MUSSELpdb | valid sp. Anodontites trigona specimens

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Page last updated
Thu 17 July 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

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species Anodontites trigona (Spix in Wagner, 1827): specimens
19 available nominal species synonyms (29 total synonyms)

Recent Taxonomic Opinions

Anodontites trigonus — Graf & Cummings (2007), Hoeh et al. (2009), Bogan (2010), IUCN (2011), Zieritz et al. (2015)
Anodontites (s.s.) trigonus — Rumi et al. (2008)
— Petit & Coan (2008)
Anodontites (s.s.) trigonus gorginae [sic] — Rumi et al. (2008)
Anodontites (s.s.) puelchanus — Rumi et al. (2008), Pereira et al. (2014), Torres et al. (2018)
Anodontites trigona — Lasso et al. (2009), Graf & Cummings (2021), Breure et al. (2022), Torres et al. (2024)
Anodontites elongatus — Mansur et al. (2020)
Anodontites puelchanus — Mansur et al. (2020)
Anodontites puelchana — Torres et al. (2024)

Range

Patagonia and the Paraná Basin, Argentina, north through the Amazon to Colombia and Ecuador.

Map of Georeferenced Specimens

species map

UF_270542
Probably Paraná

Argentina

SMF_11949 Argentinien	USNM_607081 Arroyo Malabrigo, trib. Rio San Javier, Santa Fe, Argentina	USNM_607081 Arroyo Malabrigo, trib. Rio San Javier, Santa Fe, Argentina
USNM_86604 Corrientes R. S.A.	USNM_86604 Corrientes R. S.A.	MCZ_111731 Corrientes, Argentina
SMF_11951 Corrientes, Argentinien	USNM_86518 Corrientes, S. Am.	AMS_126854 Guadalupe Lagoon, Parana River, Santa Fe, Argentina
FMNH_216436 La Matanza, Prov. Salta, Argentina	CM_MP_83 Laguna del Arroyo Malabrigo, Romang, Santa Fe, Argentina	CM_MP_84 Laguna del Arroyo Malabrigo, Romang, Santa Fe, Argentina
AMNH_100112 Laguna Guadalupe, Dpto. La Capital, Prov, Santa Fe, Argentina	CM_66.101 Laguna Guadalupe, Santa Fe, Argentina	FMNH_216472 Laguna Oca, Prov. Formosa, Argentina
FMNH_216471 Laguna, near Airport Oran, Prov. of Salta, Argentina	SMF_3822 Maloya Sumpfe, Prov Corrientes, Argentinien	SMF_3822 Maloya Sumpfe, Prov Corrientes, Argentinien
FMNH_44560 Northern Santa Fe Prov., Argentina	USNM_86605 Rio Corrientes, South America	USNM_86605 Rio Corrientes, South America
CM_61.5828 Rio Limay ?, Patagonia, Argentina	INHS_85705 Rio Parana (Río de la Plata Dr.) side channel, 17.3 mi SSW Esquina, Corrientes, Argentina, South America, -30.24806, -59.60724	SMF_11945 Rio Paraná, bei Godoy, Corrientes, Argentinien
SMF_11945 Rio Paraná, bei Godoy, Corrientes, Argentinien	SMF_4053 Rio Parana, Prov. Corrientes, Argentinien	SMF_4053 Rio Parana, Prov. Corrientes, Argentinien

