The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)
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Bivalvia | Unionida | Unionoidea | Unionidae | Ambleminae | Lampsilini genus Epioblasma Rafinesque, 1831: cladomics
28 Recent valid species (29 total species) Cladomics Summarynumber of species = 28 (29 total)
analyzed in 38 cladograms in 26 publications
maximum number of terminals in an analysis =
7 (mean = 2.21, median = 1) Basic Monophylynumber of cladograms that test basic monophyly
(i.e., more than 1 included species) = 14
maximum number of included terminals = 7, (mean =
4.29 median = 4)
proportion of cladograms that support basic monophyly
= 1
Pairwise Cladogram ClusteringThe graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.
Buhay et al., 2002 [Publ 2661], fig. 2B ** Top Monophyly Cladogram ** iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 3 | 7 | 3 | 3 | 3 | 0 | +-------------- Lampsilis
|
| +-- Cambarunio
+-----------+
--+ +-- Villosa
|
| +----------- Epioblasma_brevidens **
| |
+--+ +-------- Epioblasma_triquetra **
| |
+--+ +----- Epioblasma_capsaeformis__3 **
| |
+--+ +-- Epioblasma_capsaeformis__1 **
| |
| +-- Epioblasma_capsaeformis__2 **
+--+
+-- Epioblasma_walkeri__1 **
|
+-- Epioblasma_walkeri__2 **
Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14 ** Top Sister-Group Cladogram ** iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 4 | 6 | 627 | 88 | 26 | 6 | +-------- Potamilus_inflatus
+--------+
| | +----- Potamilus_ohiensis
| +--+
| | +-- Potamilus_amphichaenus
| +--+
+-----------------------------------+ +-- Potamilus_streckersoni
| |
| | +----- Potamilus_fragilis__2
| | +--------+
| | | | +-- Potamilus_leptodon
| | | +--+
| +--+ +-- Potamilus_fragilis__1
| |
| | +----------- Potamilus_metnecktayi
| | |
| +--+ +-------- Potamilus_alatus__3
| | |
| +--+ +----- Potamilus_alatus__2
--+ | |
| +--+----- Potamilus_purpuratus__2
| |
| | +-- Potamilus_alatus__1
| +--+
| +-- Potamilus_purpuratus__1
|
| +-- Villosa_villosa
| +-----------------------------------------------+
| | +-- Atlanticoncha
| |
| | +-- Cambarunio_nebulosus
| | +--------------------------------------------+
+--+ | +-- Ligumia
| |
| | +-------- Leaunio_umbrans
| | +-----------------------------------+
| | | | +----- Cambarunio_taeniatus
+--+ | +--+
| | | +-- Cambarunio_iris
| | +--+
| | +-- Cambarunio_simus
| |
| | +----------------------------------------- Hamiota
| | |
+--+ | +----------------------- Pachynaias
| | |
| | +--------------+ +-------------------- Sagittunio
| | | | |
| | | | | +----- Ellipsaria
| | | +--+ +-----------+
| | | | | | +-- Potamilus_capax
| | | | | +--+
| | | | | +-- Truncilla
| | | +--+
+--+ | | +----- Medionidus_acutissimus
| | | +--------+
| | | | | +-- Lemiox
| | | | +--+
| | +--+ +-- Medionidus_penicillatus
| | |
| | | +----------- Ptychobranchus
| | | |
| | +--+ +-- Cyprogenia_aberti__2
| | | +-----+
| | | | +-- Dromus
| | +--+
+--+ | +----- Cyprogenia_stegaria__2
| +--+
| | +-- Cyprogenia_aberti__1
| +--+
| +-- Cyprogenia_stegaria__1
|
| +-- Villosa_delumbis
| +--------------------------------+
| | +-- Villosa_vibex
| |
| | +-------------------- Obovaria_olivaria
| | |
| | | +-------- Obovaria_retusa
| | +-----------+ +--------+
