The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)
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Publication Dai et al. (2025): cladograms
publication summary | cladograms Complete CitationDai, Y.-T., Z.-G. Chen, S. Ouyang, X.-C. Huang & X.-P. Wu. 2025. A new tribe, genus, and species of freshwater mussel from the Changjiang River Basin in China (Bivlavia, Unionidae, Unioninae). Zoosystematics and Evolution 101(2): 779-790. Available electronically.
Cladogram Figures(click the figure number to see the cladogram) Cladogram Figure 1
+-- Gibbosula_rochechouartii
+--------------------------------------------------+
| +-- Margaritifera_dahurica
|
| +-- Pronodularia_japanensis
--+ +-----------------------------------------------+
| | +-- Lamprotula_leaii
| |
| | +-- Lampsilis_siliquoidea
+--+ +--------------------------------------------+
| | +-- Quadrula_quadrula
| |
| | +-- Anemina_harlandi
| | +-----------------------------------------+
+--+ | +-- Amuranodonta_kijaensis
| |
| | +----- Parreysia_nagpoorensis
| +--------------------------------------+
| | | +-- Coelatura_aegyptiaca
| | +--+
+--+ +-- Indonaia_andersoniana
|
| +----- Pletholophus_tenuis
| +-----------------------------------+
| | | +-- Sinanodonta_woodiana
| | +--+
| | +-- Beringiana_beringiana
| |
+--+ +-------------------------------------- Cristaria_plicata
| |
| | +-- Alasmidonta_marginata
| | +--+
| | | +-- Lasmigona_costata
| +--------------------------------+
+--+ | +-- Anodonta_anatina
| +--+
| +-- Pseudanodonta_complanata
|
| +-- Lanceolaria_gladiola
| +--------------------------------+
| | +-- Lanceolaria_oxyrhyncha
+--+
| +-------------------------------- Lepidodesma_languilati
| |
+--+ +----------------------------- Acuticosta_chinensis
| |
| | +-------------------------- Globunio_mirificus
+--+ |
| | +-------- Scriptolamprotula_tortuosa
| | |
| | +--------------+ +----- Scriptolamprotula_coreana
+--+ | | |
| | +--+----- Scriptolamprotula_fibrosa
| | |
| | | +-- Scriptolamprotula_polysticta
| | +--+
+--+ +-- Scriptolamprotula_scripta
|
| +-------------------- Middendorffinaia_mongolica
| |
| | +-- Nodularia_douglasiae
| | +--------------+
+--+ | +-- Nodularia_breviconcha
| |
| | +-- Cuneopsis_celtiformis
| | +-----------+
+--+ | +-- Cuneopsis_szechenyii
| |
| | +----- Diaurora_aurorea
| | |
| +--------+ +-- Schistodesmus_xinyuensis
| | | |
+--+ +--+-- Schistodesmus_spinosus
| |
| +-- Schistodesmus_tongpenensis
|
| +-- Pseudocuneopsis_sichuanensis
| +--------+
| | +-- Pseudocuneopsis_capitata
| |
+--+ +-- Unio_pictorum
| +-----+
| | +-- Unio_nanus
+--+
| +----- Tchangsinaia_piscicula
+--+
| +-- Inversiunio_yanagawensis
+--+
+-- Inversiunio_reinianus
Cladogram Figure 2
+-- Gibbosula_rochechouartii
+--------------------------------------------+
| +-- Margaritifera_dahurica
|
| +-- Pronodularia_japanensis
| +--+
| | +-- Lamprotula_leaii
--+ +--------------------------------------+
