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Page last updated
Thu 3 September 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

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Bivalvia | Unionida | Unionoidea | Unionidae | Parreysiinae | Indochinellini

genus Harmandia Rochebrune, 1882: cladomics
     1 Recent valid species (1 total species)

Cladomics Summary

number of species = 1 (1 total)
analyzed in 16 cladograms in 10 publications
maximum number of terminals in an analysis = 1 (mean = 1, median = 1)


Bolotov et al., 2024 [Publ 4369], fig. 2

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
110161287--
                 +-- Pseudoscabies         
  +--------------+                         
  |              +-- Indonaia              
--+                                        
  |  +-------------- Indochinella          
  +--+                                     
     |  +----------- Radiatula             
     +--+                                  
        |  +-------- Scabiellus            
        +--+                               
           |  +----- Harmandia_somboriensis **
           +--+                            
              |  +-- Unionetta             
              +--+                         
                 +-- Scabies               

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iTtTbiriSoTroTroSiD
11015633--
  +-------- Indonaia              
--+                               
  |  +----- Indochinella          
  +--+                            
     |  +-- Scabies               
     +--+                         
        +-- Harmandia_somboriensis **

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

iTtTbiriSoTroTroSiD
1101629276--
  +----- Harmandia_somboriensis **
--+                            
  |  +-- Unionetta             
  +--+                         
     +-- Scabies               

Bolotov et al., 2020 [Publ 4100], fig. S1

iTtTbiriSoTroTroSiD
1101110196--
  +-------------------- Indonaia               
  |                                            
--+  +----------------- Indochinella           
  |  |                                         
  |  |           +----- Radiatula_mouhoti      
  +--+  +--------+                             
     |  |        |  +-- Radiatula_myitkyinae   
     |  |        +--+                          
     +--+           +-- Radiatula_chaudhurii   
        |                                      
        |  +----------- Radiatula_crispisulcata
        +--+                                   
           |  +-------- Scabiellus             
           +--+                                
              |  +----- Harmandia_somboriensis ** 
              +--+                             
                 |  +-- Unionetta              
                 +--+                          
                    +-- Scabies                

Bolotov et al., 2020 [Publ 4100], fig. 1

iTtTbiriSoTroTroSiD
1101116206--
  +----------------- Indonaia              
--+                                        
  |  +-------------- Indochinella          
  +--+                                     
     |  +----------- Radiatula             
     +--+                                  
        |  +-------- Scabiellus            
        +--+                               
           |  +----- Scabies               
           +--+                            
              |  +-- Unionetta             
              +--+                         
                 +-- Harmandia_somboriensis **

Bolotov et al., 2024 [Publ 4369], fig. S1

iTtTbiriSoTroTroSiD
110162287--
                 +-- Pseudoscabies         
  +--------------+                         
  |              +-- Indonaia              
--+                                        
  |  +-------------- Indochinella          
  +--+                                     
     |  +----------- Radiatula             
     +--+                                  
        |  +-------- Scabiellus            
        +--+                               
           |  +----- Harmandia_somboriensis **
           +--+                            
              |  +-- Unionetta             
              +--+                         
                 +-- Scabies               

Jeratthitikul et al., 2019 [Publ 3922], fig. 1

iTtTbiriSoTroTroSiD
110122176--
  +----------------- Indonaia              
--+                                        
  |  +-------------- Indochinella          
  +--+                                     
     |  +----------- Radiatula             
     +--+                                  
        |  +-------- Scabiellus            
        +--+                               
           |  +----- Harmandia_somboriensis **
           +--+                            
              |  +-- Unionetta             
              +--+                         
                 +-- Scabies               

Bolotov et al., 2022 [Publ 4252], fig. 2

iTtTbiriSoTroTroSiD
110160266--
  +----------------- Indonaia              
--+                                        
  |  +-------------- Indochinella          
  +--+                                     
     |  +----------- Radiatula             
     +--+                                  
        |  +-------- Scabiellus            
        +--+                               
           |  +----- Harmandia_somboriensis **
           +--+                            
              |  +-- Unionetta             
              +--+                         
                 +-- Scabies               

