoideus_foliaceus
| | | |
| | +--+ +-- Lens_contradens
| | | +-----+
| | | | +-- Lens_eximius
| | +--+
| | | +----- Yaukthwa_elongatula
| | +--+
+--+ | +-- Yaukthwa_inlenensis
| +--+
| +-- Yaukthwa_paiensis
|
| +----------------- Schepmania_nieuwenhuisi
| |
| +--+ +-- Indopseudodon_bogani
| | | +-----------+
| | | | +-- Indopseudodon_kayinensis
| | +--+
| | | +----------- Pilsbryoconcha_exilis
| | | |
| | +--+ +-- Pseudodon_mekongi
| | | +-----+
| | | | +-- Pseudodon_vondembuschianus
| | +--+
| | | +----- Thaiconcha_callifera
+--+ +--+
| | +-- Bineurus_anodontinum
| +--+
| +-- Bineurus_loeiensis
|
| +-- Inversidens_brandtii
| +-----+
| | +-- Inversidens_rentianensis
| +--------+
| | | +----- Potomida_littoralis
| | +--+
| | | +-- Lamprotula_caveata
+--+ +--+
| +-- Lamprotula_leaii
|
| +-------------- Sinosolenaia_carinata
| |
+--+ +-- Gonidea_angulata
| +--------+
| | +-- Microcondylaea_bonellii
+--+
| +-------- Parvasolenaia_rivularis
+--+
| +----- Koreosolenaia_sitgyensis
+--+
| +-- Postolata_guangxiensis
+--+
+-- Ptychorhynchus_pfisteri
Cladogram Figure 5b
+-- Margaritifera_dahurica
+-----------------------------------+
| +-- Margaritifera_margaritifera
|
| +----------------------------------- Schepmania_nieuwenhuisi
| |
| | +-- Indopseudodon_bogani
--+ | +-----------+
| | | +-- Indopseudodon_kayinensis
| | +-----------------+
| | | | +----------- Pilsbryoconcha_exilis
| | | | |
| | | +--+ +-- Bineurus_anodontinum
| | | | +-----+
+--+ | | | +-- Bineurus_loeiensis
| | +--+
| | | +----- Thaiconcha_callifera
| | +--+
| | | +-- Pseudodon_mekongi
| | +--+
| | +-- Pseudodon_vondembuschianus
| |
| | +-------------------------- Hyriopsis_bialata
| | |
+--+ | +----- Rectidens_sumatrensis
| +--+ +-----------------+
| | | | | +-- Khairuloconcha_lunbawangorum
| | | | +--+
| | +--+ +-- Khairuloconcha_sahanae
| | |
| | | +-------------------- Ensidens_sagittarius
| | | |
| | +--+ +----------------- Hyriopsis_desowitzi
| | | |
| | +--+ +-------------- Yaukthwa_elongatula
| | | |
| | +--+ +----------- Trapezoideus_foliaceus
| | | |
| | +--+ +-- Physunio_superbus
+--+ | +-----+
| | | +-- Yaukthwa_inlenensis
| +--+
| | +----- Yaukthwa_paiensis
| +--+
| | +-- Lens_contradens
| +--+
| +-- Lens_eximius
|
| +-- Inversidens_brandtii
| +-----------------+
| | +-- Inversidens_rentianensis
| |
| | +-- Chamberlainia_somsakpanhai
+--------+ +--+
| | +-- Potomida_littoralis
| +-----------+
| | | +-- Lamprotula_caveata
| | +--+
+--+ +-- Lamprotula_leaii
|
| +-------------- Sinosolenaia_carinata
| |
+--+ +-- Gonidea_angulata
| +--------+
| | +-- Microcondylaea_bonellii
+--+
| +-------- Koreosolenaia_sitgyensis
+--+
| +----- Parvasolenaia_rivularis
+--+
| +-- Postolata_guangxiensis
+--+
+-- Ptychorhynchus_pfisteri
Cladogram Figure 5c
+-- Margaritifera_dahurica
+-----------------------------+
| +-- Margaritifera_margaritifera
|
| +----------------------------- Schepmania_nieuwenhuisi__1
| |
