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Page last updated
Thu 3 September 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

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Bivalvia | Unionida | Hyrioidea | Hyriidae | Hyriinae | Hyridellini

genus Hyridella Swainson, 1840: cladomics
     7 Recent valid species (10 total species)

Cladomics Summary

number of species = 7 (10 total)
analyzed in 44 cladograms in 26 publications
maximum number of terminals in an analysis = 3 (mean = 1.5, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 15
maximum number of included terminals = 3, (mean = 2.47 median = 2)
proportion of cladograms that support basic monophyly = 0.47

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.


PCoA Ordination


NJ Tree

Santos-Neto et al., 2016 [Publ 3739], fig. 2a_b

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
31920317110
                 +-------- Westralunio           
                 |                               
  +--------------+     +-- Hyridella_drapeta **     
  |              |  +--+                         
  |              |  |  +-- Hyridella_depressa **    
  |              +--+                            
  |                 |  +-- Diplodon_gratus       
  |                 +--+                         
  |                    +-- Diplodon_suavidicus__2
  |                                              
  |                    +-- Hyridella_australis **   
  |     +--------------+                         
  |     |              +-- Cucumerunio           
  +-----+                                        
  |     |  +-------------- Echyridella           
--+     +--+                                     
  |        |  +----------- Velesunio_ambiguus    
  |        +--+                                  
  |           |  +-------- Microdontia           
  |           +--+                               
  |              |  +----- Lortiella             
  |              +--+                            
  |                 |  +-- Alathyria             
  |                 +--+                         
  |                    +-- Velesunio_angasi      
  |                                              
  |  +-------------------- Diplodon_suavidicus__1
  |  |                                           
  +--+                 +-- Triplodon             
     |  +--------------+                         
     |  |              +-- Prisodon              
     +--+                                        
        |  +-------------- Castalia_stevensi     
        |  |                                     
        +--+           +-- Callonaia             
           |  +--------+                         
           |  |        +-- Castalia_ambigua__1   
           +--+                                  
              |  +-------- Castalia_ambigua__4   
              +--+                               
                 |  +----- Castalia_ambigua__2   
                 +--+                            
                    |  +-- Castaliella           
                    +--+                         
                       +-- Castalia_ambigua__3

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
33131970020
  +----- Hyridella_australis **
--+                         
  |  +-- Hyridella_depressa ** 
  +--+                      
     +-- Hyridella_drapeta **

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
33131950020
  +----- Hyridella_australis **
--+                         
  |  +-- Hyridella_depressa ** 
  +--+                      
     +-- Hyridella_drapeta **

Graf et al., 2015 [Publ 3619], fig. 1

iTtTbiriSoTroTroSiD
3423171119
     +-- Hyridella_depressa ** 
  +--+                      
  |  +-- Hyridella_drapeta **  
--+                         
  |  +-- Hyridella_australis **
  +--+                      
     +-- Cucumerunio

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
33131970020
  +----- Hyridella_australis **
--+                         
  |  +-- Hyridella_depressa ** 
  +--+                      
     +-- Hyridella_drapeta **

Santos-Neto et al., 2016 [Publ 3739], fig. 4

iTtTbiriSoTroTroSiD
313113171116
              +-------- Westralunio        
  +-----------+                            
  |           |  +----- Diplodon           
  |           +--+                         
  |              |  +-- Hyridella_drapeta **  
--+              +--+                      
  |                 +-- Hyridella_depressa ** 
  |                                        
  |  +----------------- Echyridella        
  |  |                                     
  +--+              +-- Cucumerunio        
     |  +-----------+                      
     |  |           +-- Hyridella_australis **
     +--+                                  
        |  +----------- Velesunio_ambiguus 
        +--+                               
           |  +-------- Alathyria          
           +--+                            
              |  +----- Velesunio_angasi   
              +--+                         
                 |  +-- Lortiella          
                 +--+                      
                    +-- Microdontia

