The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)
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Publication Campbell & Lydeard (2026): taxonomic opinions
publication summary Complete CitationCampbell, D.C. & C. Lydeard. 2026. Three decades of mussel mitochondria. Malacologia 68(1/2): 237-294.
Taxonomic OpinionsAcudonta baitiaoensis Wu, Liu & Zhang in Wu et al., 2025 Acuticosta chinensis (Lea, 1868) Acuticosta retiaria (Heude, 1883) Alasmidonta raveneliana (Lea, 1834) Alasmidonta uwharriensis Whelan, Perkins & Mays in Whelan et al., 2023 Alasmidonta varicosa (Lamarck, 1819) Alasmidonta (Bullella) arcula (Lea, 1838) = Alasmidonta arcula Alasmidonta (Decurambis) marginata Say, 1818 = Alasmidonta marginata Alasmidonta (s.s.) undulata (Say, 1817) = Alasmidonta undulata Alasminota etowaensis (Conrad, 1849) Alasminota holstonia (Lea, 1838) Alathyria jacksoni Iredale, 1934 Alathyria pertexta Iredale, 1934 Alathyria profuga (Gould, 1850) Amblema elliottii (Lea, 1857) Amblema neislerii (Lea, 1858) Amblema plicata (Say, 1817) Amuranodonta kijaensis Moskvicheva, 1973 Anemina arcaeformis (Heude, 1877) = Anemina harlandi Anodonta anatina (Linnaeus, 1758) Anodonta cygnea (Linnaeus, 1758) Anodonta exulcerata Porro, 1838 Anodonta seddoni Gürlek, Kebapçı & Lopes-Lima in Lopes-Lima et al., 2021 = Anodonta seddonae Anodontites elongata (Swainson, 1823) Anodontites guanarensis Marshall, 1927 Anodontites trapesialis (Lamarck, 1819) Anodontites trigona (Spix in Wagner, 1827) Anodontoides denigratus (Lea, 1852) Anodontoides ferussacianus (Lea, 1834) Arcidens confragosus (Say, 1829) Arcidens wheeleri (Ortmann & Walker, 1912) Aspatharia pfeifferiana (Bernardi, 1860) Assamnaia involuta (Hanley, 1856) Atlanticoncha ochracea (Say, 1817) Balwantia soleniformis (Benson, 1836) Beringiana beringiana (Middendorff, 1851) Beringiana fukuharai San, Hattori & Kondo in Lopes-Lima et al., 2020 Beringiana gosannensis San, Hattori & Kondo in Lopes-Lima et al., 2020 Beringiana japonica (Clessin, 1874) Bineurus anodontinus (Rochebrune, 1882) = Bineurus anodontinum Bineurus exilis (Morelet, 1866) Bineurus loeiensis Konopleva, Bolotov, Pfeiffer, Vikhrev, Kondakov, Gofarov, Tomilova, Tanmuangpak, & Tumpeesuwa, 2021 Bineurus mouhotii (Lea, 1863) Bineurus panhai Jeratthitikul & Sutcharit, 2023 Brachyanodon californiensis (Lea, 1852) = Anodonta californiensis Brachyanodon impurus (Say, 1829) = Brachyanodon impura Brachyanodon nuttallianus (Lea, 1838) = Anodonta nuttalliana Brachyanodon oregonensis (Lea, 1838) = Anodonta oregonensis Buldowskia flavotincta (Martens, 1905) Buldowskia iwakawai (Suzuki, 1939) Buldowskia kamiyai San, Hattori & Kondo in Lopes-Lima et al., 2020 Buldowskia shadini (Moskvicheva, 1973) Buldowskia suifunica (Lindholm, 1925) Callonaia duprei (Récluz, 1842) Cambarunio dactylus (Lea, 1840) Cambarunio hesperus Watters, 2018 Cambarunio iris (Lea, 1829) Cambarunio jonesii (Lea, 1859) Cambarunio nebulosus (Conrad, 1834) Cambarunio simus (Lea, 1838) Cambarunio taeniatus (Conrad, 1834) Castalia ambigua Lamarck, 1819 Castalia stevensi (H.B. Baker, 1930) Castaliella quadrata (G.B. Sowerby II, 1869) Caudiculatus caudiculatus (Martens, 1867) Chambardia rubens (Lamarck, 1819) Chambardia wahlbergi (Krauss, 1848) Chamberlainia duclerci (Rochebrune, 1882) Chamberlainia hainesiana (Lea, 1856) Coelatura aegyptiaca (Cailliaud, 1827) Coelatura alluaudi (Dautzenberg, 1908) Coelatura bakeri (H. Adams, 1866) Coelatura briarti (Dautzenberg, 1901) Coelatura choziensis (Preston, 1910) Coelatura choziensis bangweolica Haas, 1936 = Coelatura choziensis Coelatura gabonensis (Küster, 1862) Coelatura hauttecoeuri (Bourguignat, 1883) Coelatura hauttecoeuri kyogae Mandahl-Barth, 1954 = Coelatura hauttecoeuri Coelatura horei (E.A. Smith, 1880) Coelatura hypsiprymna (Martens, 1897) Coelatura kunenensis (Mousson, 1887) Coelatura leopoldvillensis (Putzeys, 1898) Coelatura luapulaensis (Preston, 1913) Coelatura mesafricana Pilsbry & Bequaert, 1927 Coelatura mossambicensis (Martens, 1860) Coelatura ratidota (Charmes, 1885) Coelatura