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Thu 3 September 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

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Publication Dai et al. (2024): cladograms
     publication summary | cladograms

Complete Citation

Dai, Y.-T., Z.-G. Chen, C.-L. Hu, P.-F. Ning, S. Ouyang et al. 2024. Taxonomic reassessment of Scabies (Bivalvia: Unionidae) species in China based on multilocus and mitogenomic phylogenetic analyses. Invertebrate Systematics 38(6): 1-18. Available electronically.

Cladogram Figures

(click the figure number to see the cladogram)

Cladogram Figure 3
  +----------------------------- Scabies_scobinatus        
  |                                                        
--+  +-------------------------- Unio_crassus              
  |  |                                                     
  |  |                    +----- Scriptolamprotula_scripta 
  +--+  +-----------------+                                
     |  |                 |  +-- Anodonta_anatina          
     |  |                 +--+                             
     +--+                    +-- Lepidodesma_languilati    
        |                                                  
        |  +-------------------- Middendorffinaia_mongolica
        |  |                                               
        +--+  +----------------- Inversiunio_jokohamensis  
           |  |                                            
           +--+  +-------------- Cuneopsis_celtiformis     
              |  |                                         
              +--+  +----------- Nodularia_breviconcha     
                 |  |                                      
                 +--+        +-- Nodularia_fusiformans     
                    |  +-----+                             
                    |  |     +-- Nodularia_huana           
                    +--+                                   
                       |  +----- Nodularia_nuxpersicae     
                       +--+                                
                          |  +-- Nodularia_douglasiae      
                          +--+                             
                             +-- Nodularia_nipponensis     
Cladogram Figure 5-BI
                             +-- Margaritifera_margaritifera
  +--------------------------+                              
  |                          +-- Gibbosula_laosensis        
  |                                                         
  |                       +----- Trapezidens_exolescens     
--+  +--------------------+                                 
  |  |                    |  +-- Scabies_scobinatus         
  |  |                    +--+                              
  |  |                       +-- Scabies_crispata           
  |  |                                                      
  +--+  +----------------------- Acuticosta_chinensis       
     |  |                                                   
     |  +----------------------- Anemina_harlandi           
     |  |                                                   
     |  +----------------------- Anodonta_anatina           
     +--+                                                   
        |  +-------------------- Lanceolaria_grayii         
        |  |                                                
        |  |                 +-- Scriptolamprotula_scripta  
        |  +-----------------+                              
        +--+                 +-- Lepidodesma_languilati     
           |                                                
           |  +----------------- Unio_crassus               
           |  |                                             
           +--+  +-------------- Middendorffinaia_mongolica 
              |  |                                          
              |  |  +----------- Cuneopsis_celtiformis      
              +--+  |                                       
                 |  |        +-- Inversiunio_jokohamensis   
                 |  +--------+                              
                 |  |        +-- Schistodesmus_xinyuensis   
                 +--+                                       
                    |        +-- Nodularia_huana            
                    |  +-----+                              
                    |  |     +-- Nodularia_breviconcha      
                    |  |                                    
                    +--+     +-- Nodularia_nuxpersicae      
                       |  +--+                              
                       |  |  +-- Nodularia_nipponensis      
                       +--+                                 
                          |  +-- Nodularia_douglasiae       
                          +--+                              
                             +-- Nodularia_dualobtusa       
Cladogram Figure 5-ML
                                   +-- Margaritifera_margaritifera
  +--------------------------------+                              
  |                                +-- Gibbosula_laosensis        
  |                                                               
--+                             +----- Trapezidens_exolescens     
  |  +--------------------------+                                 
  |  |                          |  +-- Scabies_scobinatus         
  |  |                          +--+                              
  +--+                             +-- Scabies_crispata           
     |                                                            
     |  +----------------------------- Anodonta_anatina           
     |  |                                                         
     +--+                          +-- Acuticosta_chinensis       
        |  +-----------------------+                              
        |  |                       +-- Anemina_harlandi           
        +--+                                                      
           |  +----------------------- Lepidodesma_languilati     
           |  |                                                   
           +--+  +-------------------- Lanceolaria_grayii         
              |  |                                                
              +--+  +----------------- Scriptolamprotula_scripta  
                 |  |                                             
                 +--+  +-------------- Unio_crassus               
                    |  |                                          
                    |  |  +----------- Middendorffinaia_mongolica 
                    +--+  |                                       
                       |  |     +----- Cuneopsis_celtiformis      
                       |  |  +--+                                 
                       +--+  |  |  +-- Inversiunio_jokohamensis   
