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Page last updated
Thu 3 September 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Search the database pages:

Bivalvia | Unionida | Unionoidea | Unionidae | Unioninae | Unionini | Aculamprotulina

genus Scriptolamprotula Modell, 1964: cladomics
     9 Recent valid species (9 total species)

Cladomics Summary

number of species = 9 (9 total)
analyzed in 114 cladograms in 64 publications
maximum number of terminals in an analysis = 6 (mean = 2.68, median = 2)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 67
maximum number of included terminals = 6, (mean = 3.87 median = 4)
proportion of cladograms that support basic monophyly = 1

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
6657625000
  +----------------- Scriptolamprotula_tortuosa **     
--+                                                 
  |  +-------------- Scriptolamprotula_tientsinensis **
  +--+                                              
     |  +----------- Scriptolamprotula_fibrosa **      
     +--+                                           
        |  +-------- Scriptolamprotula_coreana **      
        +--+                                        
           |  +----- Scriptolamprotula_scripta__2 **   
           +--+                                     
              |  +-- Scriptolamprotula_polysticta **   
              +--+                                  
                 +-- Scriptolamprotula_scripta__1 **

Wu et al., 2020 [Publ 4121], fig. 3

iTtTbiriSoTroTroSiD
553544004
  +-------- Scriptolamprotula_tortuosa **     
  |                                        
--+     +-- Scriptolamprotula_coreana **      
  |  +--+                                  
  |  |  +-- Scriptolamprotula_tientsinensis **
  +--+                                     
     |  +-- Scriptolamprotula_polysticta **   
     +--+                                  
        +-- Scriptolamprotula_scripta **

Wu et al., 2018 [Publ 3836], fig. 2

iTtTbiriSoTroTroSiD
553548006
  +----------- Scriptolamprotula_tortuosa **     
--+                                           
  |  +-------- Scriptolamprotula_scripta **      
  +--+                                        
     |  +----- Scriptolamprotula_coreana **      
     +--+                                     
        |  +-- Scriptolamprotula_fibrosa **      
        +--+                                  
           +-- Scriptolamprotula_tientsinensis **

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

iTtTbiriSoTroTroSiD
5535136004
  +-------- Scriptolamprotula_tortuosa **     
  |                                        
--+     +-- Scriptolamprotula_coreana **      
  |  +--+                                  
  |  |  +-- Scriptolamprotula_tientsinensis **
  +--+                                     
     |  +-- Scriptolamprotula_polysticta **   
     +--+                                  
        +-- Scriptolamprotula_scripta **

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iTtTbiriSoTroTroSiD
553591004
  +-------- Scriptolamprotula_tortuosa **     
  |                                        
--+     +-- Scriptolamprotula_coreana **      
  |  +--+                                  
  |  |  +-- Scriptolamprotula_tientsinensis **
  +--+                                     
     |  +-- Scriptolamprotula_polysticta **   
     +--+                                  
        +-- Scriptolamprotula_scripta **

Wu et al., 2021 [Publ 4366], fig. 3-BI

iTtTbiriSoTroTroSiD
553569004
  +-------- Scriptolamprotula_tortuosa **     
  |                                        
--+     +-- Scriptolamprotula_scripta **      
  |  +--+                                  
  |  |  +-- Scriptolamprotula_polysticta **   
  +--+                                     
     |  +-- Scriptolamprotula_coreana **      
     +--+                                  
        +-- Scriptolamprotula_tientsinensis **

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iTtTbiriSoTroTroSiD
553587004
  +-------- Scriptolamprotula_tortuosa **     
  |                                        
--+     +-- Scriptolamprotula_coreana **      
  |  +--+                                  
  |  |  +-- Scriptolamprotula_tientsinensis **
  +--+                                     
     |  +-- Scriptolamprotula_polysticta **   
     +--+                                  
        +-- Scriptolamprotula_scripta **

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
5525195005
  +-------- Scriptolamprotula_tortuosa **     
  |                                        
--+  +----- Scriptolamprotula_coreana **      
  |  |                                     
  +--+----- Scriptolamprotula_scripta **      
     |                                     
     |  +-- Scriptolamprotula_fibrosa **      
     +--+                                  
        +-- Scriptolamprotula_tientsinensis **