| +----------------------------- Schepmania_nieuwenhuisi__2
| |
--+ | +-- Indopseudodon_bogani
| | +-----------+
| | | +-- Indopseudodon_kayinensis
| | +-----------+
| | | | +----------- Pilsbryoconcha_exilis
| | | | |
| | | +--+ +-------- Bineurus_anodontinum
+--+ | | |
| | +--+-------- Bineurus_loeiensis
| | |
| | | +----- Thaiconcha_callifera
| | +--+
| | | +-- Pseudodon_mekongi
| | +--+
| | +-- Pseudodon_vondembuschianus
| |
| | +-------- Ensidens_sagittarius
| | |
| | +-------- Hyriopsis_desowitzi
+--+ |
| +-------- Rectidens_sumatrensis
| |
| +-------- Hyriopsis_bialata
| |
| +--------------+ +-- Khairuloconcha_lunbawangorum
| | +-----+
| | | +-- Khairuloconcha_sahanae
| | |
| | | +----- Physunio_superbus
| | | |
| | | +----- Yaukthwa_paiensis
| | | |
| | | +----- Yaukthwa_inlenensis
| | +--+
| | +----- Trapezoideus_foliaceus
+--+ |
| +----- Yaukthwa_elongatula
| |
| | +-- Lens_contradens
| +--+
| +-- Lens_eximius
|
| +-- Inversidens_rentianensis
| +-----------------+
| | +-- Inversidens_brandtii
| |
| | +-- Chamberlainia_somsakpanhai
+--+ +--+
| | +-- Potomida_littoralis
| +-----------+
| | | +-- Lamprotula_caveata
| | +--+
+--+ +-- Lamprotula_leaii
|
| +-------------- Sinosolenaia_carinata
| |
+--+ +----------- Gonidea_angulata
| |
+--+----------- Microcondylaea_bonellii
|
| +-------- Koreosolenaia_sitgyensis
+--+
| +----- Parvasolenaia_rivularis
+--+
| +-- Postolata_guangxiensis
+--+
+-- Ptychorhynchus_pfisteri
Cladogram Figure 6-7
+-- Margaritifera_dahurica
+-----------------------------+
| +-- Margaritifera_margaritifera
|
| +----------------------------- Chamberlainia_somsakpanhai
| |
| | +-- Khairuloconcha_lunbawangorum
| | +--------------+
--+ | | +-- Khairuloconcha_sahanae
| | |
| | | +-------- Ensidens_sagittarius
| | +--------+ +-----+
| | | | | | +----- Rectidens_sumatrensis
| | | | | +--+
| | | | | | +-- Hyriopsis_bialata
+--+ | | | +--+
| | +--+ +-- Hyriopsis_desowitzi
| | |
| | | +-- Physunio_superbus
| | | +--------+
| | | | +-- Trapezoideus_foliaceus
| | | |
| | +--+ +-- Lens_contradens
| | | +-----+
| | | | +-- Lens_eximius
| | +--+
| | | +----- Yaukthwa_elongatula
+--+ +--+
| | +-- Yaukthwa_inlenensis
| +--+
| +-- Yaukthwa_paiensis
|
| +----------------- Schepmania_nieuwenhuisi
| |
| +-----+ +-- Indopseudodon_bogani
| | | +-----------+
| | | | +-- Indopseudodon_kayinensis
| | +--+
| | | +----------- Pilsbryoconcha_exilis
| | | |
| | +--+ +-- Pseudodon_mekongi
| | | +-----+
| | | | +-- Pseudodon_vondembuschianus
| | +--+
+--+ | +----- Thaiconcha_callifera
| +--+
| | +-- Bineurus_anodontinum
| +--+
| +-- Bineurus_loeiensis
|
| +----- Potomida_littoralis
| +--------------+
| | | +-- Lamprotula_caveata
| | +--+
| | +-- Lamprotula_leaii
+--+
| +-- Inversidens_brandtii
| +--------------+
| | +-- Inversidens_rentianensis
+--+
| +-------------- Sinosolenaia_carinata
| |
+--+ +-- Gonidea_angulata
| +--------+
| | +-- Microcondylaea_bonellii