Bolivia

UF_93273 15.8 km SSW of San Rafael, Velasco Province, Santa Cruz Departemento, Bolivia 60 30’ W, 16 56’ S	UF_93311 16 km S of San Rafael, Velasco Province, Santa Cruz Departemento, Bolivia	UF_93270 3.5 km N of San Ramon, Santa Cruz Departemento, Bolivia
USNM_361150 Beni River, Rurrenebague, Bolivia, drains into the Madeira which drains into the Amazon	USNM_361150 Beni River, Rurrenebague, Bolivia, drains into the Madeira which drains into the Amazon	USNM_361150 Beni River, Rurrenebague, Bolivia, drains into the Madeira which drains into the Amazon
ZMB_MP_0178 Bolivia	INHS_18223 borrow pit (Rio Amazon - Rio Apere Dr.), 3 km E Estacion Biologica del Beni on rd. to San Ignacio, Beni, Bolivia	UF_93274 Comunidad La Esperanza, circa 40 km N of San Jose, Velasco Province, Santa Cruz Departemento, Bolivia
FMNH_21482 Ibares, Bolivia	FMNH_45716 Puerto Suarez, Departmento de Santa Fe, Bolivia, Paraguay drainage	UF_93373 Rio Ibare, 17 km SSE San Lorenzo, Marbon Province, Beni Departemento, Bolivia
UF_93360 Rio Ibare, Puerto Almecen, Cercado Province, Beni Departemento, Bolivia	UF_93362 Rio Ibare, Puerto Almecen, Cercado Province, Beni Departemento, Bolivia	UF_93362 Rio Ibare, Puerto Almecen, Cercado Province, Beni Departemento, Bolivia
UF_93363 Rio Ibare, Puerto Almecen, Cercado Province, Beni Departemento, Bolivia	UF_93364 Rio Ibare, Puerto Almecen, Cercado Province, Beni Departemento, Bolivia	FMNH_21483 Rio Ibari, Bolivia
AMNH_186421 Rio Itenez (=Guaporé), Bani, 4 km above Costa Marques, Brazil, Bolivia	FMNH_45717 Rio Mamore, Trinidad, Departmento de Benin, Bolivia	INHS_17844 Rio Matos (Rio Amazon - Rio Apere Dr.), 6 km E Estación Biologica del Beni on rd. to San Ignacio, Beni, Bolivia
INHS_17845 Rio Matos (Rio Amazon - Rio Apere Dr.), 6 km E Estación Biologica del Beni on rd. to San Ignacio, Beni, Bolivia	UF_93346 Rio Matos, 25 km E, 12 km S San Borja, Yacuma Province, Beni Departemento, Bolivia	UF_93346 Rio Matos, 25 km E, 12 km S San Borja, Yacuma Province, Beni Departemento, Bolivia
UF_93349 Rio Matos, 25 km E, 12 km S San Borja, Yacuma Province, Beni Departemento, Bolivia	UF_93349 Rio Matos, 25 km E, 12 km S San Borja, Yacuma Province, Beni Departemento, Bolivia	INHS_18226 Rio Moseruna (Rio Amazon - Rio Apere Dr.), 46 km E Estacion Biologica del Beni, on rd. to San Ignacio, Beni, Bolivia
UF_93377 Rio Parapeti, Estancia, Isla Verde, Chiquitos Province, Santa Cruz Departmento, Bolivia	UF_93377 Rio Parapeti, Estancia, Isla Verde, Chiquitos Province, Santa Cruz Departmento, Bolivia	UF_93296 Rio Tucavaca, Candelaria, Chiquitos Province, Santa Cruz Departemento, Bolivia
UF_93296 Rio Tucavaca, Candelaria, Chiquitos Province, Santa Cruz Departemento, Bolivia	UF_93308 Rio Tucavaca, Candelaria, Chiquitos Province, Santa Cruz Departemento, Bolivia	UF_93310 Rio Tucavaca, Candelaria, Chiquitos Province, Santa Cruz Departemento, Bolivia
UF_93316 Rio Uroguaito, 39 km E Concepcion, Nuflo de Chavez Province, Santa Cruz Departemento, Bolivia	UF_93336 Rio Zapacos, 3 km W Asencion, Nuflo de Chavez Province, Santa Cruz Departemento, Bolivia	MNHN_MP_0671 Santa Anna de Chiquitos, Brazil
UF_93326 Small Arroyo, Circa 10 km W of Warnes, Andres Ibanez Province, Santa Cruz Departemento, Bolivia	UMMZ_112644 Tapacany River, Tarbusia, Bolivia	UMMZ_112644 Tapacany River, Tarbusia, Bolivia