Wu et al., 2026 [Publ 4811], fig. 4

iTtTbiriSoTroTroSiD
553590004
  +-------- Scriptolamprotula_tortuosa **     
  |                                        
--+     +-- Scriptolamprotula_coreana **      
  |  +--+                                  
  |  |  +-- Scriptolamprotula_tientsinensis **
  +--+                                     
     |  +-- Scriptolamprotula_polysticta **   
     +--+                                  
        +-- Scriptolamprotula_scripta **

Lopes-Lima et al., 2020 [Publ 4080], fig. 5

iTtTbiriSoTroTroSiD
55354002
  +----------- Scriptolamprotula_tortuosa **     
--+                                           
  |  +-------- Scriptolamprotula_tientsinensis **
  +--+                                        
     |  +----- Scriptolamprotula_fibrosa **      
     +--+                                     
        |  +-- Scriptolamprotula_scripta **      
        +--+                                  
           +-- Scriptolamprotula_coreana **

Wu et al., 2022 [Publ 4238], fig. 3

iTtTbiriSoTroTroSiD
553569004
  +-------- Scriptolamprotula_tortuosa **     
  |                                        
--+     +-- Scriptolamprotula_tientsinensis **
  |  +--+                                  
  |  |  +-- Scriptolamprotula_coreana **      
  +--+                                     
     |  +-- Scriptolamprotula_polysticta **   
     +--+                                  
        +-- Scriptolamprotula_scripta **

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
5535193004
  +-------- Scriptolamprotula_tortuosa **     
  |                                        
--+     +-- Scriptolamprotula_coreana **      
  |  +--+                                  
  |  |  +-- Scriptolamprotula_scripta **      
  +--+                                     
     |  +-- Scriptolamprotula_fibrosa **      
     +--+                                  
        +-- Scriptolamprotula_tientsinensis **

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iTtTbiriSoTroTroSiD
5535117006
  +----------- Scriptolamprotula_tortuosa **     
--+                                           
  |  +-------- Scriptolamprotula_scripta **      
  +--+                                        
     |  +----- Scriptolamprotula_coreana **      
     +--+                                     
        |  +-- Scriptolamprotula_fibrosa **      
        +--+                                  
           +-- Scriptolamprotula_tientsinensis **

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iTtTbiriSoTroTroSiD
553591004
  +-------- Scriptolamprotula_tortuosa **     
  |                                        
--+     +-- Scriptolamprotula_coreana **      
  |  +--+                                  
  |  |  +-- Scriptolamprotula_tientsinensis **
  +--+                                     
     |  +-- Scriptolamprotula_polysticta **   
     +--+                                  
        +-- Scriptolamprotula_scripta **

Dai et al., 2025 [Publ 4440], fig. 1

iTtTbiriSoTroTroSiD
552540003
  +-------- Scriptolamprotula_tortuosa **  
  |                                     
--+  +----- Scriptolamprotula_coreana **   
  |  |                                  
  +--+----- Scriptolamprotula_fibrosa **   
     |                                  
     |  +-- Scriptolamprotula_polysticta **
     +--+                               
        +-- Scriptolamprotula_scripta **

Hou et al., 2025 [Publ 4445], fig. 5

iTtTbiriSoTroTroSiD
553548004
  +-------- Scriptolamprotula_tortuosa **     
  |                                        
--+     +-- Scriptolamprotula_coreana **      
  |  +--+                                  
  |  |  +-- Scriptolamprotula_tientsinensis **
  +--+                                     
     |  +-- Scriptolamprotula_polysticta **   
     +--+                                  
        +-- Scriptolamprotula_scripta **

Huang et al., 2019 [Publ 3843], fig. S1

iTtTbiriSoTroTroSiD
553542002
  +----------- Scriptolamprotula_tortuosa **     
--+                                           
  |  +-------- Scriptolamprotula_tientsinensis **
  +--+                                        
     |  +----- Scriptolamprotula_fibrosa **      
     +--+                                     
        |  +-- Scriptolamprotula_coreana **      
        +--+                                  
           +-- Scriptolamprotula_scripta **

