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Page last updated
Thu 17 July 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Bivalvia | Unionoida | Unionoidea | Unionidae | Unioninae | Unionini

genus Middendorffinaia Moskvicheva & Starobogatov, 1973: cladomics
     1 Recent valid species (1 total species)

Cladomics Summary

number of species = 1 (1 total)
analyzed in 18 cladograms in 11 publications
maximum number of terminals in an analysis = 1 (mean = 1, median = 1)

Dai et al., 2025 [Publ 4440], fig. 1

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
1101442110--
  +----------------------- Globunio                  
  |                                                  
--+  +-------------------- Aculamprotula             
  |  |                                               
  +--+  +----------------- Middendorffinaia_mongolica **
     |  |                                            
     +--+  +-------------- Nodularia                 
        |  |                                         
        +--+  +----------- Cuneopsis                 
           |  |                                      
           |  |        +-- Diaurora                  
           +--+--------+                             
              |        +-- Schistodesmus             
              |                                      
              |  +-------- Pseudocuneopsis           
              +--+                                   
                 |  +----- Unio                      
                 +--+                                
                    |  +-- Tchangsinaia              
                    +--+                             
                       +-- Inversiunio

Lopes-Lima et al., 2020 [Publ 4080], fig. 13

iTtTbiriSoTroTroSiD
1101971--
  +-- Middendorffinaia_mongolica **
--+                             
  +-- Nodularia

Sayenko et al., 2021 [Publ 4189], fig. 6

iTtTbiriSoTroTroSiD
110112123--
  +-------- Unio                      
--+                                   
  |  +----- Inversiunio               
  +--+                                
     |  +-- Middendorffinaia_mongolica **
     +--+                             
        +-- Nodularia

Hou et al., 2024 [Publ 4400], fig. 4

iTtTbiriSoTroTroSiD
110175487--
  +-------------------- Aculamprotula             
--+                                               
  |  +----------------- Unio                      
  +--+                                            
     |  +-------------- Middendorffinaia_mongolica **
     +--+                                         
        |  +----------- Pseudocuneopsis           
        +--+                                      
           |  +-------- Nodularia                 
           +--+                                   
              |  +----- Schistodesmus             
              +--+                                
                 |  +-- Tchangsinaia              
                 +--+                             
                    +-- Cuneopsis

Hou et al., 2025 [Publ 4401], fig. 4

iTtTbiriSoTroTroSiD
110127255--
  +-------------- Middendorffinaia_mongolica **
--+                                         
  |  +----------- Pseudocuneopsis           
  +--+                                      
     |  +-------- Nodularia                 
     +--+                                   
        |  +----- Schistodesmus             
        +--+                                
           |  +-- Tchangsinaia              
           +--+                             
              +-- Cuneopsis

Dai et al., 2025 [Publ 4440], fig. 2

iTtTbiriSoTroTroSiD
1101442110--
  +-------------------- Globunio                  
  |                                               
--+  +----------------- Aculamprotula             
  |  |                                            
  |  |              +-- Middendorffinaia_mongolica **
  +--+  +-----------+                             
     |  |           +-- Nodularia                 
     |  |                                         
     +--+  +----------- Cuneopsis                 
        |  |                                      
        |  |        +-- Diaurora                  
        +--+--------+                             
           |        +-- Schistodesmus             
           |                                      
           |  +-------- Pseudocuneopsis           
           +--+                                   
              |  +----- Inversiunio               
              +--+                                
                 |  +-- Tchangsinaia              
                 +--+                             
                    +-- Unio

Dai et al., 2024 [Publ 4386], fig. 3

iTtTbiriSoTroTroSiD
11011295--
  +-------- Middendorffinaia_mongolica **
--+                                   
  |  +----- Inversiunio               
  +--+                                
     |  +-- Cuneopsis                 
     +--+                             
        +-- Nodularia

Hou et al., 2024 [Publ 4400], fig. 1

iTtTbiriSoTroTroSiD
110114123--
           +----- Cuneopsis_rufescens       
  +--------+                                
  |        |  +-- Middendorffinaia_mongolica **
  |        +--+                             
--+           +-- Pseudocuneopsis           
  |                                         
  |  +----------- Tchangsinaia              
  |  |                                      
  +--+        +-- Cuneopsis_kiangsiensis    
     |  +-----+                             
     |  |     +-- Cuneopsis_celtiformis     
     +--+                                   
        |  +----- Cuneopsis_szechenyii      
        +--+                                
           |  +-- Cuneopsis_heudei          
           +--+                             
              +-- Cuneopsis_demangei