                          |  |  +--+                              
                          |  |     +-- Schistodesmus_xinyuensis   
                          |  |                                    
                          +--+     +-- Nodularia_nuxpersicae      
                             |  +--+                              
                             |  |  +-- Nodularia_nipponensis      
                             |  |                                 
                             |  |  +-- Nodularia_huana            
                             +--+--+                              
                                |  +-- Nodularia_breviconcha      
                                |                                 
                                |  +-- Nodularia_douglasiae       
                                +--+                              
                                   +-- Nodularia_dualobtusa       
Cladogram Figure 6
                             +-------- Quadrula_quadrula              
  +--------------------------+                                        
  |                          |  +----- Venustaconcha_ellipsiformis    
  |                          +--+                                     
  |                             |  +-- Lampsilis_powellii             
  |                             +--+                                  
  |                                +-- Lampsilis_siliquoidea          
  |                                                                   
  |                                +-- Chamberlainia_somsakpanhai     
  |                    +-----------+                                  
  |                    |           +-- Sinohyriopsis_cumingii         
  |                    |                                              
--+                    |     +-------- Potomida_littoralis            
  |  +-----------------+  +--+                                        
  |  |                 |  |  |  +----- Pronodularia_japanensis        
  |  |                 |  |  +--+                                     
  |  |                 |  |     |  +-- Lamprotula_caveata             
  |  |                 +--+     +--+                                  
  |  |                    |        +-- Lamprotula_leaii               
  |  |                    |                                           
  |  |                    |  +-------- Pilsbryoconcha_exilis          
  |  |                    +--+                                        
  |  |                       |  +----- Parvasolenaia_rivularis        
  +--+                       +--+                                     
     |                          |  +-- Sinosolenaia_carinata          
     |                          +--+                                  
     |                             +-- Sinosolenaia_oleivora          
     |                                                                
     |  +----------------------------- Lepidodesma_languilati         
     |  |                                                             
     |  |                 +----------- Lanceolaria_lanceolata         
     |  |  +--------------+                                           
     |  |  |              |  +-------- Acuticosta_chinensis           
     |  |  |              +--+                                        
     +--+  |                 |  +----- Sinanodonta_woodiana           
        |  |                 +--+                                     
        |  |                    |  +-- Pyganodon_grandis              
        |  |                    +--+                                  
        |  |                       +-- Utterbackia_peninsularis       
        |  |                                                          
        +--+                 +-------- Scriptolamprotula_tortuosa     
           |                 |                                        
           |  +--------------+     +-- Scriptolamprotula_polysticta   
           |  |              |  +--+                                  
           |  |              |  |  +-- Scriptolamprotula_scripta      
           |  |              +--+                                     
           |  |                 |  +-- Scriptolamprotula_coreana      
           |  |                 +--+                                  
           |  |                    +-- Scriptolamprotula_tientsinensis
           |  |                                                       
           +--+        +-------------- Unio_tumidus                   
              |  +-----+                                              
              |  |     |  +----------- Unio_nanus                     
              |  |     +--+                                           
              |  |        |  +-------- Unio_delphinus                 
              |  |        +--+                                        
              |  |           |  +----- Unio_pictorum                  
              |  |           +--+                                     
              +--+              |  +-- Unio_elongatulus               
                 |              +--+                                  
                 |                 +-- Unio_mancus                    
                 |                                                    
                 |  +----------------- Pseudocuneopsis_capitata       
                 |  |                                                 
                 |  |     +----------- Nodularia_breviconcha          
                 |  |  +--+                                           
                 +--+  |  |  +-------- Nodularia_fusiformans          
                    |  |  +--+                                        
                    |  |     |  +----- Nodularia_douglasiae           
                    |  |     +--+                                     
                    |  |        |  +-- Nodularia_nuxpersicae          
                    +--+        +--+                                  
                       |           +-- Nodularia_nipponensis          
                       |                                              
                       |           +-- Schistodesmus_lampreyanus      
                       |     +-----+                                  
                       |     |     +-- Schistodesmus_xinyuensis       
                       +-----+                                        
                             |  +----- Tchangsinaia_piscicula         
                             +--+                                     
                                |  +-- Cuneopsis_heudei               
                                +--+                                  
                                   +-- Cuneopsis_celtiformis          
 
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