UF_93346B. Rio Matos, 25 km E, 12 km S San Borja, Yacuma Province, Beni Departemento, Bolivia

Brazil

SMF_11943 A.d. Amazonas Fl.	CM_61.11131 Amazon	MCZ_MP.171 Amazon
USNM_127191 Amazon	USNM_127191 Amazon	AMNH_30865 Amazon River
ANSP_125438 Amazon River	DMNH_81458 Amazon River	FMNH_115948 Amazon River
FMNH_21479 Amazon River	FMNH_21493 Amazon River	USNM_86731 Amazon River
USNM_86731 Amazon River	USNM_86731 Amazon River	UF_93181 Amazon River, Brazil
UMMZ_112431 Amazon River, Brazil	UMMZ_112431 Amazon River, Brazil	UMMZ_112617 Amazon River, Brazil
UMMZ_112617 Amazon River, Brazil	UMMZ_112394 Amazon River, South America	UMMZ_112394 Amazon River, South America
SMF_11934 Amazonas	SMF_11934 Amazonas	INHS_79223 at mouth of rio Jarauçu (Aquiqui)
ZMB_40678 Bakairi, Kamayura	INHS_78833 beach on right bank ca. 60 km north of Vitória do Xingu	INHS_79198 beach on right bank, campsite 5 and vicinity, ca. 3.5 km southeast of Porto do Moz
MNHN_MP_0670 Brasil	ANSP_125439 Brazil	ANSP_125439 Brazil
ANSP_41853 Brazil	MCZ_MP.164 Brazil	UMMZ_112392 Brazil
UMMZ_112392 Brazil	UMMZ_112432 Brazil	UMMZ_112432 Brazil
UMMZ_112626 Brazil	UMMZ_112626 Brazil	USNM_126492 Brazil
USNM_126492 Brazil	SMF_168140 Fluss Amazon	CM_61.5827 headwaters of Rio Paraguay, Santa Rita, Matto Grosso, Brazil
CM_61.5831 headwaters of Rio Paraguay, Santa Rita, Matto Grosso, Brazil	CM_61.5832 headwaters of the Rio Paraguay, Santa Rita, Matto Grosso, Brazil	FMNH_54628 Igarapé Aniba, Amazonas, Brazil
SMF_11935 Itaituba am Amazonas	MCZ_MP.165 Javary, Brazil	AMNH_139058 Juruá River, Juao Pessoa, Brazil
DMNH_118526 Juruá River, Juao Pessoa, Brazil	DMNH_146473 Juruá River, Juao Pessoa, Brazil	DMNH_56336 Juruá River, Juao Pessoa, Brazil
MCZ_91165 Juruá River, Juao Pessoa, Brazil	SBMHN_367467 Juruá River, Juao Pessoa, Brazil	SBMHN_367477 Juruá River, Juao Pessoa, Brazil
USNM_706865 Juruá River, Juao Pessoa, Brazil	USNM_706865 Juruá River, Juao Pessoa, Brazil	USNM_853432 Juruá River, Juao Pessoa, Brazil
FMNH_30447 mouth of Rio Curucumba, into Lake Mamuru, near Obidos, Pará, Brazil	INHS_78894 off island opposite of mouth of igarapé Tucuruí, ca. 4 km northeast of Vitória do Xingu	INHS_78879 off right bank, point between campsite 5 and Porto do Moz
FMNH_126609 Rio Araguaia, Barra do Tapirape, Mato Grosso, Brazil; 10°50’S, 50°40’	ZMB_37269 Rio de Miranda, Mattogrosso	ZMB_37269 Rio de Miranda, Mattogrosso
FMNH_25885 Rio Juruá, near João Pessoa, Amazonas, Brazil	FMNH_54603 Rio Juruá, near João Pessoa, Amazonas, Brazil	MCZ_297399 Rio Maica Faz, Taparinha, Santarem, Pará, Brazil
ANSP_158784 Rio Paraguay, Descalvados, Matto Grosso, Brazil	CM_MP_77 Rio Paraguay, Sao Luis de Caceres, Matto Grosso, Brazil	SMF_11947 Rio Paraguay, Staat Matto Grosso, Brasilien
UMMZ_4392 Rio Purús, Brazil	UMMZ_4392 Rio Purús, Brazil	CM_61.5829 Rio Tapajós, Santarem, Pará, Brazil
CM_MP_75 Rio Tapajós, Santarem, Pará, Brazil	FMNH_112110 Rio Tapirape, affl. of Rio Araguaia, Mato Grosso, Brazil	ZMB_35612 Rio Tocantins
INHS_45937 Rio Xingu, ~7 km N Senador José Porfîrio, Pará, Brazil	INHS_45937 Rio Xingu, ~7 km N Senador José Porfîrio, Pará, Brazil	SBMHN_367481 S. Joai, Matto Grosso
UMMZ_112397 Southern Brazil	UMMZ_112397 Southern Brazil	UMMZ_112397 Southern Brazil
UMMZ_112438 trib. to Amazon River, 100 mi from Parà, Brazil	UMMZ_112438 trib. to Amazon River, 100 mi from Parà, Brazil	UF_270563 tributary of The Amazon River
USNM_86724 Upper Amazon, Brazil	USNM_86724 Upper Amazon, Brazil	FMNH_43783 Xingu Alto, Jacare, Mato Grosso, Brazil