Hou et al., 2025 [Publ 4432], fig. 4

iTtTbiriSoTroTroSiD
110127255--
  +-------------- Middendorffinaia_mongolica **
--+                                         
  |  +----------- Pseudocuneopsis           
  +--+                                      
     |  +-------- Nodularia                 
     +--+                                   
        |  +----- Schistodesmus             
        +--+                                
           |  +-- Tchangsinaia              
           +--+                             
              +-- Cuneopsis

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iTtTbiriSoTroTroSiD
1101120267--
  +----------------- Aculamprotula             
  |                                            
--+  +-------------- Middendorffinaia_mongolica **
  |  |                                         
  +--+  +----------- Unio                      
     |  |                                      
     +--+  +-------- Nodularia                 
        |  |                                   
        +--+  +----- Schistodesmus             
           |  |                                
           +--+----- Cuneopsis                 
              |                                
              |  +-- Tchangsinaia              
              +--+                             
                 +-- Inversiunio

Dai et al., 2024 [Publ 4386], fig. 5-ML

iTtTbiriSoTroTroSiD
110120106--
  +----------------- Aculamprotula             
--+                                            
  |  +-------------- Unio                      
  +--+                                         
     |  +----------- Middendorffinaia_mongolica **
     +--+                                      
        |  +-------- Nodularia                 
        +--+                                   
           |  +----- Cuneopsis                 
           +--+                                
              |  +-- Inversiunio               
              +--+                             
                 +-- Schistodesmus

Hou et al., 2025 [Publ 4432], fig. 3

iTtTbiriSoTroTroSiD
11011093--
  +-------- Tchangsinaia              
--+                                   
  |  +----- Pseudocuneopsis           
  +--+                                
     |  +-- Middendorffinaia_mongolica **
     +--+                             
        +-- Cuneopsis

Bolotov et al., 2020 [Publ 4084], fig. 1

iTtTbiriSoTroTroSiD
11013542--
  +----- Middendorffinaia_mongolica **
--+                                
  |  +-- Nodularia                 
  +--+                             
     +-- Unio

Klishko et al., 2019 [Publ 3886], fig. 9

iTtTbiriSoTroTroSiD
11011072--
  +----- Unio                      
--+                                
  |  +-- Middendorffinaia_mongolica **
  +--+                             
     +-- Nodularia

Konopleva et al., 2023 [Publ 4269], fig. S1_4

iTtTbiriSoTroTroSiD
110155115--
  +-------------- Middendorffinaia_mongolica **
--+                                         
  |  +----------- Aculamprotula             
  +--+                                      
     |  +-------- Unio                      
     +--+                                   
        |  +----- Nodularia                 
        +--+                                
           |  +-- Tchangsinaia              
           +--+                             
              +-- Cuneopsis

Chen et al., 2023 [Publ 4302], fig. 2

iTtTbiriSoTroTroSiD
110139169--
  +----------------- Aculamprotula             
  |                                            
  |              +-- Middendorffinaia_mongolica **
--+  +-----------+                             
  |  |           +-- Nodularia                 
  |  |                                         
  +--+  +----------- Cuneopsis                 
     |  |                                      
     |  |        +-- Diaurora                  
     +--+--------+                             
        |        +-- Schistodesmus             
        |                                      
        |  +-------- Pseudocuneopsis           
        +--+                                   
           |  +----- Inversiunio               
           +--+                                
              |  +-- Tchangsinaia              
              +--+                             
                 +-- Unio

Dai et al., 2024 [Publ 4386], fig. 5-BI

iTtTbiriSoTroTroSiD
110120106--
  +----------- Unio                      
  |                                      
--+  +-------- Middendorffinaia_mongolica **
  |  |                                   
  +--+  +----- Cuneopsis                 
     |  |                                
     +--+----- Nodularia                 
        |                                
        |  +-- Inversiunio               
        +--+                             
           +-- Schistodesmus

Chen et al., 2023 [Publ 4302], fig. 1

iTtTbiriSoTroTroSiD
110139169--
  +----------------- Aculamprotula             
  |                                            
  |              +-- Middendorffinaia_mongolica **
--+  +-----------+                             
  |  |           +-- Nodularia                 
  |  |                                         
  +--+  +----------- Cuneopsis                 
     |  |                                      
     |  |        +-- Diaurora                  
     +--+--------+                             
        |        +-- Schistodesmus             
        |                                      
        |  +-------- Pseudocuneopsis           
        +--+                                   
           |  +----- Inversiunio               
           +--+                                
              |  +-- Tchangsinaia              
              +--+                             
                 +-- Unio
 
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