| | | +-- Lampsilis_siliquoidea
| | +--+
| | +-- Quadrula_quadrula
| |
| | +-- Coelatura_aegyptiaca
+--+ |
| +--------------------------------------+-- Indonaia_andersoniana
| | |
| | +-- Parreysia_nagpoorensis
| |
| | +-- Anemina_harlandi
+--+ +-----------------------------------+
| | +-- Amuranodonta_kijaensis
| |
| | +----- Pletholophus_tenuis
| | +-----------------------------+
+--+ | | +-- Sinanodonta_woodiana
| | +--+
| | +-- Beringiana_beringiana
| |
| | +-- Alasmidonta_marginata
+--+ +--+
| | +-- Lasmigona_costata
| +--------------------------+
| | | +-- Anodonta_anatina
| | +--+
| | +-- Pseudanodonta_complanata
| |
+--+ +----------------------------- Lanceolaria_gladiola
| |
| +----------------------------- Lanceolaria_oxyrhyncha
| |
| +----------------------------- Cristaria_plicata
+--+
| +-------------------------- Acuticosta_chinensis
| |
| +-------------------------- Lepidodesma_languilati
| |
+--+ +----------------------- Globunio_mirificus
| |
| | +-------- Scriptolamprotula_tortuosa
| | |
| | +-----------+ +-- Scriptolamprotula_fibrosa
+--+ | | +--+
| | | | +-- Scriptolamprotula_coreana
| | +--+
| | | +-- Scriptolamprotula_scripta
| | +--+
| | +-- Scriptolamprotula_polysticta
+--+
| +----- Middendorffinaia_mongolica
| +-----------+
| | | +-- Nodularia_douglasiae
| | +--+
| | +-- Nodularia_breviconcha
| |
| | +-- Cuneopsis_celtiformis
+--+ +-----------+
| | +-- Cuneopsis_szechenyii
| |
| | +-------- Diaurora_aurorea
| +-----+
| | | +----- Schistodesmus_spinosus
| | +--+
+--+ | +-- Schistodesmus_xinyuensis
| +--+
| +-- Schistodesmus_tongpenensis
|
| +-- Pseudocuneopsis_sichuanensis
| +--------+
| | +-- Pseudocuneopsis_capitata
| |
+--+ +-- Inversiunio_yanagawensis
| +-----+
| | +-- Inversiunio_reinianus
+--+
| +----- Tchangsinaia_piscicula
+--+
| +-- Unio_pictorum
+--+
+-- Unio_nanus
Cladogram Figure 3
+-- Anodontites_trapesialis
+--------------------------------------------+
| +-- Mutela_dubia
|
| +-------------------------------------------- Echyridella_menziesii
| |
| | +-------------- Gibbosula_rochechouartii
| | +--------------------------+
--+ | | | +----------- Cumberlandia_monodonta
| | | +--+
| | | | +-------- Pseudunio_marocanus
| | | +--+
| | | | +----- Margaritifera_falcata
| | | +--+
| | | | +-- Margaritifera_dahurica
| | | +--+
+--+ | +-- Margaritifera_margaritifera
| |
| | +-- Quadrula_quadrula
| | +--------------------------+
| | | +-- Uniomerus_tetralasmus
| | |
| | +--------+ +-- Elliptio_complanata
| | | | +-----------------------+
| | | | | +-- Pleurobema_oviforme
| | | +--+
| | | | +----------------------- Amblema_plicata
| | | | |
+--+ | +--+ +-------------------- Popenaias_popeii
| | | |
| | +--+ +----------------- Toxolasma_parvum
| | | |
| | | | +----- Potamilus_leptodon
| | +--+ +--------+
| | | | | +-- Potamilus_streckersoni
| | | | +--+
| | +--+ +-- Potamilus_alatus
| | |
| | | +----------- Leaunio_lienosus
| | | |
| | +--+ +-- Lampsilis_cardium