PUCRS_MCP_6119. Brazil: Acre: Marechal Taumaturgo: (Amazon - Juruá): rio Tejo: -8.94684 -72.59079

INPA_1577. Brazil: Amapá: (Amazon): rio Amazonas: prox. Forte Macapá 00° 1' 47" N, 51° 2' 53'' W: 0.02972, -51.04806

PUCRS_MCP_8538. Brazil: Amazonas: Humaitá: (Amazon - Madeira): rio Madeira: -7.51509 -63.01646

INPA_909. Brazil: Amazonas: Iranduba: (Amazon - Negro): rio Negro: Catalão, entre o flutuante do INPA e a margem do rio Negro: -3.14898 -59.91798

INPA_1451. Brazil: Amazonas: Iranduba: (Amazon - Negro): rio Negro: Catalão, local 1, caminho entre flutuante do INPA e margem direita do rio Negro: -3.14898 -59.91798

INPA_1124. Brazil: Amazonas: Iranduba: (Amazon - Negro): rio Negro: Catalão, margem direita do paraná que liga o lago Catalão até o rio Negro: -3.14833 -59.91833

INPA_1325. Brazil: Amazonas: Iranduba: (Amazon - Negro): rio Negro: Catalão, praia na boca do paraná do lago Catalão: -3.14833 -59.91833

PUCRS_MCP_9081. Brazil: Goias: São Miguel do Araguaia: (Tocantins - Araguaia)

INPA_491. Brazil: Mato Grosso do Sul: Aquidauana: (Río de la Plata - Paraná - Miranda): rio Negro: na fazenda São Pedro

MHNCI_2525. Brazil: Mato Grosso do Sul: Corumbá: (Río de la Plata - Paraná - Paraguay)

PUCRS_MCP_6151. Brazil: Pará: (Amazon - Tapajós): rio Tapajós: Praia do Rio Tapajós

PUCRS_MCP_8537. Brazil: Pará: Afuá: (Amazon): Ilha do Chagas, afluente da Ilha do Pará: -1.99813 -54.93062

INPA_1521. Brazil: Para: Rio Xingu, ilha do Kaituka

PUCRS_MCP_8528. Brazil: Pará: Santarém: (Tocantins): Ponta das Pedras: -1.26962 -48.79856

INPA_630. Brazil: Pará: Terra Santa: (Amazon lake): Lago Piraruacá: -2.03830 -56.30357