Bolotov et al., 2022 [Publ 4252], fig. S1

iTtTbiriSoTroTroSiD
110160266--
  +----------------- Indonaia              
--+                                        
  |  +-------------- Indochinella          
  +--+                                     
     |  +----------- Radiatula             
     +--+                                  
        |  +-------- Scabiellus            
        +--+                               
           |  +----- Harmandia_somboriensis **
           +--+                            
              |  +-- Unionetta             
              +--+                         
                 +-- Scabies               

Chen et al., 2026 [Publ 4833], fig. 1

iTtTbiriSoTroTroSiD
110126135--
  +-------- Harmandia_somboriensis **      
  |                                     
--+     +-- Unionetta_xishuangbannaensis
  |  +--+                               
  |  |  +-- Unionetta_fabagina__2       
  +--+                                  
     |  +-- Unionetta_fabagina__1       
     +--+                               
        +-- Scabies                     

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iTtTbiriSoTroTroSiD
11015633--
  +-------- Indonaia              
--+                               
  |  +----- Indochinella          
  +--+                            
     |  +-- Scabies               
     +--+                         
        +-- Harmandia_somboriensis **

Konopleva et al., 2023 [Publ 4262], fig. 8

iTtTbiriSoTroTroSiD
110181136--
  +----------------- Indonaia              
--+                                        
  |  +-------------- Indochinella          
  +--+                                     
     |  +----------- Radiatula             
     +--+                                  
        |  +-------- Scabiellus            
        +--+                               
           |  +----- Scabies               
           +--+                            
              |  +-- Unionetta             
              +--+                         
                 +-- Harmandia_somboriensis **

Bolotov et al., 2022 [Publ 4239], fig. 2

iTtTbiriSoTroTroSiD
110157256--
  +----------------- Indonaia              
--+                                        
  |  +-------------- Indochinella          
  +--+                                     
     |  +----------- Radiatula             
     +--+                                  
        |  +-------- Scabiellus            
        +--+                               
           |  +----- Harmandia_somboriensis **
           +--+                            
              |  +-- Unionetta             
              +--+                         
                 +-- Scabies               

Bolotov et al., 2022 [Publ 4239], fig. S2

iTtTbiriSoTroTroSiD
110157256--
  +----------------- Indonaia              
--+                                        
  |  +-------------- Indochinella          
  +--+                                     
     |  +----------- Radiatula             
     +--+                                  
        |  +-------- Scabiellus            
        +--+                               
           |  +----- Harmandia_somboriensis **
           +--+                            
              |  +-- Unionetta             
              +--+                         
                 +-- Scabies               

Bolotov et al., 2022 [Publ 4239], fig. S1

iTtTbiriSoTroTroSiD
110157256--
  +----------------- Indonaia              
--+                                        
  |  +-------------- Indochinella          
  +--+                                     
     |  +----------- Radiatula             
     +--+                                  
        |  +-------- Scabiellus            
        +--+                               
           |  +----- Harmandia_somboriensis **
           +--+                            
              |  +-- Unionetta             
              +--+                         
                 +-- Scabies               

Pfeiffer et al., 2018 [Publ 3827], fig. 2

iTtTbiriSoTroTroSiD
110127166--
  +-------------------- Indonaia              
--+                                           
  |  +----------------- Indochinella          
  +--+                                        
     |  +-------------- Radiatula             
     +--+                                     
        |  +----------- Scabiellus            
        +--+                                  
           |  +-------- Unionetta_fabagina__2 
           +--+                               
              |  +----- Harmandia_somboriensis **
              +--+                            
                 |  +-- Unionetta_fabagina__1 
                 +--+                         
                    +-- Scabies               
 
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