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

iTtTbiriSoTroTroSiD
3182036281117.5
                          +----- Diplodon             
  +-----------------------+                           
  |                       |  +-- Hyridella_depressa **   
  |                       +--+                        
  |                          +-- Hyridella_drapeta **    
--+                                                   
  |                          +-- Cucumerunio          
  |  +-----------------------+                        
  |  |                       +-- Hyridella_australis **  
  |  |                                                
  +--+  +----------------------- Echyridella          
     |  |                                             
     |  |              +-------- Velesunio__1         
     +--+  +-----------+                              
        |  |           |  +----- Alathyria_jacksoni__1
        |  |           +--+                           
        |  |              |  +-- Velesunio_ambiguus   
        +--+              +--+                        
           |                 +-- Alathyria_profuga    
           |                                          
           |  +----------------- Velesunio__3         
           |  |                                       
           +--+  +-------------- Microdontia          
              |  |                                    
              +--+           +-- Lortiella            
                 |  +--------+                        
                 |  |        +-- Velesunio_angasi     
                 +--+                                 
                    |  +-------- Alathyria_pertexta__2
                    +--+                              
                       |  +----- Velesunio__2         
                       +--+                           
                          |  +-- Alathyria_jacksoni__2
                          +--+                        
                             +-- Alathyria_pertexta__1

Graf & Ó Foighil, 2000 [Publ 2649], fig. 1_2

iTtTbiriSoTroTroSiD
230390023
  +-------- Hyridella_australis__2 **
--+                               
  |  +----- Hyridella_australis__1 **
  +--+                            
     |  +-- Echyridella           
     +--+                         
        +-- Hyridella_depressa **

Walker et al., 2006 [Publ 2666], fig. 6

iTtTbiriSoTroTroSiD
2802340020.5
  +-------------- Hyridella_depressa ** 
  |                                  
  +-------------- Hyridella_australis **
--+                                  
  |  +----------- Echyridella        
  |  |                               
  |  |        +-- Velesunio_ambiguus 
  +--+  +-----+                      
     |  |     +-- Velesunio__3       
     |  |                            
     +--+  +----- Velesunio__2       
        |  |                         
        |  |  +-- Alathyria          
        +--+--+                      
           |  +-- Velesunio__1       
           |                         
           |  +-- Velesunio_angasi   
           +--+                      
              +-- Lortiella

Baker et al., 2003 [Publ 2690], fig. 5

iTtTbiriSoTroTroSiD
270260023
           +-- Hyridella_australis **
  +--------+                      
  |        +-- Diplodon           
--+                               
  |  +-------- Hyridella_depressa ** 
  +--+                            
     |  +----- Velesunio          
     +--+                         
        |  +-- Echyridella        
        +--+                      
           +-- Castalia

Hoeh et al., 2009 [Publ 2716], fig. 5

iTtTbiriSoTroTroSiD
22024400--
  +-- Hyridella_depressa ** 
--+                      
  +-- Hyridella_australis **

Hoeh et al., 2009 [Publ 2716], fig. 3

iTtTbiriSoTroTroSiD
22024500--
  +-- Hyridella_depressa ** 
--+                      
  +-- Hyridella_australis **

Baker et al., 2004 [Publ 2685], fig. 2-3

iTtTbiriSoTroTroSiD
230270022
  +----- Hyridella_depressa ** 
--+                         
  |  +-- Hyridella_australis **
  +--+                      
     +-- Diplodon

Hoeh et al., 2009 [Publ 2716], fig. 4

iTtTbiriSoTroTroSiD
22024400--
  +-- Hyridella_depressa ** 
--+                      
  +-- Hyridella_australis **

Graf & Cummings, 2006 [Publ 1968], fig. 2

iTtTbiriSoTroTroSiD
22024500--
  +-- Hyridella_depressa ** 
--+                      
  +-- Hyridella_australis **