stanleyvillensis Pilsbry & Bequaert, 1927 Coelatura stuhlmanni (Martens, 1897) Cosmopseudodon resupinatus (Martens, 1902) Cosmopseudodon wenshanensis Wu & Liu in Wu et al., 2024 = Cosmopseudodon resupinatus Cristaria beirensis Liu & Zhang, 1982 = Cristaria plicata Cristaria bellua (Morelet, 1866) Cristaria clessini (Kobelt, 1879) Cristaria plicata (Leach, 1814) Cristaria radiata Simpson, 1900 Cristaria truncata Dang, Thai, & Pham, 1980 Ctenodesma bersinara Zieritz, Rahim, Prayogo & Lopes-Lima in Zieritz et al., 2024 Ctenodesma borneensis (Issel, 1874) Ctenodesma mawonae Zieritz, Rahim, Lopes-Lima & Pfeiffer in Zieritz et al., 2024 Cucumerunio novaehollandiae (Gray, 1834) Cumberlandia monodonta (Say, 1829) Cuneopsis celtiformis (Heude, 1874) Cuneopsis celtiosimilis Hou & Wu in Hou et al., 2025 Cuneopsis demangei Haas, 1929 Cuneopsis heudei (Heude, 1874) Cuneopsis kiangsiensis Tchang & Li, 1965 Cuneopsis rufescens (Heude, 1874) Cuneopsis szechenyii (Neumayr, 1899) Cyclonaias tuberculata (Rafinesque, 1820) Cyprogenia aberti (Conrad, 1850) Cyprogenia stegaria (Rafinesque, 1820) Cyrtonaias tampicoensis (Lea, 1838) = Mesonaias tampicoensis Diaurora aurorea (Heude, 1883) Diaurora laeve Chen, Dai, Huang & Wu in Chen et al., 2023 Diplodon chilensis (Gray, 1828) Diplodon gratus (Lea, 1860) Diplodon suavidicus (Lea, 1856) Discomya radulosa (Drouët & Chaper, 1892) Dromus dromas (Lea, 1834) Echyridella aucklandica (Gray, 1843) Echyridella menziesii (Gray, 1843) Echyridella onekaka Fenwick & Marshall, 2006 Ellipsaria lineolata (Rafinesque, 1820) Elliptio angustata (Lea, 1831) Elliptio arca (Conrad, 1834) Elliptio arctata (Conrad, 1834) Elliptio complanata (Lightfoot, 1786) Elliptio congaraea (Lea, 1831) Elliptio crassidens (Lamarck, 1819) Elliptio dariensis (Lea, 1842) Elliptio downiei (Lea, 1858) Elliptio fisheriana (Lea, 1838) Elliptio folliculata (Lea, 1838) Elliptio fraterna (Lea, 1852) Elliptio fumata (Lea, 1857) Elliptio hopetonensis (Lea, 1838) Elliptio icterina (Conrad, 1834) Elliptio jayensis (Lea, 1838) Elliptio mcmichaeli Clench & Turner, 1956 Elliptio mediocris (Lea, 1863) = Elliptio complanata Elliptio nasutidus (Simpson, 1900) [sic] = Elliptio producta Elliptio nigella (Lea, 1852) Elliptio producta (Conrad, 1836) Elliptio pullata (Lea, 1857) Elliptio purpurella (Lea, 1857) Elliptio roanokensis (Lea, 1834) Elliptio shepardiana (Lea, 1834) Elliptio spinosa (Lea, 1836) Elliptoideus sloatianus (Lea, 1840) Ensidens dugasti (Morlet, 1892) Ensidens ingallsianus (Lea, 1852) Ensidens jaculus (Rochebrune, 1882) Ensidens sagittarius (Lea, 1856) Ensidens spiculus Pfeiffer, Graf, Cummings, & Page, 2021 Ensidens telus Pfeiffer, Graf, Cummings, & Page, 2021 Epioblasma brevidens (Lea, 1831) Epioblasma capsaeformis (Lea, 1834) Epioblasma obliquata (Rafinesque, 1820) Epioblasma rangiana (Lea, 1838) Epioblasma triquetra (Rafinesque, 1820) Epioblasma walkeri (Wilson & Clark, 1914) Etheria elliptica Lamarck, 1807 Eurynia dilatata (Rafinesque, 1820) Fossula fossiculifera (d’Orbigny, 1835) Fusconaia askewi (Marsh, 1896) Fusconaia burkei (Walker in Ortmann & Walker, 1922) Fusconaia cerina (Conrad, 1838) Fusconaia chunii (Lea, 1862) Fusconaia cor (Conrad, 1834) Fusconaia cuneolus (Lea, 1840) Fusconaia escambia Clench & Turner, 1956 Fusconaia flava (Rafinesque, 1820) Fusconaia hebetata (Conrad, 1854) = Fusconaia flava Fusconaia iheringi (B.H. Wright, 1898) Fusconaia masoni (Conrad, 1834) Fusconaia mitchelli (Simpson in Dall, 1896) Fusconaia ozarkensis (Call, 1887) Fusconaia subrotunda (Lea, 1831) Gibbosula crassa (Wood, 1815) Gibbosula laosensis (Lea, 1863) Gibbosula rochechouartii (Heude, 1875) Gibbosula sinensis Chen, Dai, Chen & Wu in Z.-G. Chen et al., 2025 Glebula rotundata (Lamarck, 1819) Globunio mirificus Chen, Dai, Huang & Wu in Dai, Chen, Ouyang et al., 2025 Gonidea angulata (Lea, 1838) Grandidieria burtoni (Woodward, 1859) Guiunio rarus Chen, Li, Dai & Wu in Dai et al., 2025 Hamiota altilis (Conrad, 1834) Hamiota australis (Simpson, 1900) Hamiota perovalis (Conrad, 1834) Hamiota subangulata (Lea, 1840) Harmandia somboriensis Rochebrune, 1882 Hemistena lata (Rafinesque, 1820) Huizhou xietianxiangi H. Chen, Z.