| | | | | | +----- Obovaria_choctawensis
| | | | | +--+
| | | | | | +-- Obovaria_unicolor
| | | +--+ +--+
| | | | +-- Obovaria_subrotunda
| | | |
+--+ | | +-------------- Venustaconcha
| | | |
| | +--+ +-- Epioblasma_brevidens **
| | | +--------+
| | | | +-- Epioblasma_triquetra **
| | +--+
| | | +-------- Epioblasma_rangiana__2 **
| | +--+
| | | +----- Epioblasma_rangiana__1 **
| | +--+
| | | +-- Epioblasma_capsaeformis **
| | +--+
| | +-- Epioblasma_walkeri **
| |
+--+ +-- Mesonaias
| +-----------------------+
| | +-- Obovaria_arkansasensis
| |
| +--+ +-- Lampsilis_splendida
| | | +--------------------+
| | | | +-- Lampsilis_bracteata
| | | |
| | +--+ +-- Lampsilis_fasciola
| | | +-----------------+
| | | | +-- Ortmanniana_pectorosa
| | +--+
| | | +----------------- Lampsilis_ornata
| | | |
| | +--+ +-------------- Lampsilis_ovata__2
| | | |
| | +--+ +----------- Lampsilis_dolabraeformis
| | | |
| | | | +----- Lampsilis_cardium__2
| | +--+ +--+
| | | | | +-- Lampsilis_cardium__1
+--+ | | +--+
| +--+ +-- Lampsilis_satura
| |
| | +----- Lampsilis_cariosa__2
| +--+
| | +-- Lampsilis_ovata__1
| +--+
| +-- Lampsilis_cariosa__1
|
| +----------- Ortmanniana_ligamentina__2
| +--------------+
| | | +-------- Lampsilis_abrupta
| | +--+
| | | +----- Lampsilis_higginsii__2
| | +--+
| | | +-- Ortmanniana_ligamentina__1
| | +--+
+--+ +-- Lampsilis_higginsii__1
|
| +----- Medionidus_conradicus
| +-----------------+
| | | +-- Leaunio_lienosus
| | +--+
| | +-- Leaunio_vanuxemensis
| |
+--+ +-- Lampsilis_floridensis
| +-----------------+
| | +-- Lampsilis_teres
| |
| | +-- Lampsilis_siliquoidea__1
+--+ |
| +-- Lampsilis_siliquoidea__2
| +--------------+
| | +-- Lampsilis_powellii__1
| | |
+--+ +-- Lampsilis_powellii__2
|
| +-------------- Lampsilis_sietmani
| |
+--+ +-- Lampsilis_straminea__2
| +--------+
| | +-- Lampsilis_virescens
+--+
| +-------- Lampsilis_hydiana__2
+--+
| +----- Lampsilis_hydiana__1
+--+
| +-- Lampsilis_bergmanni
+--+
+-- Lampsilis_straminea__1
Jones & Neves, 2010 [Publ 2646], fig. 16 iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 7 | 5 | 7 | 0 | 0 | 0 | 7 | +-- Epioblasma_triquetra **
+-----------+
| +-- Epioblasma_brevidens **
--+
| +----------- Epioblasma_rangiana **
+--+
| +-------- Epioblasma_capsaeformis **
+--+
| +----- Epioblasma_ahlstedti **
+--+
| +-- Epioblasma_aureola **
+--+
+-- Epioblasma_walkeri **
Jones & Neves, 2010 [Publ 2646], fig. 17 iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 7 | 1 | 7 | 0 | 0 | 0 | 6 | +----- Epioblasma_triquetra **
|
+----- Epioblasma_brevidens **
|
--+ +-- Epioblasma_capsaeformis **
| |
| +-- Epioblasma_aureola **
| |
+--+-- Epioblasma_walkeri **
|
+-- Epioblasma_rangiana **
|
+-- Epioblasma_ahlstedti **
Zanatta & Murphy, 2006 [Publ 2672], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 5 | 51 | 43 | 22 | 4 | +-- Cyrtonaias
+-----------------------------------------+
| +-- Glebula
|
--+ +----------------------------------------- Paetulunio
| |
| | +-- Obliquaria
+--+ +-----------------------------------+
| | +-- Toxolasma
| |
| | +----------------------------------- Sagittunio
+--+ |
| | +----------- Potamilus
| | |
| | +--------------------+ +-- Medionidus_conradicus
| | | | +-----+
