The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)
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Publication Wu et al. (2026): cladograms
publication summary | cladograms Complete CitationWu, R., X. Wang, X. Hu, F. Nan, X. Liu et al. 2026. Integrative taxonomic reappraisal and molecular phylogeny of freshwater mussels of the genus Lanceolaria (Bivalvia: Unionidae), with description of two new species. Zoologica Scripta: 1-24. Available electronically.
Cladogram Figures(click the figure number to see the cladogram) Cladogram Figure 3
+-- Schistodesmus_spinosus
+-----------------+
| +-- Schistodesmus_xinyuensis
|
--+ +----------------- Lanceolaria_oxyrhyncha
| |
| | +-- Lanceolaria_gladiola
+--+ +-----------+
| | +-- Lanceolaria_bilirata
| |
+--+ +----- Lanceolaria_acrorrhyncha
| +-----+
| | | +-- Lanceolaria_grayii
| | +--+
+--+ +-- Lanceolaria_danzhouensis
|
| +-------- Lanceolaria_kihirai
+--+
| +----- Lanceolaria_triformis
+--+
| +-- Lanceolaria_lanceolata
+--+
+-- Lanceolaria_helianthoides
Cladogram Figure 5
+-- Lamprotula_leaii
+--------------------------+
| +-- Potomida_littoralis
|
| +-- Schistodesmus_xinyuensis
| +-----+
--+ | +-- Schistodesmus_spinosus
| +-----------------+
| | | +----- Nodularia_fusiformans
| | +--+
| | | +-- Nodularia_douglasiae
| | +--+
+--+ +-- Nodularia_dualobtusa
|
| +-- Scriptolamprotula_scripta
| +--------------------+
| | +-- Scriptolamprotula_tientsinensis
| |
+--+ +-------------------- Lanceolaria_oxyrhyncha
| |
| | +-- Lanceolaria_bilirata
+--+ +--------------+
| | +-- Lanceolaria_gladiola
| |
+--+ +-- Lanceolaria_grayii
| +-----------+
| | +-- Lanceolaria_danzhouensis
+--+
| +----------- Lanceolaria_acrorrhyncha
+--+
| +-------- Lanceolaria_kihirai
+--+
| +----- Lanceolaria_triformis
+--+
| +-- Lanceolaria_helianthoides
+--+
+-- Lanceolaria_lanceolata
Cladogram Figure 6B
+-- Gibbosula_rochechouartii
+--------------------------------------+
| +-- Margaritifera_margaritifera
|
| +-- Lepidodesma_aligera
| +-----------------------------------+
| | +-- Lepidodesma_languilati
| |
| | +-------- Scriptolamprotula_tortuosa
| | +-----------------------+
| | | | +----- Scriptolamprotula_scripta
| | | +--+
| | | | +-- Scriptolamprotula_coreana
| | | +--+
--+ | | +-- Scriptolamprotula_tientsinensis
| | |
| | +--+ +-------- Unio_tumidus
| | | | +--------------------+
| | | | | | +----- Unio_elongatulus
| | | | | +--+
| | | | | | +-- Unio_carneus
| | | | | +--+
| | | | | +-- Unio_nanus
| | | +--+
| | | | +-------------------------- Middendorffinaia_mongolica
| | | | |
| | | | | +-------- Pseudocuneopsis_capitata
+--+ | | | +--------------+
| | | | | | +----- Pseudocuneopsis_sichuanensis
| | +--+ | +--+
| | | | | +-- Pseudocuneopsis_wuana
| | | | +--+
| | | | +-- Pseudocuneopsis_yangshuoensis
| | | |
| | +--+ +-------- Nodularia_hanensis
| | | +-----------+
| | | | | +----- Nodularia_fusiformans
| | | | +--+
| | | | | +-- Nodularia_douglasiae
| | | | +--+
| | | | +-- Nodularia_guiensis
| | +--+
| | | +----------- Schistodesmus_spinosus
| | | +-----+
| | | | | +-------- Schistodesmus_tongpenensis
| | | | +--+
| | | | | +----- Schistodesmus_xinyuensis
| | | | +--+
| | +--+ | +-- Schistodesmus_lampreyanus
+--+ | +--+
| | +-- Schistodesmus_luqiaoensis
| |
| | +-------------- Tchangsinaia_piscicula
| | |
| +--+ +----------- Cuneopsis_rufescens
| | |
| +--+ +-- Cuneopsis_heudei
| | +-----+
| | | +-- Cuneopsis_szechenyii
| +--+
| | +----- Cuneopsis_celtiformis
| +--+
| | +-- Cuneopsis_celtiosimilis
| +--+
| +-- Cuneopsis_kiangsiensis
|
| +-- Lanceolaria_bilirata
| +--------+
| | +-- Lanceolaria_gladiola
| |
| +-----------------+ +-- Lanceolaria_grayii
| | | +-----+
| | | | +-- Lanceolaria_danzhouensis
| | +--+
| | | +----- Lanceolaria_triformis
| | +--+
| | | +-- Lanceolaria_helianthoides
| | +--+
| | +-- Lanceolaria_lanceolata
| |
| | +----- Anodonta_anatina
| | +-----------+
| | | | +-- Anodonta_cygnea
+-----+ | +--+
| | +-- Anodonta_exulcerata
| +--------+
| | | +-------------- Pseudanodonta_complanata
| | | |
| | | | +----- Pyganodon_grandis
| | +--+ +-----+
| | | | | +-- Utterbackia_imbecillis
| | | | +--+
| | +--+ +-- Utterbackia_peninsularis
| | |
| | | +-------- Pterosyna_complanata
| | +--+
+--+ | +----- Prolasmidonta_heterodon
| +--+
| | +-- Alasmidonta_varicosa
| +--+
| +-- Platynaias_compressa
|
| +----- Cristaria_liboi
| +-----------------+
| | | +-- Cristaria_plicata
| | +--+
| | +-- Cristaria_truncata
| |
+--+ +----- Amuranodonta_kijaensis
| +--------------+
| | | +-- Anemina_harlandi
| | +--+
| | +-- Buldowskia_shadini
+--+
| +-- Beringiana_fukuharai
| +--------------+
| | +-- Pletholophus_guangzhouensis
+--+
| +-------------- Acudonta_baitiaoensis
| |
+--+ +----------- Sinanodonta_lucida
| |
+--+ +-- Sinanodonta_schrenkii
| +-----+
| | +-- Sinanodonta_tumens
+--+
| +----- Sinanodonta_pacifica__2
+--+
| +-- Sinanodonta_lauta
+--+
+-- Sinanodonta_pacifica__1
|