-G. Chen, H.-Q. Xiang, Y.-M. He & X.-P. Wu in Chen et al., 2025 Hyridella australis (Lamarck, 1819) Hyridella depressa (Lamarck, 1819) Hyridella drapeta (Iredale, 1934) Hyriopsis bialata Simpson, 1900 Hyriopsis delaportei (Crosse & Fischer, 1876) Hyriopsis desowitzi Brandt, 1974 Hyriopsis khoratensis Pfeiffer, Graf, Cummings, & Page, 2021 Hyriopsis kratiensis Pfeiffer, Graf, Cummings, & Page, 2021 Hyriopsis myersiana (Lea, 1856) Hyriopsis panhai Jeratthitikul, Paphatmethin, Zieritz, Lopes-Lima, & Ngor, 2021 Hyriopsis phuphaniensis Pfeiffer, Graf, Cummings, & Page, 2021 Hyriopsis sakhonensis Pfeiffer, Graf, Cummings, & Page, 2021 Indochinella pugio (Benson, 1862) Indochinella pugio daweiensis Bolotov et al., 2019 = Indochinella pugio Indochinella pugio paradoxa Bolotov et al., 2019 = Indochinella pugio Indochinella pugio pugio (Benson, 1862) = Indochinella pugio Indochinella pugio viridissima Bolotov et al., 2019 = Indochinella pugio Indonaia andersoniana (Nevill, 1877) Indonaia bonneaudii (Eydoux, 1838) Indonaia caerulea (Lea, 1831) Indonaia corbis (Hanley, 1856) Indonaia cylindrica Annandale & Prashad, 1919 Indonaia gratiosa (Philippi, 1843) Indonaia rugosa (Gmelin, 1791) Indonaia shurtleffiana (Lea, 1856) Indonaia subclathrata (Martens, 1899) Indopseudodon bogani Bolotov, Kondakov & Konopleva in Bolotov et al., 2017 Indopseudodon crebristriatus (Anthony, 1865) Indopseudodon indawgyiensis Bolotov, Konopleva, Kondakov & Vikhrev in Bolotov et al., 2023 Indopseudodon kayinensis Bolotov, Konopleva, Vikhrev, Gofarov, Lopes-Lima, Bogan, Lunn, Chan, Win, Aksenova, Tomilova, Tanmuangpak, Tumpeesuwan, & Kondakov, 2020 Indopseudodon salwenianus (Gould, 1844) Inversidens brandtii (Kobelt, 1879) Inversidens rentianensis Wu & Wu in Wu et al., 2021 Inversiunio jokohamensis (Ihering, 1893) Inversiunio reinianus (Kobelt, 1879) Inversiunio verrucosus Kondo, Hyun & Seung-Ho, 2007 Inversiunio yanagawensis (Kondo, 1982) Isannaia fortunata Jeratthitikul, Sutcharit & Prasankok, 2024 Isannaia occultata Jeratthitikul, Sutcharit & Prasankok, 2024 Khairuloconcha lunbawangorum (Issel, 1874) [sic] Khairuloconcha sahanae Zieritz, Jainih & Lopes-Lima in Zieritz et al., 2021 Koreosolenaia sitgyensis Lee, Kim, Lopes-Lima & Bogan in Lopes-Lima et al., 2020 Lamellidens chindwinensis Bolotov, Konopleva, Chan, Lunn, Win, Gofarov, Kondakov, Tomilova, & Vikhrev, 2022 Lamellidens corrianus (Lea, 1834) Lamellidens ferrugineus (Annandale, 1918) Lamellidens generosus (Gould, 1847) Lamellidens jenkinsianus (Benson, 1862) Lamellidens lamellatus (Lea, 1838) Lamellidens mainwaringi Preston, 1912 Lamellidens marginalis (Lamarck, 1819) Lamellidens nongyangensis Preston, 1912 Lamellidens savadiensis (Nevill, 1877) Lamproscapha ensiformis (Spix in Wagner, 1827) Lamprotula caveata (Heude, 1877) Lamprotula cornuumlunae (Heude, 1883) Lamprotula leaii (Gray in Griffith & Pidgeon, 1833) Lamprotula seomjinensis Kondo, Hyun, & Seung-Ho, 2007 Lampsilis bergmanni Inoue & Randklev in Inoue et al., 2020 Lampsilis bracteata (Gould, 1855) Lampsilis cardium Rafinesque, 1820 Lampsilis cariosa (Say, 1817) Lampsilis dolabraeformis (Lea, 1838) Lampsilis fasciola Rafinesque, 1820 Lampsilis hydiana (Lea, 1838) Lampsilis ornata (Conrad, 1835) Lampsilis ovata (Say, 1817) Lampsilis pectorosa (Conrad, 1834) = Ortmanniana pectorosa Lampsilis powellii (Lea, 1852) Lampsilis radiata (Gmelin, 1791) Lampsilis satura (Lea, 1852) Lampsilis sietmani Keogh & Simons, 2019 Lampsilis siliquoidea (Barnes, 1823) Lampsilis splendida (Lea, 1838) Lampsilis straminea (Conrad, 1834) Lampsilis straminea claibornensis (Lea, 1838) = Lampsilis straminea Lampsilis straminea contrarius (Conrad, 1849) = Lampsilis straminea Lampsilis virescens (Lea, 1858) Lanceolaria acrorrhyncha (Martens, 1894) Lanceolaria eucylindrica Lin, 1962 Lanceolaria gladiola (Heude, 1877) Lanceolaria grayii (Griffith & Pidgeon, 1833) Lanceolaria kihirai Kondo & Hattori, 2019 Lanceolaria lanceolata (Lea, 1856) Lanceolaria oxyrhyncha (Martens, 1861) Lanceolaria triformis (Heude, 1877) Lannanaia kokensis Jeratthitikul, Sutcharit & Prasankok, 2024 Lasmigona (Pterosyna) complanata (Barnes, 1823) = Pterosyna complanata Lasmigona (s.s.) costata (Rafinesque, 1820) = Lasmigona costata Leaunio lienosus (Conrad, 1834) Leaunio lienosus aquilonius Watters, 2018 = Leaunio lienosus Leaunio lienosus caliginosus (Lea, 1845) = Leaunio lienosus Leaunio lienosus lienosus (Conrad, 1834) = Leaunio lienosus Leaunio lienosus unicostatus (B.H. Wright, 1899) = Leaunio lienosus Leaunio ortmanni (Walker, 1925) Leaunio pataecus Watters, 2018 Leaunio umbrans (Lea, 1857) Leaunio vanuxemensis (Lea, 1838) Leguminaia anatolica Gürlek, Kebapçı & Lopes-Lima in Lopes-Lima et al., 2021 Leguminaia saulcyi (Bourguignat, 1852) Leguminaia wheatleyi (Lea, 1862) Lemiox rimosus (Rafinesque, 1831) Lens comptus (Deshayes in Deshayes & Jullien, 1876) Lens contradens (Lea, 1838) Lens eximius (Lea, 1856) Lens inornatus (Lea, 1856) Lens lugens (Drouët & Chaper, 1892) Lens maenamensis Pfeiffer, Graf, Cummings, & Page, 2021 Lens micropterus (Morelet, 1866) Lens misellus (Morelet, 1865) Lens novoselovi Konopleva, Bolotov, Spitsyn, Kondakov, Gofarov, & Vikhrev, 2019 Lens pallegoixi (G.B. Sowerby II, 1867) Lens rolfbrandti Jeratthitikul & Panha in Jeratthitikul et al., 2019 Leoparreysia burmana (Blanford, 1869) Leoparreysia canefrii Vikhrev, Bolotov & Kondakov in Bolotov et al., 2017 Leoparreysia olivacea (Prashad, 1930) Leoparreysia tavoyensis (Gould, 1843) Leoparreysia whitteni Bolotov, Konopleva, Vikrev, Lopes-Lima, Bogan, Lunn, Chan, Win, Aksenova, Gofarov, Tomilova, & Kondakov, 2019 Lepidodesma aligera (Heude, 1877) Lepidodesma languilati (Heude, 1874) Ligodonta obscura Perkins, DeVilbiss & Whelan in Perkins et al., 2025 Ligumia recta (Lamarck, 1819) Lortiella rugata (G.B. Sowerby II, 1868) Margaritifera dahurica (Middendorff, 1850) Margaritifera falcata (Gould, 1850) Margaritifera hembeli (Conrad, 1838) Margaritifera laevis (Haas, 1910) Margaritifera margaritifera (Linnaeus, 1758) Margaritifera marrianae R.I. Johnson, 1983 Margaritifera middendorffi (Rosén, 1926) Medionidus acutissimus (Lea, 1831) Medionidus conradicus (Lea, 1834) Medionidus penicillatus (Lea, 1857) Megalonaias nervosa (Rafinesque, 1820) Megalonaias nickliniana (Lea, 1834) Microcondylaea bonellii (Férussac, 1827) Microdontia anodontaeformis (Tapparone Canefri, 1883) Middendorffinaia mongolica (Middendorff, 1851) Mimo guoliangi Xiang, Lu, He, & Li, 2025 Moncetia anceyi Bourguignat, 1886 Monocondylaea minuana (d’Orbigny, 1835) Monocondylaea parchappii (d’Orbigny, 1835) Mulleria rivoli (Deshayes, 1827) Mutela dubia (Gmelin, 1791) Mutela hargeri E.A. Smith, 1908 Mutela rostrata (Rang, 1835) Mycetopoda siliquosa (Spix in Wagner, 1827) Mytella strigata (Hanley, 1843) = Mytella charruana Namkongnaia inkhavilayi Jeratthitikul, Sutcharit, Ngor, & Prasankok, 2021 Namkongnaia lemeslei (Morelet, 1875) Neotrigonia lamarckii (J.E. Gray, 1838) Neotrigonia margaritacea (Lamarck, 1804) Nephronaias tempisquensis Pilsbry, 1920 Nitia acuminata (H. Adams, 1866) Nitia monceti (Bourguignat, 1883) Nitia teretiuscula (Philippi, 1847) Nodularia breviconcha Lee, Kim, Bogan & Kondo in Lopes-Lima et al., 2020 Nodularia dorri (Wattebled, 1886) Nodularia douglasiae (Griffith & Pidgeon, 1833) Nodularia dualobtusa Wu & Liu in Wu et al., 2024 Nodularia fusiformans Wu & Liu in Wu et al., 2024 Nodularia guiensis Hou & Wu in Hou, Liu et al., 2025 Nodularia hanensis Wu & Liu in Wu et al., 2024 Nodularia huana Wu & Liu in Wu et al., 2024 Nodularia micheloti (Morlet, 1886) Nodularia nipponensis (Martens, 1877) Nodularia nuxpersicae (Dunker, 1848) Nyassunio nyassaensis (Lea, 1864) Nyeinchanconcha nyeinchani Bolotov, Konopleva, Vikhrev, Gofarov, Lopes-Lima, Bogan, Lunn, Chan, Win, Aksenova, Tomilova, Tanmuangpak, Tumpeesuwan & Kondakov, 2020 Obliquaria reflexa Rafinesque, 1820 Obovalis omiensis (Heimburg, 1884) Obovaria arkansasensis (Lea, 1862) Obovaria choctawensis (Athearn, 1964) Obovaria