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
5535195004
  +-------- Scriptolamprotula_tortuosa **     
  |                                        
--+     +-- Scriptolamprotula_coreana **      
  |  +--+                                  
  |  |  +-- Scriptolamprotula_scripta **      
  +--+                                     
     |  +-- Scriptolamprotula_fibrosa **      
     +--+                                  
        +-- Scriptolamprotula_tientsinensis **

Wu et al., 2021 [Publ 4366], fig. 3-ML

iTtTbiriSoTroTroSiD
553569004
  +-------- Scriptolamprotula_tortuosa **     
  |                                        
--+     +-- Scriptolamprotula_scripta **      
  |  +--+                                  
  |  |  +-- Scriptolamprotula_polysticta **   
  +--+                                     
     |  +-- Scriptolamprotula_coreana **      
     +--+                                  
        +-- Scriptolamprotula_tientsinensis **

Hou et al., 2024 [Publ 4400], fig. 4

iTtTbiriSoTroTroSiD
553571004
  +-------- Scriptolamprotula_tortuosa **     
  |                                        
--+     +-- Scriptolamprotula_coreana **      
  |  +--+                                  
  |  |  +-- Scriptolamprotula_tientsinensis **
  +--+                                     
     |  +-- Scriptolamprotula_polysticta **   
     +--+                                  
        +-- Scriptolamprotula_scripta **

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iTtTbiriSoTroTroSiD
553587004
  +-------- Scriptolamprotula_tortuosa **     
  |                                        
--+     +-- Scriptolamprotula_coreana **      
  |  +--+                                  
  |  |  +-- Scriptolamprotula_tientsinensis **
  +--+                                     
     |  +-- Scriptolamprotula_polysticta **   
     +--+                                  
        +-- Scriptolamprotula_scripta **

Dai et al., 2024 [Publ 4386], fig. 6

iTtTbiriSoTroTroSiD
553537004
  +-------- Scriptolamprotula_tortuosa **     
  |                                        
--+     +-- Scriptolamprotula_polysticta **   
  |  +--+                                  
  |  |  +-- Scriptolamprotula_scripta **      
  +--+                                     
     |  +-- Scriptolamprotula_coreana **      
     +--+                                  
        +-- Scriptolamprotula_tientsinensis **

Dai et al., 2025 [Publ 4440], fig. 2

iTtTbiriSoTroTroSiD
553540004
  +-------- Scriptolamprotula_tortuosa **  
  |                                     
--+     +-- Scriptolamprotula_fibrosa **   
  |  +--+                               
  |  |  +-- Scriptolamprotula_coreana **   
  +--+                                  
     |  +-- Scriptolamprotula_scripta **   
     +--+                               
        +-- Scriptolamprotula_polysticta **

Wu et al., 2021 [Publ 4366], fig. 4

iTtTbiriSoTroTroSiD
553564004
  +-------- Scriptolamprotula_tortuosa **     
  |                                        
--+     +-- Scriptolamprotula_scripta **      
  |  +--+                                  
  |  |  +-- Scriptolamprotula_polysticta **   
  +--+                                     
     |  +-- Scriptolamprotula_coreana **      
     +--+                                  
        +-- Scriptolamprotula_tientsinensis **

Wu et al., 2024 [Publ 4379], fig. 5

iTtTbiriSoTroTroSiD
553549004
  +-------- Scriptolamprotula_tortuosa **     
  |                                        
--+     +-- Scriptolamprotula_coreana **      
  |  +--+                                  
  |  |  +-- Scriptolamprotula_tientsinensis **
  +--+                                     
     |  +-- Scriptolamprotula_polysticta **   
     +--+                                  
        +-- Scriptolamprotula_scripta **

Dai et al., 2025 [Publ 4440], fig. 3

iTtTbiriSoTroTroSiD
553583004
  +-------- Scriptolamprotula_tortuosa **     
  |                                        
--+     +-- Scriptolamprotula_polysticta **   
  |  +--+                                  
  |  |  +-- Scriptolamprotula_scripta **      
  +--+                                     
     |  +-- Scriptolamprotula_coreana **      
     +--+                                  
        +-- Scriptolamprotula_tientsinensis **