+--+
| +-------- Koreosolenaia_sitgyensis
+--+
| +----- Parvasolenaia_rivularis
+--+
| +-- Postolata_guangxiensis
+--+
+-- Ptychorhynchus_pfisteri
Cladogram Figure 8a_S1
+-------------------------------------------------- Neotrigonia_margaritacea
|
| +----------------------------------------------- Echyridella_menziesii
| |
--+ | +-- Anodontites_trapesialis
| | +-----------------------------------------+
| | | +-- Mutela_dubia
| | |
| | | +-- Margaritifera_dahurica
+--+ | +--------------------------------------+
| | | +-- Margaritifera_margaritifera
| | |
| | | +-- Quadrula_quadrula
| | | +--------------------------+
| | | | +-- Uniomerus_tetralasmus
| | | |
+--+ | +--------+ +-- Elliptio_complanata
| | | | +-----------------------+
| | | | | +-- Pleurobema_oviforme
| | | +--+
| | | | +----------------------- Amblema_plicata
| | | | |
| | | +--+ +-------------------- Popenaias_popeii
| | | | |
| | | +--+ +----------------- Toxolasma_parvum
| | | | |
| | | | | +----- Potamilus_leptodon
+--+ | +--+ +--------+
| | | | | +-- Potamilus_alatus
| | | | +--+
| | | | +-- Potamilus_streckersoni
| | +--+
| | | +----------- Leaunio_lienosus
| | | |
| | | | +----- Lampsilis_ornata
| | +--+ +--+
| | | | | +-- Lampsilis_cardium
| | | | +--+
| | +--+ +-- Lampsilis_cariosa
| | |
| | | +----- Venustaconcha_ellipsiformis
| | +--+
| | | +-- Lampsilis_powellii
| | +--+
| | +-- Lampsilis_siliquoidea
| |
+--+ +----- Chamberlainia_somsakpanhai
| +--------------------+
| | | +-- Sinohyriopsis_cumingii
| | +--+
| | +-- Sinohyriopsis_schlegelii
| |
| | +-- Hyriopsis_bialata
| | +--+
| +--------+ | +-- Rectidens_sumatrensis
| | | +-----------------+
| | | | | +-- Lens_contradens
| | | | +--+
| | | | +-- Physunio_superbus
| | | |
| | | | +----------- Potomida_littoralis
| | +--+ +--------+
| | | | | +-------- Pronodularia_japanensis
| | | | +--+
| | | | | +----- Lamprotula_leaii
| | | | +--+
| | | | | +-- Lamprotula_caveata
| | +--+ +--+
| | | +-- Lamprotula_cornuumlunae
| | |
| | | +-- Pseudodon_vondembuschianus
| | | +--------------+
| | | | +-- Pilsbryoconcha_exilis
| | +--+
| | | +-------------- Inversidens_rentianensis
| | | |
+--+ +--+ +-- Parvasolenaia_rivularis
| | +--------+
| | | +-- Ptychorhynchus_pfisteri
| +--+
| | +-------- Microcondylaea_bonellii
| +--+
| | +----- Sinosolenaia_carinata
| +--+
| | +-- Sinosolenaia_iridinea
| +--+
| +-- Sinosolenaia_oleivora
|
| +-------------------------------- Lamellidens_jenkinsianus
| |
| | +-- Lepidodesma_aligera
| | +--------------------------+
| | | +-- Lepidodesma_languilati
| | |
| | | +-------- Acuticosta_chinensis
| | | +--------------+
| | | | | +----- Lanceolaria_gladiola
| | | | +--+
+--+ | | | +-- Lanceolaria_grayii
| | | +--+
| | | +-- Lanceolaria_lanceolata
| | |
| | | +-------------- Cristaria_plicata
| | +--+ +-----+
| | | | | | +----------- Anemina_arcaeformis
| | | | | +--+
| | | | | | +-------- Beringiana_fukuharai