unicolor (Lea, 1845) Obovaria (Pseudoon) olivaria (Rafinesque, 1820) = Obovaria olivaria Obovaria (Rotundaria) subrotunda (Rafinesque, 1820) = Obovaria subrotunda Obovaria (s.s.) retusa (Lamarck, 1819) = Obovaria retusa Ortmanniana abrupta (Say, 1831) = Lampsilis abrupta Ortmanniana higginsii (Lea, 1857) = Lampsilis higginsii Ortmanniana ligamentina (Lamarck, 1819) Pachynaias spheniopsis (Morelet, 1849) Paetulunio fabalis (Lea, 1831) Parreysia corrugata (Müller, 1774) Parreysia keralaensis Bolotov, Pasupuleti & Subba Rao in Bolotov et al., 2022 Parreysia marcens (Benson, 1862) Parreysia nagpoorensis (Lea, 1860) Parreysia plagiosoma (Benson, 1862) Parreysia rakhinensis Bolotov, Konopleva, Vikhrev, Gofarov, Lopes-Lima, Bogan, Lunn, Chan, Win, Aksenova, Tomilova, Tanmuangpak, Tumpeesuwan, & Kondakov, 2020 Parvasolenaia rivularis (Heude, 1877) Parvaspina collina (Conrad, 1837) Parvaspina lanceolata (Lea, 1828) Parvaspina steinstansana (Johnson & Clarke, 1983) Physunio massini (Morelet, 1864) Physunio modelli Brandt, 1974 Physunio pellucidus Pfeiffer, Graf, Cummings, & Page, 2021 Physunio superbus (Lea, 1843) Pilsbryoconcha acuta Jeratthitikul & Prasankok in Jeratthitikul et al., 2022 Pilsbryoconcha carinifera (Conrad, 1837) Pilsbryoconcha compressa (Martens, 1860) = Pilsbryoconcha exilis Pilsbryoconcha exilis (Lea, 1838) Pilsbryoconcha hoikaab Jeratthitikul & Prasankok in Jeratthitikul et al., 2022 Pilsbryoconcha kittitati Jeratthitikul & Prasankok in Jeratthitikul et al., 2022 Pilsbryoconcha linguaeformis (Morelet, 1875) Pilsbryoconcha mekongiana Jeratthitikul & Prasankok in Jeratthitikul et al., 2022 Pilsbryoconcha schomburgki (Martens, 1860) Platynaias compressa (Lea, 1829) Platynaias decorata (Lea, 1852) Platynaias subviridis (Conrad, 1835) Plectomerus dombeyanus (Valenciennes in Humboldt & Bonpland, 1827) Plethobasus cooperianus (Lea, 1834) Plethobasus cyphyus (Rafinesque, 1820) Pletholophus guangzhouensis Dai, Chen, Huang & Wu in Dai et al., 2024 Pletholophus honglinhensis Bogan, Do, Froufe, & Lopes-Lima, 2023 Pletholophus reinianus (Martens, 1875) Pletholophus tenuis (Griffith & Pidgeon, 1833) Pleurobema athearni Gangloff, Williams, & Feminella, 2006 Pleurobema beadleianum (Lea, 1861) Pleurobema clava (Lamarck, 1819) Pleurobema cordatum (Rafinesque, 1820) Pleurobema decisum (Lea, 1831) Pleurobema georgianum (Lea, 1841) Pleurobema hanleyianum (Lea, 1852) Pleurobema oviforme (Conrad, 1834) Pleurobema perovatum (Conrad, 1834) Pleurobema plenum (Lea, 1840) Pleurobema pyriforme (Lea, 1857) Pleurobema riddellii (Lea, 1862) Pleurobema rubellum (Conrad, 1834) Pleurobema sintoxia (Rafinesque, 1820) Pleurobema stabile (Lea, 1861) Pleurobema strodeanum (B.H. Wright, 1898) Pleurobema taitianum (Lea, 1834) Pleuronaia barnesiana (Lea, 1838) Pleuronaia dolabelloides (Lea, 1840) Pleuronaia estabrookianus Lea, 1845 = Pleuronaia estabrookiana Pleuronaia gibber (Lea, 1838) Popenaias berezai Inoue, Cummings, Tiemann, Miller, Johnson, Smith, & Randklev, 2020 Popenaias popeii (Lea, 1857) Popenaias semirasa (Pilsbry, 1910) Postolata guangxiensis Dai, Huang, Guo & Wu in Dai et al., 2023 Postolata longjiangensis Liu & Wu in Liu et al., 2024 Potamilus alatus (Say, 1817) Potamilus amphichaenus (Frierson, 1898) Potamilus capax (Green, 1832) Potamilus fragilis (Rafinesque, 1820) Potamilus inflatus (Lea, 1831) Potamilus leptodon (Rafinesque, 1820) Potamilus metnecktayi R.I. Johnson, 1998 Potamilus ohiensis (Rafinesque, 1820) Potamilus purpuratus (Lamarck, 1819) Potamilus streckersoni Smith, Johnson, Inoue, Doyle, & Randklev, 2019 Potomida acarnanica (Kobelt, 1879) Potomida littoralis (Cuvier, 1798) Potomida semirugata (Lamarck, 1819) Prisodon obliquus Schumacher, 1817 Prisodon syrmatophorus (Gmelin, 1791) Prisodontopsis aviculaeformis F.R. Woodward, 1991 Prisodontopsis mweruensis (E.A. Smith, 1908) Prolasmidonta heterodon (Lea, 1829) Pronodularia japanensis (Lea, 1859) Pseudanodonta complanata (Rossmässler, 1835) Pseudocuneopsis capitata (Heude, 1874) Pseudocuneopsis liuovatus (He & Zhuang, 2013)