| | | +-----+
| | | | +-- Lampsilis_ornata
| | +--+
| | | +----- Venustaconcha_ellipsiformis
| | +--+
| | | +-- Lampsilis_powellii
| | +--+
| | +-- Lampsilis_siliquoidea
| |
+--+ +----- Chamberlainia_somsakpanhai
| +-----------------+
| | | +-- Sinohyriopsis_cumingii
| | +--+
| | +-- Sinohyriopsis_schlegelii
| |
| | +-- Lens_contradens
| +-----------+ +--+
| | | | +-- Physunio_superbus
| | | +--------------+
| | | | | +-- Hyriopsis_bialata
| | | | +--+
| | | | +-- Rectidens_sumatrensis
| | | |
| | +--+ +-------- Potomida_littoralis
| | | +--------+
| | | | | +----- Pronodularia_japanensis
| | | | +--+
| | | | | +-- Lamprotula_caveata
| | | | +--+
| | +--+ +-- Lamprotula_leaii
| | |
| | | +-- Pseudodon_mekongi
| | | +-----------+
| | | | +-- Pilsbryoconcha_exilis
| | | |
| | +--+ +----------- Inversidens_rentianensis
| | | |
| | | | +----- Parvasolenaia_rivularis
+--+ +--+ +--+
| | | | +-- Postolata_guangxiensis
| | | +--+
| +--+ +-- Ptychorhynchus_pfisteri
| |
| | +----- Microcondylaea_bonellii
| +--+
| | +-- Sinosolenaia_carinata
| +--+
| +-- Sinosolenaia_oleivora
|
| +-------------------------------- Lepidodesma_languilati
| |
| | +-------- Acuticosta_chinensis
| | +--------+
| | | | +----- Lanceolaria_gladiola
| | | +--+
| | | | +-- Lanceolaria_lanceolata
| | | +--+
| | | +-- Lanceolaria_grayii
| | +-----------+
| | | | +-------- Cristaria_plicata
+--+ | | +-----+
| | | | | +----- Anemina_harlandi
| | | | +--+
| | | | | +-- Sinanodonta_lucida
| | | | +--+
| | +--+ +-- Sinanodonta_woodiana
| | |
| | | +-- Anodonta_anatina
| | | +--------+
| | | | +-- Anodonta_cygnea
| | +--+
| | | +-------- Platynaias_compressa
+--+ +--+
| | +----- Pyganodon_grandis
| +--+
| | +-- Utterbackia_imbecillis
| +--+
| +-- Utterbackia_peninsularis
|
| +-------------------------- Globunio_mirificus
| |
| | +-------- Scriptolamprotula_tortuosa
| | |
| | +--------------+ +-- Scriptolamprotula_polysticta
| | | | +--+
| | | | | +-- Scriptolamprotula_scripta
+--+ | +--+
| | | +-- Scriptolamprotula_coreana
| | +--+
| | +-- Scriptolamprotula_tientsinensis
| |
| | +-------------- Unio_tumidus
| | +-----+
+--+ | | +----------- Unio_nanus
| | +--+
| | | +-------- Unio_delphinus
| | +--+
| | | +----- Unio_pictorum
| | +--+
| | | +-- Unio_elongatulus
| | +--+
| | +-- Unio_mancus
| |
+--+ +-------- Nodularia_breviconcha
| +--+
| | | +----- Nodularia_fusiformans
| | +--+
| | | +-- Nodularia_nuxpersicae
| +-----+ +--+
| | | +-- Nodularia_douglasiae
| | |
| | | +-------- Pseudocuneopsis_capitata
| | +--+
| | | +----- Pseudocuneopsis_yangshuoensis
| | +--+
+--+ | +-- Pseudocuneopsis_sichuanensis
| +--+
| +-- Pseudocuneopsis_yemaoi
|
| +----- Schistodesmus_spinosus
| +--------+
| | | +-- Schistodesmus_xinyuensis
| | +--+
+--+ +-- Schistodesmus_tongpenensis
|
| +----------- Tchangsinaia_piscicula
+--+
| +-------- Cuneopsis_rufescens
+--+
| +----- Cuneopsis_celtiformis
+--+
| +-- Cuneopsis_heudei
+--+
+-- Cuneopsis_demangei
|