Chen et al., 2025 [Publ 4448], fig. 3

iTtTbiriSoTroTroSiD
553581004
  +-------- Scriptolamprotula_tortuosa **     
  |                                        
--+     +-- Scriptolamprotula_coreana **      
  |  +--+                                  
  |  |  +-- Scriptolamprotula_tientsinensis **
  +--+                                     
     |  +-- Scriptolamprotula_polysticta **   
     +--+                                  
        +-- Scriptolamprotula_scripta **

Wang et al., 2025 [Publ 4488], fig. 3B

iTtTbiriSoTroTroSiD
553560004
  +-------- Scriptolamprotula_tortuosa **     
  |                                        
--+     +-- Scriptolamprotula_coreana **      
  |  +--+                                  
  |  |  +-- Scriptolamprotula_tientsinensis **
  +--+                                     
     |  +-- Scriptolamprotula_polysticta **   
     +--+                                  
        +-- Scriptolamprotula_scripta **

Huang et al., 2019 [Publ 3843], fig. 1

iTtTbiriSoTroTroSiD
553540006
  +----------- Scriptolamprotula_tortuosa **     
--+                                           
  |  +-------- Scriptolamprotula_scripta **      
  +--+                                        
     |  +----- Scriptolamprotula_coreana **      
     +--+                                     
        |  +-- Scriptolamprotula_fibrosa **      
        +--+                                  
           +-- Scriptolamprotula_tientsinensis **

Wu et al., 2022 [Publ 4238], fig. 2d

iTtTbiriSoTroTroSiD
553534004
  +-------- Scriptolamprotula_tortuosa **     
  |                                        
--+     +-- Scriptolamprotula_tientsinensis **
  |  +--+                                  
  |  |  +-- Scriptolamprotula_coreana **      
  +--+                                     
     |  +-- Scriptolamprotula_polysticta **   
     +--+                                  
        +-- Scriptolamprotula_scripta **

Konopleva et al., 2023 [Publ 4269], fig. S1_4

iTtTbiriSoTroTroSiD
442452005
  +-------- Scriptolamprotula_tortuosa **     
--+                                        
  |  +----- Scriptolamprotula_scripta **      
  +--+                                     
     |  +-- Scriptolamprotula_tientsinensis **
     +--+                                  
        +-- Scriptolamprotula_coreana **

Wu et al., 2020 [Publ 4121], fig. 2

iTtTbiriSoTroTroSiD
442430005
  +-------- Scriptolamprotula_tortuosa **     
--+                                        
  |  +----- Scriptolamprotula_polysticta **   
  +--+                                     
     |  +-- Scriptolamprotula_fibrosa **      
     +--+                                  
        +-- Scriptolamprotula_tientsinensis **

Wu et al., 2019 [Publ 3864], fig. 3

iTtTbiriSoTroTroSiD
442442005
  +-------- Scriptolamprotula_tortuosa **     
--+                                        
  |  +----- Scriptolamprotula_scripta **      
  +--+                                     
     |  +-- Scriptolamprotula_tientsinensis **
     +--+                                  
        +-- Scriptolamprotula_coreana **

Wu et al., 2020 [Publ 4121], fig. S1

iTtTbiriSoTroTroSiD
442430005
  +-------- Scriptolamprotula_tortuosa **     
--+                                        
  |  +----- Scriptolamprotula_polysticta **   
  +--+                                     
     |  +-- Scriptolamprotula_tientsinensis **
     +--+                                  
        +-- Scriptolamprotula_fibrosa **

Huang et al., 2019 [Publ 3843], fig. 3

iTtTbiriSoTroTroSiD
442452005
  +-------- Scriptolamprotula_tortuosa **     
--+                                        
  |  +----- Scriptolamprotula_scripta **      
  +--+                                     
     |  +-- Scriptolamprotula_tientsinensis **
     +--+                                  
        +-- Scriptolamprotula_coreana **

Wu et al., 2020 [Publ 4147], fig. 3

iTtTbiriSoTroTroSiD
442426005
  +-------- Scriptolamprotula_tortuosa **     
--+                                        
  |  +----- Scriptolamprotula_scripta **      
  +--+                                     
     |  +-- Scriptolamprotula_tientsinensis **
     +--+                                  
        +-- Scriptolamprotula_coreana **