INPA_848. Brazil: Pará: Terra Santa: (Amazon lake): Lago Piraruacá: -2.03830 -56.30357

INPA_1360. Brazil: Tocantins: Caseara: (Tocantins - Araguaia): rio Araguaia: -9.23073 -49.96436

Colombia

ANSP_164552 Cali, Colombia	CM_61.1545 canal, 40 mi S Cartegena, Colombia	MCZ_MP.180 Cartagena
UMMZ_112408 Cartagena, Colombia, South America	UMMZ_112408 Cartagena, Colombia, South America	USNM_26100 Cartagena, So. America
AMNH_30872 Carthagena	USNM_86684 Carthegena, U.S. Colombia	USNM_86684 Carthegena, U.S. Colombia
USNM_86684A Carthegena, U.S. Colombia	CM_73969 Cesar River, at La Brillantina, Colombia	CM_73971 Cesar River, at La Brillantina, Colombia
FMNH_179991 Cga. de Repelon, between Luruaco & Santa Rosa, towards Canal del Dique, Departmento de Bolivar, Colombia	UMMZ_112434 Colombia, South America	UMMZ_112434 Colombia, South America
SMF_11940 Frontino Kolumbien	SMF_83378 Frontino, New Granada	UF_29685 Hacienda Atenas, Florida (nr. Palmira), Valle del Cauca, Colombia
USNM_809472 Hacienda Atenas, Florida Valle, Colombia	MCZ_157416 Hacienda La Maria, Municipo de Buga, Departmento de Valle del Cauca, Colombia	ZMB_35610 Mocoa
USNM_599515 near Codazzi, Dept. Magdalena, Colombia	USNM_599515 near Codazzi, Dept. Magdalena, Colombia	ANSP_125433 Ocana, N. Grenada
ZMB_MP_0167 Puerto Nacional, New Grenada	USNM_543460 Quebrada Boquete, 5 km N of Becerril, Dept. Magdalena, Colombia	USNM_543481 Quebrada Hondo, tributary to Rio Rancheria, Dept. Magdalena, Colombia
USNM_543481 Quebrada Hondo, tributary to Rio Rancheria, Dept. Magdalena, Colombia	ANSP_166126 Quebrada Zarzal, North of El Centro	FMNH_114041 Region de la Laguna “La Raya” Municipio Achí, bajo Rio Cauca, Departmento de Bolivar, Colombia
FMNH_114155 Region de la Laguna “La Raya” Municipio Achí, bajo Rio Cauca, Departmento de Bolivar, Colombia	USNM_364263 Rio Canalete, 0.5 mi above La Union, Dept. Bolivar, U.S. Colombia	USNM_364263 Rio Canalete, 0.5 mi above La Union, Dept. Bolivar, U.S. Colombia
USNM_517848 Rio Cauca, Antioquia, Colombia	ZMB_MP_0176 Rio Choco, Nueva Grenada	MNHN_MP_0356 Rio Magdalena
MNHN_MP_0356 Rio Magdalena	MCZ_134967 Rio Putumayo, Pto. Asis, Colombia	ANSP_171279 Rio Sinu, near Jaraguay, Colombia
ANSP_171279 Rio Sinu, near Jaraguay, Colombia	USNM_590652 Simiti, near Rio Magdalena, D. Bolivar, Colombia	USNM_590652 Simiti, near Rio Magdalena, D. Bolivar, Colombia

INPA_1406. Colombia: Vichada:(Orinoco - Guaviare): rio Guaviare: Laguna Paluame, margem esquerda rio Guaviare, próximo confluência Guaviare-Inírida: 4.05008 -67.75001

Costa Rica

ANSP_41844 Costa Rica	BMNH_79-7-22-39-42 Costa Rica	UMMZ_112410 Costa Rica
UMMZ_122410 Costa Rica	USNM_382688 L. Tortuguero, Nivel del Mar, Costa Rica	ZMB_42487 Lagune d Bebero, Costa Rica
UF_270337 Limon, Costa Rica	MCZ_294790 Rio Saxaola, 2 km SE Bribri, SE Costa Rica	ANSP_171330 Rio Temesque, Filadelphia, Costa Rica
ANSP_171330 Rio Temesque, Filadelphia, Costa Rica