+--+ | | | +-- Ptychobranchus
| | +--+
| | | +----- Medionidus_acutissimus
| | | |
| | +--+----- Lemiox
| | |
| | | +-- Dromus
+--+ +--+
| +-- Cyprogenia
|
| +----- Lampsilis_teres
| +-----------------------+
| | | +-- Truncilla
| | +--+
| | +-- Ellipsaria
| |
+--+ +-------------------------- Leaunio
| |
| | +----------- Hamiota
| | |
| | +-----------+ +-- Lampsilis_fasciola
+--+ | | +-----+
| | | | +-- Ortmanniana_pectorosa
| | +--+
| | | +----- Lampsilis_ornata
| | +--+
| | | +-- Lampsilis_ovata
+--+ +--+
| +-- Lampsilis_cardium
|
| +-------- Lampsilis_siliquoidea
| +-----------+
| | | +----- Ortmanniana_ligamentina
| | +--+
| | | +-- Cambarunio
+--+ +--+
| +-- Ligumia
|
| +----------------- Venustaconcha
+--+
| +-------------- Obovaria
+--+
| +----------- Epioblasma_brevidens **
+--+
| +-------- Epioblasma_triquetra **
+--+
| +----- Epioblasma_rangiana **
+--+
| +-- Epioblasma_walkeri **
+--+
+-- Epioblasma_capsaeformis **
Jones et al., 2006 [Publ 2681], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 4 | 6 | 0 | 0 | 0 | 6 | +-- Epioblasma_triquetra **
+--------+
| +-- Epioblasma_brevidens **
--+
| +-------- Epioblasma_capsaeformis__2 **
+--+
| +----- Epioblasma_capsaeformis__1 **
+--+
| +-- Epioblasma_walkeri **
+--+
+-- Epioblasma_rangiana **
Zanatta & Murphy, 2006 [Publ 2672], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 5 | 52 | 43 | 22 | 4 | +-- Paetulunio
+-----------------------------------------+
| +-- Toxolasma
|
| +----- Obliquaria
| +-----------------------------------+
--+ | | +-- Glebula
| | +--+
| | +-- Cyrtonaias
| |
| | +----- Ptychobranchus
| | +--+
+--+ | | +-- Dromus
| | +--+
| +-----------------------------+ +-- Cyprogenia
| | |
| | | +----- Medionidus_conradicus
| | +--+
| | | +-- Medionidus_acutissimus
| | +--+
+--+ +-- Lemiox
|
| +----------------------------------- Sagittunio
| |
| +----------------------------------- Potamilus
| |
| | +-- Truncilla
+--+ +-----------------------------+
| | +-- Ellipsaria
| |
| | +-- Ortmanniana_pectorosa
| | +-----+
| | | +-- Lampsilis_fasciola
+--+ +--------------------+
| | | +----- Lampsilis_ornata
| | +--+
| | | +-- Lampsilis_ovata
| | +--+
| | +-- Lampsilis_cardium
+--+
| +-- Ligumia
| +--+
| | +-- Cambarunio
| +--------------------+
| | | +-- Lampsilis_teres
| | +--+
+--+ +-- Hamiota
|
| +----------------------- Leaunio
| |
+--+ +-------------------- Ortmanniana_ligamentina
| |
+--+ +----------------- Lampsilis_siliquoidea
| |
| | +-------------- Obovaria_olivaria
+--+ |
| +-------------- Venustaconcha
| |
| | +----- Obovaria_unicolor
+--+--------+
| | +-- Obovaria_subrotunda
| +--+
| +-- Obovaria_arkansasensis
|
| +----------- Epioblasma_brevidens **
+--+
| +-------- Epioblasma_triquetra **
+--+
| +----- Epioblasma_rangiana **
+--+
| +-- Epioblasma_walkeri **
+--+
+-- Epioblasma_capsaeformis **
Campbell et al., 2005 [Publ 2660], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 3 | 5 | 98 | 39 | 22 | 5 | +----- Obliquaria
+-----------+
| | +-- Cyrtonaias
| +--+
| +-- Glebula
|
| +-------------- Sagittunio
| |
| +-------------- Leaunio
| |
| +-------------- Potamilus
| |
| +-------------- Hamiota
| |
| | +-- Ellipsaria
| +-----------+
--+ | +-- Truncilla
| |
| | +----- Lampsilis_teres
| +--------+
