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Thu 17 July 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

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Publication Wu et al. (2024): cladograms
     publication summary | cladograms

Complete Citation

Wu, R., L. Liu, L. Zhang, A.E. Bogan, G. Niu et al. 2024. Taxonomic revision of two species in the genus Ptychorhynchus Simpson, 1900 (Bivalvia: Unionidae: Gonideinae), with description of a new species. Invertebrate Systematics 38, IS24014: 1-17. Available electronically.

Cladogram Figures

(click the figure number to see the cladogram)

Cladogram Figure 4
     +-------- Nodularia_breviconcha      
     |                                    
  +--+     +-- Nodularia_douglasiae__1    
  |  |  +--+                              
  |  |  |  +-- Nodularia_dualobtusa       
  |  +--+                                 
  |     |  +-- Nodularia_douglasiae__2    
--+     +--+                              
  |        +-- Nodularia_nuxpersicae      
  |                                       
  |        +-- Postolata_guangxiensis     
  |  +-----+                              
  |  |     +-- Obovalis_omiensis          
  +--+                                    
     |  +----- Ptychorhynchus_pfisteri    
     +--+                                 
        |  +-- Cosmopseudodon_wenshanensis
        +--+                              
           +-- Cosmopseudodon_resupinatus 
Cladogram Figure 5
                                   +-- Margaritifera_dahurica      
  +--------------------------------+                               
  |                                +-- Margaritifera_margaritifera 
  |                                                                
  |  +-------------------------------- Chamberlainia_somsakpanhai  
  |  |                                                             
  |  |                             +-- Khairuloconcha_sahanae      
  |  |              +--------------+                               
--+  |              |              +-- Khairuloconcha_lunbawangorum
  |  |              |                                              
  |  |              |        +-------- Ensidens_sagittarius        
  |  |  +-----------+  +-----+                                     
  |  |  |           |  |     |  +----- Rectidens_sumatrensis       
  |  |  |           |  |     +--+                                  
  |  |  |           |  |        |  +-- Hyriopsis_bialata           
  +--+  |           |  |        +--+                               
     |  |           +--+           +-- Hyriopsis_desowitzi         
     |  |              |                                           
     |  |              |           +-- Physunio_superbus           
     |  |              |  +--------+                               
     |  |              |  |        +-- Trapezoideus_foliaceus      
     |  |              |  |                                        
     |  |              +--+        +-- Lens_contradens             
     |  |                 |  +-----+                               
     |  |                 |  |     +-- Lens_eximius                
     |  |                 +--+                                     
     +--+                    |  +----- Yaukthwa_elongatula         
        |                    +--+                                  
        |                       |  +-- Yaukthwa_inlenensis         
        |                       +--+                               
        |                          +-- Yaukthwa_paiensis           
        |                                                          
        |                       +----- Potomida_littoralis         
        |  +--------------------+                                  
        |  |                    |  +-- Lamprotula_leaii            
        |  |                    +--+                               
        |  |                       +-- Lamprotula_caveata          
        |  |                                                       
        |  |        +----------------- Schepmania_nieuwenhuisi     
        |  |        |                                              
        |  |  +-----+              +-- Indopseudodon_bogani        
        +--+  |     |  +-----------+                               
           |  |     |  |           +-- Indopseudodon_kayinensis    
           |  |     +--+                                           
           |  |        |  +----------- Pilsbryoconcha_exilis       
           |  |        |  |                                        
           |  |        +--+        +-- Pseudodon_mekongi           
           |  |           |  +-----+                               
           |  |           |  |     +-- Pseudodon_vondembuschianus  
           +--+           +--+                                     
              |              |  +----- Thaiconcha_callifera        
              |              +--+                                  
              |                 |  +-- Bineurus_anodontinum        
              |                 +--+                               
              |                    +-- Bineurus_loeiensis          
              |                                                    
              |  +-------------------- Sinosolenaia_carinata       
              |  |                                                 
              +--+                 +-- Gonidea_angulata            
                 |  +--------------+                               
                 |  |              +-- Microcondylaea_bonellii     
                 +--+                                              
                    |  +-------------- Koreosolenaia_sitgyensis    
                    |  |                                           
                    +--+  +----------- Parvasolenaia_rivularis     
                       |  |                                        
                       +--+  +-------- Postolata_guangxiensis      
                          |  |                                     
                          +--+     +-- Cosmopseudodon_resupinatus  
                             |  +--+                               
                             |  |  +-- Cosmopseudodon_wenshanensis 
                             +--+                                  
                                |  +-- Ptychorhynchus_pfisteri     
                                +--+                               
                                   +-- Obovalis_omiensis           
Cladogram Figure 6
                                   +-- Margaritifera_dahurica      
  +--------------------------------+                               
  |                                +-- Margaritifera_margaritifera 
  |                                                                
  |                                +-- Quadrula_quadrula           