| | | | | +--+
+--+ | | | | +----- Sinanodonta_lucida
| | | | +--+
| | | | | +-- Sinanodonta_tumens
| | +--+ +--+
| | | +-- Sinanodonta_woodiana
| | |
| | | +-------- Pseudanodonta_complanata
| | | +--------+
| | | | | +----- Anodonta_anatina
| | | | +--+
| | | | | +-- Anodonta_cygnea
| | | | +--+
| | +--+ +-- Anodonta_exulcerata
| | |
| | | +-------------- Anodonta_nuttalliana
| | | |
| | | | +----- Pyganodon_grandis
+--+ +--+ +-----+
| | | | +-- Utterbackia_imbecillis
| | | +--+
| +--+ +-- Utterbackia_peninsularis
| |
| | +-------- Lasmigona_complanata
| +--+
| | +----- Prolasmidonta_heterodon
| +--+
| | +-- Alasmidonta_varicosa
| +--+
| +-- Platynaias_compressa
|
| +-------- Aculamprotula_tortuosa
| |
| +--------------+ +-- Aculamprotula_coreana
| | | +--+
| | | | +-- Aculamprotula_tientsinensis
| | +--+
| | | +-- Aculamprotula_polysticta
| | +--+
| | +-- Aculamprotula_scripta
| |
+--+ +-------------- Unio_tumidus
| +-----+
| | | +----------- Unio_nanus
| | +--+
| | | +-------- Unio_elongatulus
| | +--+
| | | +----- Unio_pictorum
| | +--+
| | | +-- Unio_delphinus
+--+ +--+
| +-- Unio_mancus
|
| +-- Nodularia_breviconcha
| +--------------+
| | +-- Nodularia_douglasiae
| |
| | +-- Pseudocuneopsis_capitata
| +--------------+
+--+ +-- Pseudocuneopsis_sichuanensis
|
| +-- Schistodesmus_lampreyanus
| +-----------+
| | +-- Schistodesmus_spinosus
+--+
| +----------- Tchangsinaia_piscicula
+--+
| +-------- Cuneopsis_rufescens
+--+
| +----- Cuneopsis_celtiformis
+--+
| +-- Cuneopsis_demangei
+--+
+-- Cuneopsis_heudei
Cladogram Figure 8b_S2
+-------------------------------------------------- Neotrigonia_margaritacea
|
| +----------------------------------------------- Echyridella_menziesii
| |
--+ | +-- Anodontites_trapesialis
| | +-----------------------------------------+
| | | +-- Mutela_dubia
| | |
| | | +-- Margaritifera_dahurica
+--+ | +--------------------------------------+
| | | +-- Margaritifera_margaritifera
| | |
| | | +-- Quadrula_quadrula
| | | +--------------------------+
| | | | +-- Uniomerus_tetralasmus
| | | |
+--+ | +--------+ +-- Elliptio_complanata
| | | | +-----------------------+
| | | | | +-- Pleurobema_oviforme
| | | +--+
| | | | +----------------------- Amblema_plicata
| | | | |
| | | +--+ +-------------------- Popenaias_popeii
| | | | |
| | | +--+ +----------------- Toxolasma_parvum
| | | | |
| | | | | +----- Potamilus_leptodon
+--+ | +--+ +--------+
| | | | | +-- Potamilus_alatus
| | | | +--+
| | +--+ +-- Potamilus_streckersoni
| | |
| | | +----------- Leaunio_lienosus
| | | |
| | +--+ +-------- Venustaconcha_ellipsiformis
| | | |
| | | | +-- Lampsilis_powellii
| | +--+-----+
| | | +-- Lampsilis_siliquoidea
| | |
| | | +----- Lampsilis_ornata
| | +--+
| | | +-- Lampsilis_cardium
| | +--+
| | +-- Lampsilis_cariosa
| |
+--+ +----- Chamberlainia_somsakpanhai
| +--------------------+
| | | +-- Sinohyriopsis_cumingii
| | +--+
| | +-- Sinohyriopsis_schlegelii