Ecuador

DMNH_173915 Ecuador	ZMB_MP_0179 Ecuador	UMMZ_112411 Ecuador, South America
UMMZ_112411 Ecuador, South America	UMMZ_112411 Ecuador, South America	USNM_127201 Equador
USNM_127201 Equador	IRSNB_MP_0917 Equatuer	IRSNB_MP_0906 Laguna formen por les rivieres Napo & Coca, laguna de Samona, a 18 minutes de Coca, Ecuador
MNHN_MP_0413 lagunas du Napo a in la Coca, Ecuador	ANSP_125432 Napo River, Ecuador	ANSP_125432 Napo River, Ecuador
ZMB_50834 Putumayo, Ecuador	MCZ_293524 Rio Napo, downstream from Coca at “Lake Taracoa”, Ecuador	FMNH_21480 Rio Napo, Ecuador
IRSNB_MP_0950 Rio Napo, Ecuador	MNHN_MP_0355 Sarayacu, Equateur

Guatemala

UMMZ_112639 Rio Conchins, Quirigua, Guatemala	UMMZ_112639 Rio Conchins, Quirigua, Guatemala	CM_61.6198 Rio Conchins, Quirigua, Maya Farm, Guatemala
ZMB_47547 Rio Michatoyac, Paso San Antonio, Guatemala

Honduras

UF_221537
Rio Choluteca, 3 km NNW of San Juan De Flores, Francisco Morazón Department, Honduras

UF_220956
Rio Choluteca, 3 km NNW of San Juan Del Flores, Francisco Morazón Department, Honduras

Panama

MCZ_MP.202
3 mi E Agua Dulce, in irrigation ditch on Sr. Mendez Panama

INHS_21692
Rio Membrillo, ca. 5 km upstream Sinai, Darien, Panama

Paraguay

MCZ_62480 Arojo Dona Juana, Villarica, Paraguay	FMNH_21478 Arroyo Mbobo, Villarrica, Paraguay	UMMZ_219678 Asuncion Bay, Asuncion, Paraguay
SMF_11946 Ob. Rio Paraná bei Puerto Bertoni, Paraguay	ZMB_MP_0080 Paraguay	ZMB_MP_0158 Paraguay
USNM_348736 Paraguay R., Asuncion, Paraguay	UMMZ_112435 Paraguay, South America	UMMZ_112435 Paraguay, South America
UMMZ_112435 Paraguay, South America	SMF_14078 Rio Paraguay	SMF_5309 Rio Paraguay, 22° S Br.
SMF_11944 Rio Paraguay, bei Asuncion, Paraguay	SMF_641 Rio Paraguay, bei Concepcion, Paraguay	SMF_11948 Rio Paraguay, nahe Mündung des Rio Apa
FMNH_120662 Rio Paraguay, Paraguay	DMNH_173141 Rio Paraná, Paraguay	FMNH_50769 Rio Pilcomayo, 3 mi N Linares, Paraguayan Chaco
UMMZ_219658 Rio Tebicuary, between Florida and Reciefe, Paraguay	CM_61.5826 swamp of Lambare, Asuncion, Paraguay	SMF_11952 Villarica, Arroyo Tebicuary, Paraguay
BMNH_1934-7-3-22-23 Villarica, Paraguay	AMNH_88043 Villarrica, Paraguay