Sonowal et al., 2021 [Publ 4221], fig. 3b

iTtTbiriSoTroTroSiD
11011400--
-- Hyridella_australis **

Walker et al., 2006 [Publ 2666], fig. 3

iTtTbiriSoTroTroSiD
11011400--
-- Hyridella **

Whelan et al., 2011 [Publ 3380], fig. 1-ML

iTtTbiriSoTroTroSiD
11013500--
-- Hyridella_australis **

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

iTtTbiriSoTroTroSiD
11013900--
-- Hyridella_australis **

Lopes-Lima et al., 2018 [Publ 3829], fig. 1

iTtTbiriSoTroTroSiD
11012700--
-- Hyridella_australis **

Sano et al., 2020 [Publ 4138], fig. 3

iTtTbiriSoTroTroSiD
11016700--
-- Hyridella_australis **

Walker et al., 2006 [Publ 2666], fig. 4

iTtTbiriSoTroTroSiD
11011400--
-- Hyridella **

Huang et al., 2018 [Publ 3828], fig. 3_5

iTtTbiriSoTroTroSiD
11012800--
-- Hyridella_australis **

Graf, 2000 [Publ 2647], fig. 1

iTtTbiriSoTroTroSiD
11011600--
-- Hyridella **

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iTtTbiriSoTroTroSiD
110112100--
-- Hyridella_depressa **

Sonowal et al., 2021 [Publ 4221], fig. 3a

iTtTbiriSoTroTroSiD
11011400--
-- Hyridella_australis **

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B

iTtTbiriSoTroTroSiD
11012900--
-- Hyridella_depressa **

Marshall et al., 2014 [Publ 3593], fig. 1

iTtTbiriSoTroTroSiD
11011311--
  +-- Cucumerunio        
--+                      
  +-- Hyridella_australis **

Graf, 2002 [Publ 2651], fig. 2

iTtTbiriSoTroTroSiD
11011700--
-- Hyridella_depressa **

Sano et al., 2020 [Publ 4138], fig. 4

iTtTbiriSoTroTroSiD
11016700--
-- Hyridella_australis **

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iTtTbiriSoTroTroSiD
11015600--
-- Hyridella_australis **

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A

iTtTbiriSoTroTroSiD
11012900--
-- Hyridella_depressa **

Combosch et al., 2017 [Publ 3806], fig. 1

iTtTbiriSoTroTroSiD
11011400--
-- Hyridella_australis **

Wu et al., 2020 [Publ 4121], fig. 2

iTtTbiriSoTroTroSiD
11013300--
-- Hyridella_australis **

Combosch et al., 2017 [Publ 3806], fig. 2

iTtTbiriSoTroTroSiD
11011400--
-- Hyridella_australis **

Walker et al., 2006 [Publ 2666], fig. 2

iTtTbiriSoTroTroSiD
11011400--
-- Hyridella **

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iTtTbiriSoTroTroSiD
11015600--
-- Hyridella_australis **

Lopes-Lima et al., 2018 [Publ 3829], fig. 3

iTtTbiriSoTroTroSiD
11012700--
-- Hyridella_australis **

Whelan et al., 2011 [Publ 3380], fig. 1-MP

iTtTbiriSoTroTroSiD
11013500--
-- Hyridella_australis **

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iTtTbiriSoTroTroSiD
11017900--
-- Hyridella_depressa **

Sonowal et al., 2021 [Publ 4221], fig. 4

iTtTbiriSoTroTroSiD
11011500--
-- Hyridella_australis **

Chen et al., 2025 [Publ 4455], fig. 1

iTtTbiriSoTroTroSiD
11012500--
-- Hyridella_australis **

Wu et al., 2020 [Publ 4121], fig. S1

iTtTbiriSoTroTroSiD
11013300--
-- Hyridella_australis **

Sano et al., 2020 [Publ 4138], fig. 2

iTtTbiriSoTroTroSiD
11016700--
-- Hyridella_australis **
 
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