+ Pseudocuneopsis heqing Wang et al., 2025 Pseudocuneopsis perflora Chen, Xiang, He, Huang & Wu in H. Chen et al., 2025 Pseudocuneopsis sichuanensis Huang, Dai, Chen & Wu in Wu et al., 2022 Pseudocuneopsis wuana Liu & Wu in Liu et al., 2023 Pseudocuneopsis yangshuoensis Wu & Liu in Wu et al., 2023 Pseudocuneopsis yemaoi Dai, Chen, Huang & Wu in Dai et al., 2024 Pseudodon aeneolus Drouët & Chaper, 1892 = Cosmopseudodon aeneolus Pseudodon cambodjensis (Fischer, 1891) Pseudodon inoscularis (Gould, 1844) Pseudodon lenyanensis Bolotov, Konopleva, Vikhrev, Gofarov, Lopes-Lima, Bogan, Lunn, Chan, Win, Aksenova, Tomilova, Tanmuangpak, Tumpeesuwan, & Kondakov, 2020 Pseudodon mekongi Bolotov, Konopleva, Vikhrev, Gofarov, Lopes-Lima, Bogan, Lunn, Chan, Win, Aksenova, Tomilova, Tanmuangpak, Tumpeesuwan, & Kondakov, 2020 Pseudodon septuaginta Jeratthitikul, Phuhong et al., 2025 Pseudodon vagulus (Petit, 1865) Pseudodon vondembuschianus (Lea, 1840) Pseudodon vondembuschianus laosicus (Bolotov, Konopleva, Vikhrev, Gofarov, Lopes-Lima, Bogan, Lunn, Chan, Win, Aksenova, Tomilova, Tanmuangpak, Tumpeesuwan & Kondakov, 2020) = Pseudodon vondembuschianus Pseudodon vondembuschianus tapienicus Konopleva, Lheknim, Sriwoon, Kondakov, Vikhrev & Bolotov in Konopleva et al., 2023 = Pseudodon vondembuschianus Pseudodon vondembuschianus thasaenicus Konopleva, Lheknim, Sriwoon, Kondakov, Vikhrev & Bolotov in Konopleva et al., 2023 = Pseudodon vondembuschianus Pseudodontoideus connasaugaensis (Lea, 1858) Pseudodontoideus subvexus (Conrad, 1834) Pseudomulleria dalyi (E.A. Smith, 1898) Pseudopostolata angula Dai, Chen, Huang & Wu in Dai et al., 2024 Pseudoscabies scobina (Hanley, 1856) Pseudospatha tanganyicensis (E.A. Smith, 1880) Pseudunio auricularius (Spengler, 1793) Pseudunio homsensis (Lea, 1865) Pseudunio marocanus (Pallary, 1918) Psoronaias guatemalensis (Simpson, 1900) Psoronaias percompressa (Martens, 1887) = Psoronaias crocodilorum Psoronaias profunda (Simpson, 1914) = Psoronaias crocodilorum Psoronaias semigranosa (von dem Busch in Philippi, 1843) Psoronaias usumasintae (Crosse & Fischer, 1892) = Psoronaias crocodilorum Ptychobranchus fasciolaris (Rafinesque, 1820) Ptychobranchus greenii (Conrad, 1834) Ptychorhynchus pfisteri (Heude, 1874) Pustulosa infucata (Conrad, 1834) Pustulosa kieneriana (Lea, 1852) Pustulosa kleiniana (Lea, 1852) Pustulosa necki (Burlakova, Karatayev, Lopes-Lima & Bogan in Burlakova et al., 2018) Pustulosa nodulata (Rafinesque, 1820) Pustulosa petrina (Gould, 1855) Pustulosa pustulosa (Lea, 1831) Pustulosa succissa (Lea, 1852) Pyganodon cataracta (Say, 1817) Pyganodon fragilis (Lamarck, 1819) Pyganodon grandis (Say, 1829) Pyganodon lacustris (Lea, 1857) Quadrula fragosa (Conrad, 1836) Quadrula nobilis (Conrad, 1854) Quadrula quadrula (Rafinesque, 1820) Quadrula verrucosa (Rafinesque, 1820) Radiatula chaudhurii (Preston, 1912) Radiatula chindwinensis Bolotov, Konopleva, Vikrev, Lopes-Lima, Bogan, Lunn, Chan,Win, Aksenova, Gofarov, Tomilova, & Kondakov, 2019 Radiatula crispisulcata (Benson, 1862) Radiatula kachinensis Bolotov, Konopleva, Chan, Lunn, Win, Gofarov, Kondakov, Tomilova, & Vikhrev, 2022 Radiatula mouhoti Vikhrev, Bolotov & Konopleva in Bolotov et al., 2017 Radiatula mouhoti haungthayawensis Bolotov et al., 2019 = Radiatula mouhoti Radiatula myitkyinae (Prashad, 1930) Radiatula myitthanensis Bolotov, Konopleva, Vikrev, Lopes-Lima, Bogan, Lunn, Chan,Win, Aksenova, Gofarov, Tomilova, & Kondakov, 2019 Rectidens lauris Zieritz, Anwari, Diba & Pfeiffer in Zieritz et al., 2024 Rectidens sumatrensis (Dunker, 1852) Reginaia ebenus (Lea, 1831) Reginaia rotulata (B.H. Wright, 1899) Sagittunio nasutus (Say, 1817) Sagittunio subrostratus (Say, 1831) Sagittunio vaughanianus (Lea, 1838) Scabiellus songkramensis (Kongim & Panha in Kongim et al., 2015) Scabies anceps (Deshayes in Deshayes & Jullien, 1876) Scabies crispata (Gould, 1843) Scabies humilis (Lea, 1856) Scabies mandarinus (Morelet, 1864) Scabies nucleus (Lea, 1856) Scabies phaselus (Lea, 1856) Scabies pilata (Lea, 1866) Scabies scobinata (Lea, 