  |              +-----------------+                               
  |              |                 +-- Uniomerus_tetralasmus       
  |              |                                                 
  |  +-----------+                 +-- Elliptio_complanata         
  |  |           |  +--------------+                               
  |  |           |  |              +-- Pleurobema_oviforme         
  |  |           +--+                                              
  |  |              |  +-------------- Popenaias_popeii            
  |  |              +--+                                           
--+  |                 |  +----------- Toxolasma_parvum            
  |  |                 +--+                                        
  |  |                    |  +-------- Potamilus_alatus            
  |  |                    +--+                                     
  |  |                       |  +----- Leaunio_lienosus            
  |  |                       +--+                                  
  |  |                          |  +-- Lampsilis_cardium           
  |  |                          +--+                               
  |  |                             +-- Venustaconcha_ellipsiformis 
  |  |                                                             
  |  |     +-------------------------- Lamellidens_marginalis      
  |  |     |                                                       
  |  |     |  +----------------------- Lepidodesma_languilati      
  |  |     |  |                                                    
  +--+  +--+  |        +-------------- Lanceolaria_lanceolata      
     |  |  |  |        |                                           
     |  |  |  |        |        +----- Anodonta_cygnea             
     |  |  |  |  +-----+  +-----+                                  
     |  |  +--+  |     |  |     |  +-- Alasmidonta_varicosa        
     |  |     |  |     |  |     +--+                               
     |  |     |  |     +--+        +-- Utterbackia_imbecillis      
     |  |     |  |        |                                        
     |  |     |  |        |  +-------- Cristaria_plicata           
     |  |     |  |        +--+                                     
     |  |     +--+           |  +----- Anemina_arcaeformis         
     |  |        |           +--+                                  
     |  |        |              |  +-- Beringiana_fukuharai        
     |  |        |              +--+                               
     |  |        |                 +-- Sinanodonta_lucida          
     |  |        |                                                 
     |  |        |  +----------------- Aculamprotula_tortuosa      
     |  |        |  |                                              
     |  |        +--+  +-------------- Unio_tumidus                
     |  |           |  |                                           
     |  |           +--+  +----------- Pseudocuneopsis_sichuanensis
     +--+              |  |                                        
        |              |  |     +----- Nodularia_breviconcha       
        |              +--+  +--+                                  
        |                 |  |  |  +-- Nodularia_douglasiae        
        |                 |  |  +--+                               
        |                 +--+     +-- Nodularia_nipponensis       
        |                    |                                     
        |                    |  +----- Tchangsinaia_piscicula      
        |                    +--+                                  
        |                       |  +-- Cuneopsis_heudei            
        |                       +--+                               
        |                          +-- Schistodesmus_lampreyanus   
        |                                                          
        |                       +----- Chamberlainia_somsakpanhai  
        |  +--------------------+                                  
        |  |                    |  +-- Sinohyriopsis_cumingii      
        |  |                    +--+                               
        |  |                       +-- Sinohyriopsis_schlegelii    
        |  |                                                       
        |  |                       +-- Hyriopsis_bialata           
        |  |                    +--+                               
        +--+                    |  +-- Rectidens_sumatrensis       
           |  +-----------------+                                  
           |  |                 |  +-- Lens_contradens             
           |  |                 +--+                               
           |  |                    +-- Physunio_superbus           
           |  |                                                    
           |  |           +----------- Potomida_littoralis         
           +--+  +--------+                                        
              |  |        |  +-------- Pronodularia_japanensis     
              |  |        +--+                                     
              |  |           |  +----- Lamprotula_leaii            
              |  |           +--+                                  
              |  |              |  +-- Lamprotula_caveata          
              |  |              +--+                               
              +--+                 +-- Lamprotula_cornuumlunae     
                 |                                                 
                 |                 +-- Pseudodon_vondembuschianus  
                 |  +--------------+                               
                 |  |              +-- Pilsbryoconcha_exilis       
                 |  |                                              
                 |  |        +-------- Microcondylaea_bonellii     
                 +--+  +-----+                                     
                    |  |     |  +----- Sinosolenaia_carinata       
                    |  |     +--+                                  
                    |  |        |  +-- Sinosolenaia_iridinea       
                    +--+        +--+                               
                       |           +-- Sinosolenaia_oleivora       
                       |                                           
                       |  +----------- Parvasolenaia_rivularis     
                       +--+                                        
                          |  +-------- Ptychorhynchus_pfisteri     
                          +--+                                     
                             |  +----- Postolata_guangxiensis      
                             +--+                                  
                                |  +-- Cosmopseudodon_resupinatus  
                                +--+                               
                                   +-- Cosmopseudodon_wenshanensis 
 
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