| | | +-- Ortmanniana_ligamentina
| | +--+
| | +-- Lampsilis_siliquoidea
| |
| | +----- Villosa
| +--------+
| | | +-- Ligumia
| | +--+
| | +-- Cambarunio
| |
| | +-------- Ortmanniana_pectorosa
| +-----+
+--+ | +----- Lampsilis_ornata
| +--+
| | +-- Lampsilis_cardium
| +--+
| +-- Lampsilis_ovata
|
| +-- Ptychobranchus
| +-----+
| | +-- Venustaconcha_ellipsiformis__1
| |
+-----+ +----- Medionidus_conradicus
| | |
| | | +-- Lemiox
| +--+--+
| | +-- Medionidus_acutissimus
| |
| | +-- Dromus
| +--+
| +-- Cyprogenia
|
| +----------- Obovaria_olivaria
| |
| +----------- Venustaconcha_ellipsiformis__2
| |
| | +----- Obovaria_unicolor
| +-----+
+--+ | +-- Obovaria_arkansasensis
| +--+
| +-- Obovaria_subrotunda
|
| +-- Epioblasma_brevidens **
| +-----+
| | +-- Epioblasma_triquetra **
+--+
| +----- Epioblasma_walkeri__1 **
+--+
| +-- Epioblasma_walkeri__2 **
+--+
+-- Epioblasma_capsaeformis **
Buhay et al., 2002 [Publ 2661], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 5 | 1 | 1 | 1 | 3 | +----------- Lampsilis_siliquoidea__1
|
+----------- Lampsilis_siliquoidea__2
--+
| +-------- Epioblasma_brevidens **
| |
+--+ +----- Epioblasma_triquetra **
| |
+--+ +-- Epioblasma_capsaeformis **
| |
+--+-- Epioblasma_walkeri__1 **
|
+-- Epioblasma_walkeri__2 **
Buhay et al., 2002 [Publ 2661], fig. 2A iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 1 | 7 | 3 | 3 | 3 | 3 | +-------- Lampsilis
|
| +-- Cambarunio
+-----+
| +-- Villosa
--+
| +----- Epioblasma_triquetra **
| |
| +----- Epioblasma_brevidens **
| |
+--+ +-- Epioblasma_capsaeformis__1 **
| |
| +-- Epioblasma_capsaeformis__2 **
| |
+--+-- Epioblasma_capsaeformis__3 **
|
+-- Epioblasma_walkeri__1 **
|
+-- Epioblasma_walkeri__2 **
Campbell et al., 2005 [Publ 2660], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 3 | 5 | 98 | 39 | 22 | 3 | +-------- Toxolasma
+-----------------------+
| | +----- Obliquaria
| +--+
| | +-- Cyrtonaias
| +--+
| +-- Glebula
|
| +-------------- Potamilus
| |
| | +-------- Medionidus_acutissimus
--+ | +--+
| +--------------+ | | +----- Lemiox
| | | | +--+
| | | | | +-- Dromus
| | | | +--+
| | +--+ +-- Cyprogenia
| | |
| | | +-- Sagittunio
| | | +--+
| | | | +-- Medionidus_conradicus
| | +-----+
+--+ | +-- Ptychobranchus
| +--+
| +-- Venustaconcha_ellipsiformis__2
|
| +-- Ellipsaria
| +-----------------------+
| | +-- Truncilla
| |
| | +----------------------- Villosa
| | |
+--+ | +-- Obovaria_olivaria
| | +--------------+
| | | +-- Venustaconcha_ellipsiformis__1
| | |
| | +--+ +----- Obovaria_unicolor
| | | | +--------+
+--+ | | | | +-- Obovaria_arkansasensis
| | | | +--+
| | +--+ +-- Obovaria_subrotunda
| | |
| | | +----------- Epioblasma_brevidens **
| | +--+
| | | +-------- Epioblasma_triquetra **
| | +--+
| | | +----- Epioblasma_walkeri__2 **
+--+ +--+
| | +-- Epioblasma_capsaeformis **
| +--+
| +-- Epioblasma_walkeri__1 **
|
| +-------- Hamiota
| |
| +-----+ +-- Lampsilis_cardium
| | | +--+
| | | | +-- Lampsilis_ovata
| | +--+
| | | +-- Ortmanniana_pectorosa
+-----+ +--+
| +-- Lampsilis_ornata
|
| +-- Ligumia
| +--------+
| | +-- Cambarunio
+--+
| +-------- Lampsilis_teres
+--+
| +----- Ortmanniana_ligamentina
+--+
| +-- Lampsilis_siliquoidea
+--+
+-- Leaunio
Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 28 | 10 | 8 | -- | +----------- Ligumia
|
+----------- Truncilla
|
| +-- Ptychobranchus
+--------+
--+ +-- Sagittunio
|
| +----- Lampsilis_siliquoidea
| +--+
| | | +-- Epioblasma_triquetra **
| | +--+
| | +-- Epioblasma_brevidens **
+--+
| +----- Ortmanniana
| |
| +----- Cambarunio
+--+
+----- Leaunio
|
| +-- Lampsilis_cardium
+--+
+-- Lampsilis_fasciola
Grobler et al., 2011 [Publ 3359], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 15 | 14 | 9 | -- | +-------- Lemiox
+-----+
| | +----- Medionidus
| +--+
| | +-- Dromus
| +--+
--+ +-- Cyprogenia
|
| +----------- Potamilus
| |
| | +----- Lampsilis_siliquoidea
| | +--+
+--+ | | +-- Lampsilis_higginsii
| | +--+
| | +-- Ortmanniana
| |
| | +----- Lampsilis_fasciola
+--+--+
| | +-- Lampsilis_cardium
| +--+
| +-- Lampsilis_ovata
|
| +----- Cambarunio
| |
+--+----- Ligumia
|
| +-- Epioblasma_brevidens **
+--+
+-- Epioblasma_capsaeformis **
Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 28 | 10 | 8 | -- | +-------------------- Truncilla
|
--+ +----------------- Ligumia
| |
| | +-- Ptychobranchus
+--+ +-----------+
| | +-- Sagittunio
| |
+--+ +----------- Lampsilis_siliquoidea
| |
| | +----- Ortmanniana
+--+ +--+
| | | +-- Lampsilis_cardium
| | +--+
| | +-- Lampsilis_fasciola
+--+
| +-- Epioblasma_brevidens **
| +--+
| | +-- Epioblasma_triquetra **
+--+
| +-- Cambarunio
+--+
+-- Leaunio
Boyer et al., 2011 [Publ 2718], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 34 | 15 | 9 | -- | +----------- Toxolasma
|
| +-------- Obliquaria
--+ |
| | +----- Obovaria
| | |
| | +----- Lampsilis_cardium
+--+ |
| +----- Potamilus
| |
| | +-- Epioblasma_triquetra **
| +--+
| | +-- Lampsilis_siliquoidea
+--+
| +-- Ortmanniana
+--+
| +-- Lampsilis_higginsii
|
| +-- Ligumia
+--+
| +-- Lampsilis_teres
|
| +-- Ellipsaria
+--+
+-- Truncilla
Baker et al., 2004 [Publ 2685], fig. 2-3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 8 | 0 | 0 | -- | -- Epioblasma_triquetra **
Plazzi et al., 2011 [Publ 3676], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 17 | 2 | 2 | -- | +----- Lampsilis
--+
| +-- Epioblasma_rangiana **
+--+
+-- Venustaconcha
Inoue et al., 2020 [Publ 4128], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 22 | 11 | 11 | -- | +-- Glebula
+--------+
| +-- Cyrtonaias
|
--+ +-------- Potamilus
| |
| | +-- Ptychobranchus
+--+ +--+
| | +-- Truncilla
| |
| | +-- Ortmanniana
+--+ |
| +-- Epioblasma_triquetra **
| |
| +-- Lampsilis
| |
+--+-- Ligumia
|
+-- Cambarunio
|
+-- Pachynaias
|
+-- Mesonaias
Boyer et al., 2011 [Publ 2718], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 37 | 15 | 9 | -- | +----------------- Potamilus
|
| +-- Ellipsaria
--+ +-----------+
| | +-- Truncilla
| |
+--+ +----------- Lampsilis_cardium
| |
| | +----- Ligumia
| +-----+
+--+ | +-- Obovaria
| +--+
| +-- Epioblasma_triquetra **
|
| +-------- Lampsilis_teres
+--+
| +----- Lampsilis_siliquoidea
+--+
| +-- Ortmanniana
+--+
+-- Lampsilis_higginsii
Ouyang et al., 2015 [Publ 4493], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 30 | 2 | 2 | -- | +-- Epioblasma_triquetra **
--+
+-- Lampsilis
Baker et al., 2003 [Publ 2690], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 7 | 0 | 0 | -- | -- Epioblasma_triquetra **
