The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)
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Publication Wu et al. (2026): cladograms
publication summary | cladograms Complete CitationWu, R., X. Wang, F. Nan, L. Cheng & J. An. 2026. An integrative taxonomic reassessment of the Chinese endemic genus Acuticosta (Bivalvia: Unionidae), with the description of a new genus. Contributions to Zoology 95: 25-51. Available electronically.
Cladogram Figures(click the figure number to see the cladogram) Cladogram Figure 3
+-- Margaritifera_margaritifera
+--------------------+
| +-- Gibbosula_rochechouartii
|
--+ +-------------------- Amblema_plicata
| |
| | +-------- Gonidea_angulata
| | +--------+
+--+ | | +----- Inversidens_trisulcata
| | +--+
| | | +-- Inversidens_brandtii
| | +--+
+--+ +-- Inversidens_rentianensis
|
| +-------------- Unio_pictorum
| |
| | +-- Lanceolaria_gladiola
+--+ +--------+
| | +-- Lanceolaria_lanceolata
| |
| | +----- Globunio_mirificus
+--+ +--+
| | | +-- Pseudoacuticosta_retiaria
| | +--+
| | +-- Huizhou_xietianxiangi
+--+
| +-- Acuticosta_ovata
| +--+
| | +-- Acuticosta_chinensis
+--+
| +-- Brachyanodon_californiensis
+--+
+-- Brachyanodon_nuttalliana
Cladogram Figure 4
+-- Gibbosula_rochechouartii
+--------------------------------------------+
| +-- Margaritifera_margaritifera
|
| +----- Chamberlainia_somsakpanhai
| +-----------------------------+
| | | +-- Sinohyriopsis_cumingii
| | +--+
| | +-- Sinohyriopsis_schlegelii
| |
| | +-- Hyriopsis_bialata
| | +--+
| | | +-- Rectidens_sumatrensis
| +--------+ +--------------------------+
| | | | | +-- Lens_contradens
| | | | +--+
| | | | +-- Physunio_superbus
| | | |
| | | | +----- Potomida_semirugata__2
| | | | +-----+
| | | | | | +-- Potomida_acarnanica
| | +--+ | +--+
| | | +-----------------+ +-- Potomida_semirugata__1
| | | | |
| | | | | +-------- Pronodularia_japanensis
--+ | | | +--+
| | | | | +----- Lamprotula_leaii
| | | | +--+
| | | | | +-- Lamprotula_caveata
| | +--+ +--+
| | | +-- Lamprotula_cornuumlunae
| | |
| | | +-- Pilsbryoconcha_exilis
| | | +-----------------------+
| | | | +-- Pseudodon_mekongi
| | | |
| | | | +-- Inversidens_rentianensis
| | +--+ +--------------------+
| | | | +-- Inversidens_trisulcata
| | | |
| | | | +-------- Sinosolenaia_carinata
| | +--+ +-----------+
| | | | | +----- Sinosolenaia_recognita
| | | | +--+
| | | | | +-- Sinosolenaia_iridinea
| | +--+ +--+
| | | +-- Sinosolenaia_oleivora
| | |
| | | +----------------- Parvasolenaia_rivularis
+--+ +--+
| | +-------------- Pseudopostolata_angula
| +--+
| | +----------- Koreosolenaia_sitgyensis
| +--+
| | +-------- Ptychorhynchus_pfisteri
| +--+
| | +----- Cosmopseudodon_resupinatus
| +--+
| | +-- Postolata_guangxiensis
| +--+
| +-- Postolata_longjiangensis
|
| +-- Lepidodesma_aligera
| +--------------------------------------+
| | +-- Lepidodesma_languilati
| |
| | +-- Acuticosta_chinensis