Wu et al., 2018 [Publ 4143], fig. 2

iTtTbiriSoTroTroSiD
442418005
  +-------- Scriptolamprotula_tortuosa **     
--+                                        
  |  +----- Scriptolamprotula_scripta **      
  +--+                                     
     |  +-- Scriptolamprotula_coreana **      
     +--+                                  
        +-- Scriptolamprotula_tientsinensis **

Wu et al., 2020 [Publ 4147], fig. 4

iTtTbiriSoTroTroSiD
442410005
  +-------- Scriptolamprotula_tortuosa **     
--+                                        
  |  +----- Scriptolamprotula_scripta **      
  +--+                                     
     |  +-- Scriptolamprotula_tientsinensis **
     +--+                                  
        +-- Scriptolamprotula_fibrosa **

Froufe et al., 2019 [Publ 3936], fig. 3

iTtTbiriSoTroTroSiD
331341004
  +----- Scriptolamprotula_tortuosa **     
--+                                     
  |  +-- Scriptolamprotula_tientsinensis **
  +--+                                  
     +-- Scriptolamprotula_coreana **

Sano et al., 2020 [Publ 4138], fig. 3

iTtTbiriSoTroTroSiD
331365004
  +----- Scriptolamprotula_tortuosa **     
--+                                     
  |  +-- Scriptolamprotula_tientsinensis **
  +--+                                  
     +-- Scriptolamprotula_coreana **

Soroka & Burzynski, 2017 [Publ 3837], fig. 1

iTtTbiriSoTroTroSiD
331320004
  +----- Scriptolamprotula_tortuosa **     
--+                                     
  |  +-- Scriptolamprotula_tientsinensis **
  +--+                                  
     +-- Scriptolamprotula_coreana **

Sano et al., 2020 [Publ 4138], fig. 2

iTtTbiriSoTroTroSiD
331365004
  +----- Scriptolamprotula_tortuosa **     
--+                                     
  |  +-- Scriptolamprotula_tientsinensis **
  +--+                                  
     +-- Scriptolamprotula_coreana **

Froufe et al., 2019 [Publ 3936], fig. 6

iTtTbiriSoTroTroSiD
331341004
  +----- Scriptolamprotula_tortuosa **     
--+                                     
  |  +-- Scriptolamprotula_tientsinensis **
  +--+                                  
     +-- Scriptolamprotula_coreana **

Zhou et al., 2007 [Publ 2658], fig. 2

iTtTbiriSoTroTroSiD
331315004
  +----- Scriptolamprotula_tortuosa **     
--+                                     
  |  +-- Scriptolamprotula_tientsinensis **
  +--+                                  
     +-- Scriptolamprotula_scripta **

Fukata & Iigo, 2020 [Publ 4106], fig. 1

iTtTbiriSoTroTroSiD
331351004
  +----- Scriptolamprotula_tortuosa **     
--+                                     
  |  +-- Scriptolamprotula_tientsinensis **
  +--+                                  
     +-- Scriptolamprotula_coreana **

Wu et al., 2021 [Publ 4366], fig. 6

iTtTbiriSoTroTroSiD
331388004
  +----- Scriptolamprotula_tortuosa **     
--+                                     
  |  +-- Scriptolamprotula_tientsinensis **
  +--+                                  
     +-- Scriptolamprotula_scripta **

Cha et al., 2018 [Publ 3916], fig. 1

iTtTbiriSoTroTroSiD
331335004
  +----- Scriptolamprotula_tortuosa **     
--+                                     
  |  +-- Scriptolamprotula_coreana **      
  +--+                                  
     +-- Scriptolamprotula_tientsinensis **

Aunins et al., 2018 [Publ 3863], fig. 1

iTtTbiriSoTroTroSiD
331337004
  +----- Scriptolamprotula_tortuosa **     
--+                                     
  |  +-- Scriptolamprotula_coreana **      
  +--+                                  
     +-- Scriptolamprotula_tientsinensis **

Huang et al., 2018 [Publ 3828], fig. 6

iTtTbiriSoTroTroSiD
331339004
  +----- Scriptolamprotula_tortuosa **     
--+                                     
  |  +-- Scriptolamprotula_coreana **      
  +--+                                  
     +-- Scriptolamprotula_tientsinensis **

