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Page last updated
Thu 3 September 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

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Bivalvia | Unionida | Unionoidea | Unionidae | Parreysiinae | Indochinellini

genus Harmandia Rochebrune, 1882: cladomics
     1 Recent valid species (1 total species)

Cladomics Summary

number of species = 1 (1 total)
analyzed in 15 cladograms in 9 publications
maximum number of terminals in an analysis = 1 (mean = 1, median = 1)


Bolotov et al., 2024 [Publ 4369], fig. 2

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
110161287--
                 +-- Pseudoscabies         
  +--------------+                         
  |              +-- Indonaia              
--+                                        
  |  +-------------- Indochinella          
  +--+                                     
     |  +----------- Radiatula             
     +--+                                  
        |  +-------- Scabiellus            
        +--+                               
           |  +----- Harmandia_somboriensis **
           +--+                            
              |  +-- Unionetta             
              +--+                         
                 +-- Scabies               

Jeratthitikul et al., 2019 [Publ 3922], fig. 1

iTtTbiriSoTroTroSiD
110122176--
  +----------------- Indonaia              
--+                                        
  |  +-------------- Indochinella          
  +--+                                     
     |  +----------- Radiatula             
     +--+                                  
        |  +-------- Scabiellus            
        +--+                               
           |  +----- Harmandia_somboriensis **
           +--+                            
              |  +-- Unionetta             
              +--+                         
                 +-- Scabies               

Bolotov et al., 2024 [Publ 4369], fig. S1

iTtTbiriSoTroTroSiD
110162287--
                 +-- Pseudoscabies         
  +--------------+                         
  |              +-- Indonaia              
--+                                        
  |  +-------------- Indochinella          
  +--+                                     
     |  +----------- Radiatula             
     +--+                                  
        |  +-------- Scabiellus            
        +--+                               
           |  +----- Harmandia_somboriensis **
           +--+                            
              |  +-- Unionetta             
              +--+                         
                 +-- Scabies               

Bolotov et al., 2022 [Publ 4239], fig. 2

iTtTbiriSoTroTroSiD
110157256--
  +----------------- Indonaia              
--+                                        
  |  +-------------- Indochinella          
  +--+                                     
     |  +----------- Radiatula             
     +--+                                  
        |  +-------- Scabiellus            
        +--+                               
           |  +----- Harmandia_somboriensis **
           +--+                            
              |  +-- Unionetta             
              +--+                         
                 +-- Scabies               

Bolotov et al., 2020 [Publ 4100], fig. S1

iTtTbiriSoTroTroSiD
1101110196--
  +-------------------- Indonaia               
  |                                            
--+  +----------------- Indochinella           
  |  |                                         
  |  |           +----- Radiatula_mouhoti      
  +--+  +--------+                             
     |  |        |  +-- Radiatula_myitkyinae   
     |  |        +--+                          
     +--+           +-- Radiatula_chaudhurii   
        |                                      
        |  +----------- Radiatula_crispisulcata
        +--+                                   
           |  +-------- Scabiellus             
           +--+                                
              |  +----- Harmandia_somboriensis ** 
              +--+                             
                 |  +-- Unionetta              
                 +--+                          
                    +-- Scabies                

Pfeiffer et al., 2018 [Publ 3827], fig. 2

iTtTbiriSoTroTroSiD
110127166--
  +-------------------- Indonaia              
--+                                           
  |  +----------------- Indochinella          
  +--+                                        
     |  +-------------- Radiatula             
     +--+                                     
        |  +----------- Scabiellus            
        +--+                                  
           |  +-------- Unionetta_fabagina__2 
           +--+                               
              |  +----- Harmandia_somboriensis **
              +--+                            
                 |  +-- Unionetta_fabagina__1 
                 +--+                         
                    +-- Scabies               

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

iTtTbiriSoTroTroSiD
1101631276--
  +----- Harmandia_somboriensis **
--+                            
  |  +-- Unionetta             
  +--+                         
     +-- Scabies               

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iTtTbiriSoTroTroSiD
11015633--
  +-------- Indonaia              
--+                               
  |  +----- Indochinella          
  +--+                            
     |  +-- Scabies               
     +--+                         
        +-- Harmandia_somboriensis **

Bolotov et al., 2022 [Publ 4252], fig. 2

iTtTbiriSoTroTroSiD
110160266--
  +----------------- Indonaia              
--+                                        
  |  +-------------- Indochinella          
  +--+                                     
     |  +----------- Radiatula             
     +--+                                  
        |  +-------- Scabiellus            
        +--+                               
           |  +----- Harmandia_somboriensis **
           +--+                            
              |  +-- Unionetta             
              +--+                         
                 +-- Scabies               

Bolotov et al., 2022 [Publ 4252], fig. S1

iTtTbiriSoTroTroSiD
110160266--
  +----------------- Indonaia              
--+                                        
  |  +-------------- Indochinella          
  +--+                                     
     |  +----------- Radiatula             
     +--+                                  
        |  +-------- Scabiellus            
        +--+                               
           |  +----- Harmandia_somboriensis **
           +--+                            
              |  +-- Unionetta             
              +--+                         
                 +-- Scabies               

Konopleva et al., 2023 [Publ 4262], fig. 8

iTtTbiriSoTroTroSiD
110181136--
  +----------------- Indonaia              
--+                                        
  |  +-------------- Indochinella          
  +--+                                     
     |  +----------- Radiatula             
     +--+                                  
        |  +-------- Scabiellus            
        +--+                               
           |  +----- Scabies               
           +--+                            
              |  +-- Unionetta             
              +--+                         
                 +-- Harmandia_somboriensis **

Bolotov et al., 2022 [Publ 4239], fig. S2

iTtTbiriSoTroTroSiD
110157256--
  +----------------- Indonaia              
--+                                        
  |  +-------------- Indochinella          
  +--+                                     
     |  +----------- Radiatula             
     +--+                                  
        |  +-------- Scabiellus            
        +--+                               
           |  +----- Harmandia_somboriensis **
           +--+                            
              |  +-- Unionetta             
              +--+                         
                 +-- Scabies               

Bolotov et al., 2020 [Publ 4100], fig. 1

iTtTbiriSoTroTroSiD
1101116206--
  +----------------- Indonaia              
--+                                        
  |  +-------------- Indochinella          
  +--+                                     
     |  +----------- Radiatula             
     +--+                                  
        |  +-------- Scabiellus            
        +--+                               
           |  +----- Scabies               
           +--+                            
              |  +-- Unionetta             
              +--+                         
                 +-- Harmandia_somboriensis **

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iTtTbiriSoTroTroSiD
11015633--
  +-------- Indonaia              
--+                               
  |  +----- Indochinella          
  +--+                            
     |  +-- Scabies               
     +--+                         
        +-- Harmandia_somboriensis **

Bolotov et al., 2022 [Publ 4239], fig. S1

iTtTbiriSoTroTroSiD
110157256--
  +----------------- Indonaia              
--+                                        
  |  +-------------- Indochinella          
  +--+                                     
     |  +----------- Radiatula             
     +--+                                  
        |  +-------- Scabiellus            
        +--+                               
           |  +----- Harmandia_somboriensis **
           +--+                            
              |  +-- Unionetta             
              +--+                         
                 +-- Scabies               
 
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