Peru

AMNH_89109 along water line at Tierra Blanca, Peru	AMNH_89111_2 along water line at Tierra Blanca, Peru	FMNH_120331 Huacabamba, Peru
SMF_11941 Huagabamba Peru, Amzaonas Geb.	ZMB_40556 Huagamba	FMNH_21481 Huayabama, Peru
UMMZ_112399 Huayabamba, Peru	UMMZ_112399 Huayabamba, Peru	AMNH_146580 Huazabamba, East Peru
ANSP_232407 Huazabamba, East Peru	SMF_168137 Huazabamba, Peru	USNM_339714 Huazabamba, Peru
SMF_162031 Laguna Yarinacocha, bei Pucallpa, O. Peru	FMNH_216446 Laguna Yarinacocha, Pucallpa, Peru	FMNH_47103 Laguna Yarinacocha, Pucallpa, Peru
AMNH_89223 Lake Yarina-Cocha, Pucallpa, Peru	ANSP_125431 Maranon River, Amazon R.	USNM_86606 Maranon, Brazil
MCZ_MP.168 Pachitea River, upstream end of Chonta Isla, Chacra, Peru	ANSP_125440 Peru	CM_61.59 Peru
CM_MP_78 Peru	UF_270535 Peru	UMMZ_112400 Peru, South America
UMMZ_112400 Peru, South America	UMMZ_112439 Peru, South America	UMMZ_112439 Peru, South America
MUSM_MP_004 Peru: Pucallpa, Yarinacocha	DMNH_195828 Rio Calleria, Colonia Calleria, 15 km from Ucayali, Loreto, Peru	FMNH_114172 Rio Calleria, Colonia Calleria, 15 km from Ucayali, Loreto, Peru
MCZ_266558 Rio Calleria, Colonia, Calleria, 15 km from Ucayali, Loreto, Peru	SMF_11938 Rio Estacamento, Peru	MCZ_17036 Rio Huallaga, Peru
SMF_11942 Rio Huallaga, Peru, Amazonas Geb.	INHS_23406 Rio Marañon, 1 mi S Nauta, in large shallow oxbow lake S of southern braid of main channel, Loreto, Peru	UMMZ_112437 Rio Maranon, Peru, South America
UMMZ_112437 Rio Maranon, Peru, South America	FMNH_216668 Rio Samiria, N. trib of the Amazon River, between Rio Huallaga and Ucayali, Peru	AMNH_89123 Rio Ucayali, Peru
UMMZ_112433 Rio Ucayali, Peru	UMMZ_112433 Rio Ucayali, Peru	UMMZ_112433 Rio Ucayali, Peru
UMMZ_112433 Rio Ucayali, Peru	SMF_11936 Rio Ucayali, Peru, Amazonas Geb.	ZMB_35611 Sanango Cocha, Hachual
ZMB_37655 Ucayali	ZMB_25666 Ucayali, E. Peru	ZMB_69757 Yarina Cocha am Ucayali

unknown

ANSP_125434 [no locality available - South America]	ANSP_85115 [no locality available - South America]	CM_1279 [no locality available - South America]
FMNH_21495 [no locality available - South America]	UMMZ_4390 [no locality available - South America]	UMMZ_4390 [no locality available - South America]
UMMZ_4390 [no locality available - South America]	ZMB_MP_0151 [no locality available - South America]	ZMB_MP_0153 [no locality available - South America]
SMF_11939 Ganges [no - South America]	BMNH_MP_292 India ? [no - South America]	SMF_11937 Ob. Amazonas
SMF_11950 Ob. Amazonas	ANSP_125457 S. Amer.	ANSP_125457 S. Amer.
UMMZ_110137 Siliguri, East Bengal, India [No - South America]	ANSP_41841 So. America	ANSP_41858 So. America
MCZ_MP.163 South Amercia	ANSP_125448 South America	MCZ_2064 South America
MCZ_MP.151 South America	MCZ_MP.167 South America	UMMZ_23177 South America
UMMZ_23177 South America	UMMZ_23178 South America	UMMZ_23178 South America
SMF_11933 Unterer [lower] Amazonas	SMF_11933 Unterer [lower] Amazonas	ANSP_125430 Upper Amazon R.
ANSP_125430 Upper Amazon R.	AMNH_30864 Upper Amazon River	DMNH_78643 Upper Amazon River

Venezuela

ZMB_32664. Maracaibo