1856) = Scabies scobinatus Schepmania nieuwenhuisi (Schepman, 1898) Schistodesmus lampreyanus (Baird & Adams, 1867) Schistodesmus luqiaoensis Hou & Wu in Hou et al., 2025 Schistodesmus spinosus (Simpson, 1900) Schistodesmus tongpenensis Hou & Wu in Hou et al., 2025 Schistodesmus xinyuensis Hou & Wu in Hou et al., 2025 Scriptolamprotula coreana (Martens, 1886) = Aculamprotula coreana Scriptolamprotula fibrosa (Heude, 1877) = Aculamprotula fibrosa Scriptolamprotula polysticta (Heude, 1877) = Aculamprotula polysticta Scriptolamprotula scripta (Heude, 1875) = Aculamprotula scripta Scriptolamprotula tientsinensis (Crosse & Debeaux, 1863) = Aculamprotula tientsinensis Scriptolamprotula tortuosa (Lea, 1865) = Aculamprotula tortuosa Simpsonaias ambigua (Say, 1825) Simpsonunio floridensis (Lea, 1852) = Lampsilis floridensis Simpsonunio teres (Rafinesque, 1820) = Lampsilis teres Sinanodonta angula (Tchang, Li, & Liu, 1965) Sinanodonta calipygos (Kobelt, 1879) Sinanodonta elliptica (Heude, 1878) Sinanodonta jourdyi (Morlet, 1886) Sinanodonta kyushuensis Sano & Kondo, 2025 = Sinanodonta lauta Sinanodonta lauta (Martens, 1877) Sinanodonta lucida (Heude, 1877) Sinanodonta pacifica (Heude, 1878) Sinanodonta plana Sano & Kondo, 2025 = Sinanodonta calipygos Sinanodonta schrenkii (Lea, 1870) Sinanodonta tumens (Haas, 1910) Sinanodonta woodiana (Lea, 1834) Sinohyriopsis cumingii (Lea, 1852) Sinohyriopsis goliath (Rolle, 1904) Sinohyriopsis schlegelii (Martens, 1861) Sinosolenaia carinata (Heude, 1877) Sinosolenaia iridinea (Heude, 1874) Sinosolenaia oleivora (Heude, 1877) Sinosolenaia recognita (Heude, 1877) Songkhlanaia moreleti (Crosse & Fischer, 1876) Songkhlanaia songkhramensis Jeratthitikul, Sutcharit, & Prasankok, 2025 Songkhlanaia tamodienica Konopleva, Lheknim, Sriwoon, Kondakov, Vikhrev & Bolotov in Konopleva et al., 2023 Sphenonaias explicata (Morelet, 1849) = Mesonaias explicata Strophitus howellsi Smith, Kiser, Johnson & Randklev, 2023 Strophitus pascagoulaensis Smith, Johnson, Pfeiffer, & Gangloff, 2018 Strophitus radiatus (Conrad, 1834) Strophitus undulatus (Say, 1817) Strophitus williamsi Smith, Johnson, Pfeiffer, & Gangloff, 2018 Sulcularia viridis (Rafinesque, 1820) Sundadontina brandti Bolotov, Konopleva, Vikhrev, Gofarov, Lopes-Lima, Bogan, Lunn, Chan, Win, Aksenova, Tomilova, Tanmuangpak, Tumpeesuwan & Kondakov, 2020 Sundadontina cumingii (Lea, 1851) Sundadontina harmandi (Crosse & Fischer, 1876) Sundadontina plugpomenica Konopleva, Lheknim, Sriwoon, Kondakov, Vikhrev & Bolotov in Konopleva et al., 2023 Sundadontina sulcata (Rochebrune, 1882) Sundadontina tanintharyiensis Bolotov, Konopleva, Vikhrev, Gofarov, Lopes-Lima, Bogan, Lunn, Chan, Win, Aksenova, Tomilova, Tanmuangpak, Tumpeesuwan, & Kondakov, 2020 Sundadontina taskaevi Bolotov, Konopleva, Vikhrev, Gofarov, Lopes-Lima, Bogan, Lunn, Chan, Win, Aksenova, Tomilova, Tanmuangpak, Tumpeesuwan, & Kondakov, 2020 Sundadontina tumida (Morelet, 1866) Tamsiella schroeteriana (Lea, 1852) Tamsiella tamsiana (Dunker, 1858) Tchangsinaia pisciculus (Heude, 1874) = Tchangsinaia piscicula Thaiconcha callifera (Martens, 1860) Thaiconcha munelliptica Konopleva, Bolotov, Pfeiffer, Vikhrev, Kondakov, Gofarov, Tomilova, Tanmuangpak, & Tumpeesuwan, 2021 Thaiconcha thaiensis Konopleva, Bolotov, Pfeiffer, Vikhrev, Kondakov, Gofarov, Tomilova, Tanmuangpak, & Tumpeesuwan, 2021 Theliderma cylindrica (Say, 1817) Theliderma johnsoni Bogan & Lopes-Lima in Lopes-Lima & Bogan, 2019 Theliderma metanevra (Rafinesque, 1820) Tongchuania mingiana Chen, Xiang, He, Huang & Wu in H. Chen et al., 2025 Toxolasma cylindrellus (Lea, 1868) Toxolasma germanum (Lea, 1861) = Toxolasma paulum Toxolasma glans (Lea, 1831) = Toxolasma lividum Toxolasma granulatum (Lea, 1861) = Toxolasma paulum Toxolasma lividum Rafinesque, 1831 Toxolasma parvum (Barnes, 1823) Toxolasma paulum (Lea, 1840) Toxolasma pullus (Conrad, 1838) Toxolasma texasiense (Lea, 1857) Trapezidens angustior (Hanley & Theobald, 1876) Trapezidens dolichorhynchus (Tapparone Canefri, 1889) Trapezidens exolescens (Gould, 1843) Trapezidens mogaungensis Bolotov, Konopleva, Chan, Lunn, Win, Gofarov, Kondakov, Tomilova, & Vikhrev, 2022 Trapezidens scutum (G.B. Sowerby II, 1868) Trapezidens yeti Konopleva, Bolotov, Kondakov, Kononov, Gofarov, Tomilova, Lunn, Chan, Win, & Vikhrev, 2020 Trapezoideus foliaceus (Gould, 1843) Trapezoideus lenya Bolotov, Konopleva, Vikhrev, Gofarov, Lopes-Lima, Bogan, Lunn, Chan, Win, Aksenova, Tomilova, Tanmuangpak, Tumpeesuwan, & Kondakov, 2020 Trapezoideus mitanensis Bolotov, Konopleva, Than Win, Kondakov & Vikhrev in Bolotov et al., 2022 Trapezoideus subviridus [sic] Pfeiffer, Graf, Cummings & Page, 2021 = Trapezoideus subviridis Trapezoideus thachiadensis Konopleva, Lheknim, Sriwoon, Kondakov, Vikhrev & Bolotov in Konopleva et al., 2023 Triplodon corrugatus (Lamarck, 1819) Truncilla cognata (Lea, 1860) Truncilla donaciformis (Lea, 1828) Truncilla macrodon (Lea, 1859) Truncilla truncata Rafinesque, 1820 Unio bruguierianus Bourguignat, 1853 Unio caffer Krauss, 1848 Unio carneus Küster, 1854 Unio crassus Philipsson in Retzius, 1788 Unio damascensis Lea, 1863 Unio delicatus Lea, 1863 Unio delphinus Spengler, 1793 Unio desectus Westerlund in Westerlund & Blanc, 1879 Unio durieui Deshayes, 1847 Unio elongatulus Pfeiffer, 1825 Unio eucirrus Bourguignat, 1857 Unio foucauldianus Pallary, 1936 Unio gibbus Spengler, 1793 Unio gontierii Bourguignat, 1856 Unio hueti Bourguignat, 1855 Unio ionicus Drouët, 1879 Unio mancus Lamarck, 1819 Unio mancus mancus Lamarck, 1819 = Unio mancus Unio mancus requienii Michaud, 1831 = Unio mancus Unio mancus turtonii Payraudeau, 1826 = Unio mancus Unio mardinensis Lea, 1865 Unio nanus Lamarck, 1819 Unio pictorum (Linnaeus, 1758) Unio ravoisieri Deshayes, 1848 Unio sesirmensis Kobelt, 1913 Unio terminalis Bourguignat, 1852 Unio tigridis Bourguignat, 1852 Unio tumidiformis Castro, 1885 Unio tumidus Philipsson in Retzius, 1788 Unio vicarius Westerlund in Westerlund & Blanc, 1879 Uniomerus carolinianus (Bosc, 1801) Uniomerus declivis (Say, 1831) Uniomerus tetralasmus (Say, 1831) Unionetta fabagina (Deshayes in Deshayes & Jullien, 1876) Utterbackia imbecillis (Say, 1829) Utterbackia peggyae (R.I. Johnson, 1965) Utterbackia peninsularis Bogan & Hoeh, 1995 Utterbackiana couperiana (Lea, 1840) Utterbackiana hartfieldorum (Williams, Bogan, & Garner, 2009) Utterbackiana heardi (Gordon & Hoeh, 1995) Utterbackiana implicata (Say, 1829) Utterbackiana suborbiculata (Say, 1831) Velesunio ambiguus (Philippi, 1847) Velesunio angasi (G.B. Sowerby II, 1867) Venustaconcha constricta (Conrad, 1838) Venustaconcha ellipsiformis (Conrad, 1836) Venustaconcha pleasii (Marsh, 1891) Venustaconcha trabalis (Conrad, 1834) Venustaconcha troostensis (Lea, 1834) Villosa delumbis (Conrad, 1834) Villosa vibex (Conrad, 1834) Villosa villosa (B.H. Wright, 1898) Westralunio carteri Iredale, 1934 Westralunio inbisi inbisi Klunzinger, Whisson, Zieritz, Benson, Stewart, & Kirkendale, 2022 = Westralunio inbisi Westralunio inbisi meridiemus Klunzinger, Whisson, Zieritz, Benson, Stewart, & Kirkendale, 2022 = Westralunio inbisi Yaukthwa avaensis Bolotov, Konopleva, Vikhrev, Gofarov, Lopes-Lima, Bogan, Lunn, Chan, Win, Aksenova, Tomilova, Tanmuangpak, Tumpeesuwan, & Kondakov, 2020 Yaukthwa baniensis Bolotov, Konopleva, Vikhrev, Gofarov, Lopes-Lima, Bogan, Lunn, Chan, Win, Aksenova, Tomilova, Tanmuangpak, Tumpeesuwan, & Kondakov, 2020 Yaukthwa dalliana (Frierson, 1913) Yaukthwa elongatula (Bolotov, Konopleva, Vikhrev, Lopes-Lima, Bogan, Lunn, Chan,Win, Aksenova, Gofarov, Tomilova, & Kondakov, 2019) Yaukthwa inlenensis Konopleva, Pfeiffer, Vikhrev, Kondakov, Gofarov, Aksenova, Lunn, Chan, & Bolotov, 2019 Yaukthwa nesemanni (Konopleva, Vikhrev & Bolotov in Bolotov et al., 2017) Yaukthwa paiensis Konopleva, Pfeiffer, Vikhrev, Kondakov, Gofarov, Aksenova, Lunn, Chan, & Bolotov, 2019 Yaukthwa panhai (Konopleva, Bolotov & Kondakov in Bolotov et al., 2017) Yaukthwa rectangularis (Tapparone Canefri, 1889) Yaukthwa zayleymanensis (Preston, 1912)
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