Sano et al., 2020 [Publ 4138], fig. 4

iTtTbiriSoTroTroSiD
331365004
  +----- Scriptolamprotula_tortuosa **     
--+                                     
  |  +-- Scriptolamprotula_tientsinensis **
  +--+                                  
     +-- Scriptolamprotula_coreana **

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

iTtTbiriSoTroTroSiD
331328004
  +----- Scriptolamprotula_tortuosa **     
--+                                     
  |  +-- Scriptolamprotula_tientsinensis **
  +--+                                  
     +-- Scriptolamprotula_scripta **

Wu et al., 2016 [Publ 3711], fig. 1

iTtTbiriSoTroTroSiD
331318004
  +----- Scriptolamprotula_tortuosa **     
--+                                     
  |  +-- Scriptolamprotula_coreana **      
  +--+                                  
     +-- Scriptolamprotula_tientsinensis **

Zhou et al., 2007 [Publ 2658], fig. 1

iTtTbiriSoTroTroSiD
331318004
  +----- Scriptolamprotula_tortuosa **     
--+                                     
  |  +-- Scriptolamprotula_tientsinensis **
  +--+                                  
     +-- Scriptolamprotula_scripta **

Wu et al., 2024 [Publ 4379], fig. 4_6

iTtTbiriSoTroTroSiD
331320004
  +----- Scriptolamprotula_tortuosa **     
--+                                     
  |  +-- Scriptolamprotula_scripta **      
  +--+                                  
     +-- Scriptolamprotula_tientsinensis **

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

iTtTbiriSoTroTroSiD
331345004
  +----- Scriptolamprotula_tortuosa **     
--+                                     
  |  +-- Scriptolamprotula_tientsinensis **
  +--+                                  
     +-- Scriptolamprotula_coreana **

Wu et al., 2021 [Publ 4366], fig. 2_I

iTtTbiriSoTroTroSiD
331388004
  +----- Scriptolamprotula_tortuosa **     
--+                                     
  |  +-- Scriptolamprotula_scripta **      
  +--+                                  
     +-- Scriptolamprotula_tientsinensis **

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

iTtTbiriSoTroTroSiD
22023700--
  +-- Scriptolamprotula_tientsinensis **
--+                                  
  +-- Scriptolamprotula_tortuosa **

Liu et al., 2023 [Publ 4310], fig. 5

iTtTbiriSoTroTroSiD
22022200--
  +-- Scriptolamprotula_tientsinensis **
--+                                  
  +-- Scriptolamprotula_scripta **

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

iTtTbiriSoTroTroSiD
22023500--
  +-- Scriptolamprotula_tientsinensis **
--+                                  
  +-- Scriptolamprotula_tortuosa **

Lopes-Lima et al., 2017 [Publ 3792], fig. 2

iTtTbiriSoTroTroSiD
22023000--
  +-- Scriptolamprotula_tortuosa **
--+                             
  +-- Scriptolamprotula_coreana **

Chen et al., 2020 [Publ 4170], fig. 1

iTtTbiriSoTroTroSiD
22023200--
  +-- Scriptolamprotula_coreana ** 
--+                             
  +-- Scriptolamprotula_tortuosa **

Liu et al., 2023 [Publ 4310], fig. 4

iTtTbiriSoTroTroSiD
22022200--
  +-- Scriptolamprotula_scripta **      
--+                                  
  +-- Scriptolamprotula_tientsinensis **

Zhou et al., 2017 [Publ 3788], fig. 2

iTtTbiriSoTroTroSiD
22021400--
  +-- Scriptolamprotula_tientsinensis **
--+                                  
  +-- Scriptolamprotula_tortuosa **

Wen et al., 2017 [Publ 3795], fig. 3

iTtTbiriSoTroTroSiD
22021400--
  +-- Scriptolamprotula_coreana ** 
--+                             
  +-- Scriptolamprotula_tortuosa **

Dai et al., 2024 [Publ 4358], fig. 4

iTtTbiriSoTroTroSiD
22025100--
  +-- Scriptolamprotula_scripta **      
--+                                  
  +-- Scriptolamprotula_tientsinensis **

