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Page last updated
Thu 3 September 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

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Bivalvia | Unionoida | Unionoidea | Unionidae | Parreysiinae | Indochinellini | Indonaia

species Indonaia gratiosa (Philippi, 1843): synonymy
     45 specimens + 1 Genbank voucher

Unio gratiosus Philippi, 1843
‘Parreyss’ Philippi, 1843. Abbild. Beschr. Conch. 1: 20, pl. 1, fig. 5.

Nodularia (s.s.) gratiosaSimpson (1900), Simpson (1914), Preston (1915)
Iredale (1934)
Ensidens (Uniandra) gratiosusHaas (1969)
Contradens gratiosusGraf & Cummings (2007), Zieritz et al. (2018), Konopleva et al. (2019), Graf & Cummings (2021)
Uniandra gratiosusBogan (2010), IUCN (2012)
Radiatula gratiosusPfeiffer et al. (2021)
Indonaia (cylindrica-group) gratiosaBolotov et al. (2022), Bolotov et al. (2024)
Indonaia gratiosaBolotov et al. (2022)

Unio occatus Lea, 1860
Lea, 1860. Proc. Acad. Nat. Sci. 12: 307.
Lea, 1863. Jour. Acad. Nat. Sci. 5 [O. 10]: 398, pl. 50, fig. 304.
Type(s): LECTOTYPE USNM_84077, Bengal, India.

Nodularia (s.s.) occataSimpson (1900), Preston (1912), Simpson (1914), Preston (1915)
Indonaia occataAnnandale & Prashad (1921), Haas (1969), Graf & Cummings (2021)
Parreysia (Radiatula) occataSubba Rao (1989), Graf & Cummings (2007), Ramakrishna & Dey (2007)
Radiatula (Indonaia) occataNesemann et al. (2003)
Radiatula occataNesemann et al. (2007), Zieritz et al. (2018)
Parreysia occataBogan (2010), IUCN (2010)
Indonaia (cylindrica-group) gratiosaBolotov et al. (2022)

Unio macilentus Benson, 1862
Benson, 1862. Ann. Mag. Nat. Hist. 10: 187.

Nodularia (s.s.) occataSimpson (1900), Simpson (1914), Preston (1915)
Indonaia occataHaas (1969)
Indonaia (cylindrica-group) gratiosaBolotov et al. (2022)

Unio siliguriensis Preston, 1908
Preston, 1908. Rec. Ind. Mus. 2: 47.
Preston, 1912. Rec. Ind. Mus. 7: 293.
Preston, 1915. Fauna Brit. India: 148.

Nodularia (Radiatula) lima var. siliguriensisPreston (1912)
Nodularia (sec. Radiatula) lima var. siliguriensisSimpson (1914), Preston (1915)
Radiatula limaHaas (1969)
Parreysia (Radiatula) limaSubba Rao (1989), Ramakrishna & Dey (2007)
Indonaia (cylindrica-group) gratiosaBolotov et al. (2022)

Unio rugosus Hanley & Theobald, 1876
Hanley & Theobald, 1876. Conch. Indica (8): 62, pl. 154, fig. 3.
(misidentified reference to Unio rugosus Barnes, 1823)

Nodularia (s.s.) occataSimpson (1900), Simpson (1914), Preston (1915)
Indonaia occataHaas (1969)

Taxonomic History (since 1900)

Nodularia (s.s.) gratiosaSimpson (1900), Simpson (1914), Preston (1915)
Nodularia (s.s.) occataSimpson (1900), Preston (1912), Simpson (1914), Preston (1915)
Nodularia (Radiatula) lima var. siliguriensisPreston (1912)
Nodularia (sec. Radiatula) lima var. siliguriensisSimpson (1914), Preston (1915)
Indonaia occataAnnandale & Prashad (1921), Haas (1969), Graf & Cummings (2021)
Iredale (1934)
Ensidens (Uniandra) gratiosusHaas (1969)
Radiatula limaHaas (1969)
Parreysia (Radiatula) occataSubba Rao (1989), Graf & Cummings (2007), Ramakrishna & Dey (2007)
Parreysia (Radiatula) limaSubba Rao (1989), Ramakrishna & Dey (2007)
Radiatula (Indonaia) occataNesemann et al. (2003)
Contradens gratiosusGraf & Cummings (2007), Zieritz et al. (2018), Konopleva et al. (2019), Graf & Cummings (2021)
Radiatula occataNesemann et al. (2007), Zieritz et al. (2018)
Uniandra gratiosusBogan (2010), IUCN (2012)
Parreysia occataBogan (2010), IUCN (2010)
Radiatula gratiosusPfeiffer et al. (2021)
Indonaia (cylindrica-group) gratiosaBolotov et al. (2022), Bolotov et al. (2024)
Indonaia gratiosaBolotov et al. (2022)

 
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