Hoeh et al., 2009 [Publ 2716], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 45 | 6 | 6 | -- | +----- Truncilla
|
+----- Ptychobranchus
--+
| +-- Ortmanniana
| |
| +-- Cambarunio
+--+
+-- Ligumia
|
+-- Epioblasma_triquetra **
Inoue et al., 2020 [Publ 4128], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 21 | 10 | 10 | -- | +----------------------- Cyrtonaias
|
--+ +-------------------- Ptychobranchus
| |
+--+ +----------------- Ligumia
| |
+--+ +-------------- Potamilus
| |
+--+ +----------- Pachynaias
| |
| | +-- Epioblasma_triquetra **
+--+ +-----+
| | +-- Truncilla
| |
+--+ +-- Mesonaias
| +--+
| | +-- Cambarunio
+--+
| +-- Ortmanniana
+--+
+-- Lampsilis
Ouyang et al., 2015 [Publ 4493], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 30 | 2 | 2 | -- | +----- Cyrtonaias
--+
| +-- Lampsilis
+--+
+-- Epioblasma_triquetra **
Inoue et al., 2018 [Publ 3858], fig. 1_3a-c iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 104 | 10 | 7 | -- | -- Epioblasma_rangiana **
Campbell & Lydeard, 2012 [Publ 3309], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 72 | 14 | 10 | -- | +----------------- Obliquaria
|
| +----- Potamilus
--+ +--------+
| | | +-- Dromus
| | +--+
+--+ +-- Lemiox
|
| +----------- Lampsilis_ovata
| |
+--+ +-------- Lampsilis_virescens
| |
+--+ +-- Epioblasma_rangiana **
| +--+
| | +-- Obovaria_arkansasensis
+--+
| +-- Obovaria_olivaria
+--+
+-- Venustaconcha
Szumowski et al., 2012 [Publ 3569], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 15 | 14 | 9 | -- | +----------------------- Ellipsaria
|
--+ +-------------------- Truncilla
| |
+--+ +----------------- Obliquaria
| |
| | +-------------- Potamilus
+--+ |
| | +----- Lampsilis_ornata
| | +-----+
| | | | +-- Lampsilis_cardium
+--+ | +--+
| | +-- Lampsilis_ovata
| |
| | +-------- Lampsilis_teres
| +--+
+--+ | +----- Lampsilis_siliquoidea
| +--+
| | +-- Ortmanniana
| +--+
| +-- Lampsilis_higginsii
|
| +----- Hamiota
| |
+-----+ +-- Epioblasma_triquetra **
| |
+--+-- Cambarunio
|
+-- Ligumia
Graf & Cummings, 2006 [Publ 1968], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 46 | 7 | 7 | -- | +-------------- Obliquaria
|
+-------------- Truncilla
--+
+-------------- Ptychobranchus
|
| +----------- Ligumia
+--+
| +-------- Cambarunio
+--+
| +----- Epioblasma_triquetra **
+--+
| +-- Lampsilis
+--+
+-- Ortmanniana
Jadhav & Jamkhedkar, 2009 [Publ 3697], fig. 11 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 24 | 7 | 7 | -- | +----------- Truncilla
|
--+ +-- Lampsilis
| +-----+
| | +-- Epioblasma_triquetra **
+--+
| +----- Ligumia
+--+
| +-- Obliquaria
+--+
+-- Ortmanniana
Smith et al., 2019 [Publ 3951], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 40 | 35 | 19 | -- | +----------------------- Obliquaria
|
| +-- Medionidus
| +--+
| | +-- Lemiox
--+ +--------------+
| | | +-- Cyprogenia
| | +--+
| | +-- Ptychobranchus
| |
+--+ +-------- Potamilus
| +--------+
| | | +----- Atlanticoncha
| | +--+
| | | +-- Ellipsaria
| | +--+
+--+ +-- Truncilla
|
| +-- Hamiota
| +-----------+
| | +-- Villosa
| |
+--+ +----------- Ligumia
| |
| | +----- Venustaconcha
+--+ +--+
| | | +-- Obovaria
| | +--+
+--+ +-- Epioblasma_rangiana **
|
| +----- Ortmanniana_pectorosa
+--+