Wu et al., 2023 [Publ 4296], fig. 4

iTtTbiriSoTroTroSiD
22022100--
  +-- Scriptolamprotula_scripta **      
--+                                  
  +-- Scriptolamprotula_tientsinensis **

Fonseca et al., 2016 [Publ 3775], fig. 1

iTtTbiriSoTroTroSiD
22022900--
  +-- Scriptolamprotula_tortuosa **
--+                             
  +-- Scriptolamprotula_coreana **

Dai et al., 2024 [Publ 4386], fig. 5-BI

iTtTbiriSoTroTroSiD
11012100--
-- Scriptolamprotula_scripta **

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

iTtTbiriSoTroTroSiD
11017500--
-- Scriptolamprotula_tortuosa **

Chen et al., 2025 [Publ 4487], fig. 2

iTtTbiriSoTroTroSiD
11014000--
-- Scriptolamprotula_fibrosa **

Chen et al., 2023 [Publ 4302], fig. 2

iTtTbiriSoTroTroSiD
11014000--
-- Scriptolamprotula_fibrosa **

Xue et al., 2016 [Publ 3659], fig. 1

iTtTbiriSoTroTroSiD
11011400--
-- Scriptolamprotula_tortuosa **

Sonowal et al., 2021 [Publ 4221], fig. 3a

iTtTbiriSoTroTroSiD
11011400--
-- Scriptolamprotula_tientsinensis **

Guerra et al., 2019 [Publ 4168], fig. 1-M

iTtTbiriSoTroTroSiD
11012100--
-- Scriptolamprotula_tortuosa **

Chen et al., 2025 [Publ 4487], fig. 3

iTtTbiriSoTroTroSiD
11014000--
-- Scriptolamprotula_fibrosa **

Bolotov et al., 2017 [Publ 3761], fig. S2

iTtTbiriSoTroTroSiD
110113700--
-- Scriptolamprotula_tortuosa **

Bolotov et al., 2017 [Publ 3764], fig. S1

iTtTbiriSoTroTroSiD
110112300--
-- Scriptolamprotula_tortuosa **

Sonowal et al., 2021 [Publ 4221], fig. 3b

iTtTbiriSoTroTroSiD
11011400--
-- Scriptolamprotula_tientsinensis **

Soroka & Burzynski, 2018 [Publ 3762], fig. 3

iTtTbiriSoTroTroSiD
1101600--
-- Scriptolamprotula_tortuosa **

Wu et al., 2022 [Publ 4238], fig. 2e

iTtTbiriSoTroTroSiD
11011700--
-- Scriptolamprotula_tortuosa **

Burzynski & Soroka, 2018 [Publ 3862], fig. 5a

iTtTbiriSoTroTroSiD
11011900--
-- Scriptolamprotula_tortuosa **

Guerra et al., 2019 [Publ 4168], fig. 2

iTtTbiriSoTroTroSiD
11014000--
-- Scriptolamprotula_tortuosa **

Wu et al., 2022 [Publ 4238], fig. 2a_S1

iTtTbiriSoTroTroSiD
11011100--
-- Scriptolamprotula_tientsinensis **

Dai et al., 2024 [Publ 4386], fig. 3

iTtTbiriSoTroTroSiD
11011300--
-- Scriptolamprotula_scripta **

Burzynski et al., 2017 [Publ 3774], fig. 1-top

iTtTbiriSoTroTroSiD
11011200--
-- Scriptolamprotula_tortuosa **

Guerra et al., 2017 [Publ 3772], fig. 2b

iTtTbiriSoTroTroSiD
11012400--
-- Scriptolamprotula_tortuosa **

Wang et al., 2018 [Publ 3865], fig. 2

iTtTbiriSoTroTroSiD
11013000--
-- Scriptolamprotula_tientsinensis **

Burzynski et al., 2017 [Publ 3774], fig. 1-bottom

iTtTbiriSoTroTroSiD
11011200--
-- Scriptolamprotula_tortuosa **

Araujo et al., 2017 [Publ 3801], fig. 2

iTtTbiriSoTroTroSiD
11011300--
-- Scriptolamprotula_tortuosa **

