The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)
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Bivalvia | Unionida | Unionoidea | Unionidae | Parreysiinae | Indochinellini genus Harmandia Rochebrune, 1882: cladomics 1 Recent valid species (1 total species) Cladomics Summarynumber of species = 1 (1 total) analyzed in 16 cladograms in 10 publications maximum number of terminals in an analysis =
1 (mean = 1, median = 1)
Bolotov et al., 2024 [Publ 4369], fig. 2 ** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram ** | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 61 | 28 | 7 | -- |
+-- Pseudoscabies
+--------------+
| +-- Indonaia
--+
| +-------------- Indochinella
+--+
| +----------- Radiatula
+--+
| +-------- Scabiellus
+--+
| +----- Harmandia_somboriensis **
+--+
| +-- Unionetta
+--+
+-- Scabies
Pfeiffer et al., 2019 [Publ 3937], fig. 6 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 56 | 3 | 3 | -- |
+-------- Indonaia
--+
| +----- Indochinella
+--+
| +-- Scabies
+--+
+-- Harmandia_somboriensis **
Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 629 | 27 | 6 | -- |
+----- Harmandia_somboriensis **
--+
| +-- Unionetta
+--+
+-- Scabies
Bolotov et al., 2020 [Publ 4100], fig. S1 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 110 | 19 | 6 | -- |
+-------------------- Indonaia
|
--+ +----------------- Indochinella
| |
| | +----- Radiatula_mouhoti
+--+ +--------+
| | | +-- Radiatula_myitkyinae
| | +--+
+--+ +-- Radiatula_chaudhurii
|
| +----------- Radiatula_crispisulcata
+--+
| +-------- Scabiellus
+--+
| +----- Harmandia_somboriensis **
+--+
| +-- Unionetta
+--+
+-- Scabies
Bolotov et al., 2020 [Publ 4100], fig. 1 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 116 | 20 | 6 | -- |
+----------------- Indonaia
--+
| +-------------- Indochinella
+--+
| +----------- Radiatula
+--+
| +-------- Scabiellus
+--+
| +----- Scabies
+--+
| +-- Unionetta
+--+
+-- Harmandia_somboriensis **
Bolotov et al., 2024 [Publ 4369], fig. S1 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 62 | 28 | 7 | -- |
+-- Pseudoscabies
+--------------+
| +-- Indonaia
--+
| +-------------- Indochinella
+--+
| +----------- Radiatula
+--+
| +-------- Scabiellus
+--+
| +----- Harmandia_somboriensis **
+--+
| +-- Unionetta
+--+
+-- Scabies
Jeratthitikul et al., 2019 [Publ 3922], fig. 1 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 22 | 17 | 6 | -- |
+----------------- Indonaia
--+
| +-------------- Indochinella
+--+
| +----------- Radiatula
+--+
| +-------- Scabiellus
+--+
| +----- Harmandia_somboriensis **
+--+
| +-- Unionetta
+--+
+-- Scabies
Bolotov et al., 2022 [Publ 4252], fig. 2 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 60 | 26 | 6 | -- |
+----------------- Indonaia
--+
| +-------------- Indochinella
+--+
| +----------- Radiatula
+--+
| +-------- Scabiellus
+--+
| +----- Harmandia_somboriensis **
+--+
| +-- Unionetta
+--+
+-- Scabies
Bolotov et al., 2022 [Publ 4252], fig. S1 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 60 | 26 | 6 | -- |
+----------------- Indonaia
--+
| +-------------- Indochinella
+--+
| +----------- Radiatula
+--+
| +-------- Scabiellus
+--+
| +----- Harmandia_somboriensis **
+--+
| +-- Unionetta
+--+
+-- Scabies
Chen et al., 2026 [Publ 4833], fig. 1 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 26 | 13 | 5 | -- |
+-------- Harmandia_somboriensis **
|
--+ +-- Unionetta_xishuangbannaensis
| +--+
| | +-- Unionetta_fabagina__2
+--+
| +-- Unionetta_fabagina__1
+--+
+-- Scabies
Pfeiffer et al., 2019 [Publ 3937], fig. 4-5 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 56 | 3 | 3 | -- |
+-------- Indonaia
--+
| +----- Indochinella
+--+
| +-- Scabies
+--+
+-- Harmandia_somboriensis **
Konopleva et al., 2023 [Publ 4262], fig. 8 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 81 | 13 | 6 | -- |
+----------------- Indonaia
--+
| +-------------- Indochinella
+--+
| +----------- Radiatula
+--+
| +-------- Scabiellus
+--+
| +----- Scabies
+--+
| +-- Unionetta
+--+
+-- Harmandia_somboriensis **
Bolotov et al., 2022 [Publ 4239], fig. 2 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 57 | 25 | 6 | -- |
+----------------- Indonaia
--+
| +-------------- Indochinella
+--+
| +----------- Radiatula
+--+
| +-------- Scabiellus
+--+
| +----- Harmandia_somboriensis **
+--+
| +-- Unionetta
+--+
+-- Scabies
Bolotov et al., 2022 [Publ 4239], fig. S2 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 57 | 25 | 6 | -- |
+----------------- Indonaia
--+
| +-------------- Indochinella
+--+
| +----------- Radiatula
+--+
| +-------- Scabiellus
+--+
| +----- Harmandia_somboriensis **
+--+
| +-- Unionetta
+--+
+-- Scabies
Bolotov et al., 2022 [Publ 4239], fig. S1 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 57 | 25 | 6 | -- |
+----------------- Indonaia
--+
| +-------------- Indochinella
+--+
| +----------- Radiatula
+--+
| +-------- Scabiellus
+--+
| +----- Harmandia_somboriensis **
+--+
| +-- Unionetta
+--+
+-- Scabies
Pfeiffer et al., 2018 [Publ 3827], fig. 2 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 27 | 16 | 6 | -- |
+-------------------- Indonaia
--+
| +----------------- Indochinella
+--+
| +-------------- Radiatula
+--+
| +----------- Scabiellus
+--+
| +-------- Unionetta_fabagina__2
+--+
| +----- Harmandia_somboriensis **
+--+
| +-- Unionetta_fabagina__1
+--+
+-- Scabies
|