| +-- Ortmanniana_ligamentina
+--+
+-- Lampsilis
Hewitt et al., 2019 [Publ 3962], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 9 | 6 | 1 | -- | +-------------------- Lampsilis_siliquoidea
|
| +----------------- Lampsilis_fasciola
--+ |
| | +-- Epioblasma_rangiana **
| +--------------+
+--+ +-- Lampsilis_radiata
|
| +-------------- Lampsilis_ornata
| |
+--+ +-- Lampsilis_cardium__1
| +--------+
| | +-- Lampsilis_ovata__1
+--+
| +-------- Lampsilis_cardium__6
| |
+--+ +----- Lampsilis_cardium__5
| |
+--+ +-- Lampsilis_cardium__2
| |
| +-- Lampsilis_ovata__2
+--+
+-- Lampsilis_cardium__3
|
+-- Lampsilis_cardium__4
Zanatta & Harris, 2013 [Publ 3530], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 4 | 3 | 2 | -- | +----- Lampsilis
--+
| +-- Epioblasma_rangiana **
+--+
+-- Venustaconcha
Campbell & Lydeard, 2012 [Publ 3309], fig. 1-2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 72 | 14 | 10 | -- | +-------------- Potamilus_alatus
|
+-------------- Potamilus_capax
|
+-------------- Glebula
|
| +-- Toxolasma
+-----------+
--+ +-- Paetulunio
|
| +-- Dromus
+-----------+
| +-- Lemiox
|
| +-- Lampsilis_ovata
| +--------+
| | +-- Obliquaria
+--+
| +-------- Lampsilis_virescens
| |
+--+ +-- Epioblasma_rangiana **
| +--+
| | +-- Obovaria_arkansasensis
+--+
| +-- Obovaria_olivaria
+--+
+-- Venustaconcha
Hoeh et al., 2009 [Publ 2716], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 46 | 7 | 7 | -- | +----------------- Ptychobranchus
--+
| +-------------- Truncilla
+--+
| +----------- Ortmanniana
+--+
| +-------- Lampsilis
+--+
| +----- Cambarunio
+--+
| +-- Ligumia
+--+
+-- Epioblasma_triquetra **
Lane et al., 2016 [Publ 3735], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 4 | 4 | 2 | -- | +-- Paetulunio
|
--+-- Epioblasma_capsaeformis **
|
+-- Venustaconcha
Hoeh et al., 2009 [Publ 2716], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 45 | 7 | 7 | -- | +----------------- Ptychobranchus
--+
| +-------------- Truncilla
+--+
| +----------- Ortmanniana
+--+
| +-------- Lampsilis
+--+
| +----- Cambarunio
+--+
| +-- Ligumia
+--+
+-- Epioblasma_triquetra **
Pfeiffer et al., 2019 [Publ 3866], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 30 | 11 | 11 | -- | +-- Cyrtonaias
+--------------------+
| +-- Glebula
--+
| +-------------------- Potamilus
| |
+--+ +-- Truncilla
| +--------------+
| | +-- Ptychobranchus
+--+
| +-------------- Pachynaias
| |
+--+ +----------- Mesonaias
| |
+--+ +-- Lampsilis
| +-----+
| | +-- Ortmanniana
+--+
| +----- Cambarunio
+--+
| +-- Epioblasma_triquetra **
+--+
+-- Ligumia
Burlakova et al., 2019 [Publ 3899], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 32 | 25 | 17 | -- | +----------------- Toxolasma
|
| +-------------- Potamilus
| |
| | +-- Ellipsaria
| +-----------+
--+ | +-- Truncilla
| |
| | +----- Medionidus
| +--------+
| | | +-- Lemiox
| | +--+
+--+ +-- Dromus
|
| +----------- Cambarunio
| |
| +----------- Leaunio
| |
| +----------- Villosa
| |
| | +-- Sagittunio
| +--------+
+--+ +-- Lampsilis_ovata
|
| +----- Venustaconcha
+-----+
| | +-- Epioblasma_rangiana **
| +--+
| +-- Obovaria
|
| +-------- Lampsilis_teres
+--+
| +----- Lampsilis_siliquoidea
+--+
| +-- Lampsilis_hydiana
+--+
+-- Lampsilis_straminea
|