| |
| | +-- Hyriopsis_bialata
| | +--+
| +--------+ | +-- Rectidens_sumatrensis
| | | +-----------------+
| | | | | +-- Lens_contradens
| | | | +--+
| | | | +-- Physunio_superbus
| | | |
| | | | +----------- Potomida_littoralis
| | +--+ +--------+
| | | | | +-------- Pronodularia_japanensis
| | | | +--+
| | | | | +----- Lamprotula_leaii
| | | | +--+
| | | | | +-- Lamprotula_caveata
| | +--+ +--+
| | | +-- Lamprotula_cornuumlunae
| | |
| | | +-- Pseudodon_vondembuschianus
| | | +--------------+
| | | | +-- Pilsbryoconcha_exilis
| | +--+
| | | +-------------- Inversidens_rentianensis
| | | |
+--+ +--+ +-- Parvasolenaia_rivularis
| | +--------+
| | | +-- Ptychorhynchus_pfisteri
| +--+
| | +-------- Microcondylaea_bonellii
| +--+
| | +----- Sinosolenaia_carinata
| +--+
| | +-- Sinosolenaia_iridinea
| +--+
| +-- Sinosolenaia_oleivora
|
| +-------------------------------- Lamellidens_jenkinsianus
| |
| | +-- Lepidodesma_aligera
| | +--------------------------+
| | | +-- Lepidodesma_languilati
| | |
| | | +-------- Acuticosta_chinensis
| | | +--------------+
| | | | | +----- Lanceolaria_gladiola
| | | | +--+
+--+ | | | +-- Lanceolaria_grayii
| | | +--+
| | | +-- Lanceolaria_lanceolata
| | |
| | +--+ +-------------- Cristaria_plicata
| | | | +-----+
| | | | | | +----------- Anemina_arcaeformis
| | | | | +--+
| | | | | | +-------- Beringiana_fukuharai
| | | | | +--+
+--+ | | | | +----- Sinanodonta_lucida
| | | | +--+
| | +--+ | +-- Sinanodonta_tumens
| | | +--+
| | | +-- Sinanodonta_woodiana
| | |
| | | +----------------- Anodonta_nuttalliana
| | | |
| | | | +-------- Pseudanodonta_complanata
| | | | +-----+
| | +--+ | | +----- Anodonta_anatina
| | | | +--+
| | | | | +-- Anodonta_cygnea
| | | | +--+
| | +--+ +-- Anodonta_exulcerata
| | |
| | | +----- Pyganodon_grandis
+--+ | +-----+
| | | | +-- Utterbackia_imbecillis
| | | +--+
| +--+ +-- Utterbackia_peninsularis
| |
| | +-------- Lasmigona_complanata
| +--+
| | +----- Prolasmidonta_heterodon
| +--+
| | +-- Alasmidonta_varicosa
| +--+
| +-- Platynaias_compressa
|
| +-------- Aculamprotula_tortuosa
| |
| +--------------+ +-- Aculamprotula_coreana
| | | +--+
| | | | +-- Aculamprotula_tientsinensis
| | +--+
| | | +-- Aculamprotula_polysticta
| | +--+
| | +-- Aculamprotula_scripta
| |
+--+ +-------------- Unio_tumidus
| +-----+
| | | +----------- Unio_nanus
| | +--+
| | | +-------- Unio_elongatulus
| | +--+
| | | +----- Unio_pictorum
| | +--+
| | | +-- Unio_delphinus
+--+ +--+
| +-- Unio_mancus
|
| +-- Nodularia_breviconcha
| +--------------+
| | +-- Nodularia_douglasiae
| |
| | +-- Pseudocuneopsis_capitata
| +--------------+
+--+ +-- Pseudocuneopsis_sichuanensis
|
| +-- Schistodesmus_lampreyanus
| +-----------+
| | +-- Schistodesmus_spinosus
+--+
| +----------- Tchangsinaia_piscicula
+--+
| +-------- Cuneopsis_rufescens
+--+
| +----- Cuneopsis_celtiformis
+--+
| +-- Cuneopsis_demangei
+--+
+-- Cuneopsis_heudei
|