Dai et al., 2024 [Publ 4386], fig. 5-ML

iTtTbiriSoTroTroSiD
11012100--
-- Scriptolamprotula_scripta **

Kim et al., 2021 [Publ 4197], fig. 1

iTtTbiriSoTroTroSiD
11011700--
-- Scriptolamprotula_tientsinensis **

Guerra et al., 2019 [Publ 4168], fig. 2-M

iTtTbiriSoTroTroSiD
11012100--
-- Scriptolamprotula_tortuosa **

Chen et al., 2023 [Publ 4302], fig. 1

iTtTbiriSoTroTroSiD
11014000--
-- Scriptolamprotula_fibrosa **

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

iTtTbiriSoTroTroSiD
11017500--
-- Scriptolamprotula_tortuosa **

Campbell & Lydeard, 2026 [Publ 4810], fig. 4

iTtTbiriSoTroTroSiD
11014600--
-- Scriptolamprotula_tortuosa **

Huang et al., 2018 [Publ 3828], fig. 3_5

iTtTbiriSoTroTroSiD
11012800--
-- Scriptolamprotula_tientsinensis **

Fukata & Iigo, 2019 [Publ 4108], fig. 1

iTtTbiriSoTroTroSiD
11011600--
-- Scriptolamprotula_tientsinensis **

Burzynski & Soroka, 2018 [Publ 3862], fig. 5b

iTtTbiriSoTroTroSiD
1101900--
-- Scriptolamprotula_tortuosa **

Rosenberg et al., 1997 [Publ 2720], fig. 2B

iTtTbiriSoTroTroSiD
11012100--
-- Scriptolamprotula_tortuosa **

Dai et al., 2023 [Publ 4285], fig. 5

iTtTbiriSoTroTroSiD
11015200--
-- Scriptolamprotula_tientsinensis **

Ouyang et al., 2015 [Publ 4493], fig. 1

iTtTbiriSoTroTroSiD
11013000--
-- Scriptolamprotula_tortuosa **

Ouyang et al., 2011 [Publ 3408], fig. 1

iTtTbiriSoTroTroSiD
11011200--
-- Scriptolamprotula_tortuosa **

Wu et al., 2024 [Publ 4388], fig. 6

iTtTbiriSoTroTroSiD
11015200--
-- Scriptolamprotula_tortuosa **

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iTtTbiriSoTroTroSiD
11017900--
-- Scriptolamprotula_tortuosa **

Sonowal et al., 2021 [Publ 4221], fig. 4

iTtTbiriSoTroTroSiD
11011500--
-- Scriptolamprotula_tientsinensis **

Sano et al., 2020 [Publ 4138], fig. 1

iTtTbiriSoTroTroSiD
11013300--
-- Scriptolamprotula_tortuosa **

Guerra et al., 2019 [Publ 4168], fig. 1

iTtTbiriSoTroTroSiD
11014000--
-- Scriptolamprotula_tortuosa **

Bolotov et al., 2017 [Publ 3761], fig. S1

iTtTbiriSoTroTroSiD
110113700--
-- Scriptolamprotula_tortuosa **

Soroka & Burzynski, 2018 [Publ 3762], fig. 3-M

iTtTbiriSoTroTroSiD
1101600--
-- Scriptolamprotula_tortuosa **

Guerra et al., 2017 [Publ 3772], fig. 2a

iTtTbiriSoTroTroSiD
11012400--
-- Scriptolamprotula_tortuosa **

Ouyang et al., 2015 [Publ 4493], fig. 2

iTtTbiriSoTroTroSiD
11013000--
-- Scriptolamprotula_tortuosa **

Wang et al., 2018 [Publ 3846], fig. 2

iTtTbiriSoTroTroSiD
11013000--
-- Scriptolamprotula_tientsinensis **

Huang et al., 2019 [Publ 3843], fig. 3-M

iTtTbiriSoTroTroSiD
11012200--
-- Scriptolamprotula_tortuosa **

Huang et al., 2018 [Publ 3828], fig. 6-M

iTtTbiriSoTroTroSiD
11011600--
-- Scriptolamprotula_tortuosa **
 
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