| | +-----+
| | | +-- Acuticosta_ovata
| | +--------------------+
| | | | +----- Lanceolaria_gladiola
| | | +--+
| | | | +-- Lanceolaria_grayii
| | | +--+
| | | +-- Lanceolaria_lanceolata
| | |
| | | +----- Anodonta_anatina
| | | +-----------+
| | | | | +-- Anodonta_cygnea
| | | | +--+
| | +--------+ | +-- Anodonta_exulcerata
| | | | +--------+
| | | | | | +-------------- Pseudanodonta_complanata
| | | | | | |
| | | | | | | +----- Pyganodon_grandis
+--+ | | | +--+ +-----+
| | | | | | | +-- Utterbackia_imbecillis
| | | | | | +--+
| | | | +--+ +-- Utterbackia_peninsularis
| | | | |
| | | | | +-------- Pterosyna_complanata
| | | | +--+
| | +--+ | +----- Prolasmidonta_heterodon
| | | +--+
| | | | +-- Alasmidonta_varicosa
| | | +--+
| | | +-- Platynaias_compressa
| | |
| | | +----- Cristaria_plicata__2
| | | +-----------------+
| | | | | +-- Cristaria_plicata__1
| | | | +--+
| | | | +-- Cristaria_truncata
| | | |
| | +--+ +----- Amuranodonta_kijaensis
| | | +--------------+
| | | | | +-- Anemina_harlandi
| | | | +--+
+--+ | | +-- Buldowskia_shadini
| +--+
| | +-- Beringiana_fukuharai
| | +--------------+
| | | +-- Pletholophus_guangzhouensis
| +--+
| | +-------------- Acudonta_baitiaoensis
| | |
| +--+ +----------- Sinanodonta_lucida
| | |
| +--+ +-- Sinanodonta_schrenkii
| | +-----+
| | | +-- Sinanodonta_tumens
| +--+
| | +----- Sinanodonta_woodiana
| +--+
| | +-- Sinanodonta_lauta
| +--+
| +-- Sinanodonta_pacifica
|
| +----- Pseudoacuticosta_retiaria
| +-----------------------------+
| | | +-- Globunio_mirificus
| | +--+
| | +-- Huizhou_xietianxiangi
| |
| | +-------- Scriptolamprotula_tortuosa
| | |
+--+ +-----------------------+ +-- Scriptolamprotula_coreana
| | | +--+
| | | | +-- Scriptolamprotula_tientsinensis
| | +--+
| | | +-- Scriptolamprotula_polysticta
| | +--+
| | +-- Scriptolamprotula_scripta
+--+
| +-------- Unio_tumidus
| +--------------------+
| | | +----- Unio_elongatulus
| | +--+
| | | +-- Unio_carneus
| | +--+
| | +-- Unio_crassus
+--+
| +-------------------------- Middendorffinaia_mongolica
| |
| | +-------- Pseudocuneopsis_capitata
| | +--------------+
| | | | +----- Pseudocuneopsis_sichuanensis
+--+ | +--+
| | | +-- Pseudocuneopsis_wuana
| | +--+
| | +-- Pseudocuneopsis_yangshuoensis
| |
+--+ +-------- Nodularia_hanensis
| +-----------+
| | | +----- Nodularia_fusiformans
| | +--+
| | | +-- Nodularia_douglasiae
| | +--+
| | +-- Nodularia_guiensis
+--+
| +----------- Schistodesmus_spinosus
| +-----+
| | | +-------- Schistodesmus_tongpenensis
| | +--+
| | | +----- Schistodesmus_xinyuensis
| | +--+
+--+ | +-- Schistodesmus_lampreyanus
| +--+
| +-- Schistodesmus_luqiaoensis
|
| +-------------- Tchangsinaia_piscicula
| |
+--+ +----------- Cuneopsis_rufescens
| |
+--+ +-- Cuneopsis_heudei
| +-----+
| | +-- Cuneopsis_szechenyii
+--+
| +----- Cuneopsis_celtiformis
+--+
| +-- Cuneopsis_celtiosimilis
+--+
+-- Cuneopsis_kiangsiensis
|
