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Page last updated
Thu 7 August 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Bivalvia | Unionoida | Hyrioidea | Hyriidae | Velesunioninae

genus Lortiella Modell, 1942: cladomics
     3 Recent valid species (3 total species)

Cladomics Summary

number of species = 3 (3 total)
analyzed in 22 cladograms in 13 publications
maximum number of terminals in an analysis = 1 (mean = 1, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 0
maximum number of included terminals = , (mean = median = )
proportion of cladograms that support basic monophyly = 0

Graf et al., 2015 [Publ 3619], fig. 1

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
11011964--
  +----------- Westralunio        
  |                               
  |     +----- Alathyria_jacksoni 
--+  +--+                         
  |  |  |  +-- Velesunio          
  |  |  +--+                      
  +--+     +-- Alathyria_profuga  
     |                            
     |  +----- Alathyria_pertexta 
     +--+                         
        |  +-- Microdontia        
        +--+                      
           +-- Lortiella_froggatti **

Bolotov et al., 2017 [Publ 3761], fig. S2

iTtTbiriSoTroTroSiD
110113853--
     +----- Velesunio          
  +--+                         
  |  |  +-- Alathyria_jacksoni 
  |  +--+                      
--+     +-- Alathyria_profuga  
  |                            
  |  +----- Microdontia        
  +--+                         
     |  +-- Alathyria_pertexta 
     +--+                      
        +-- Lortiella_froggatti **

Bolotov et al., 2016 [Publ 3743], fig. 4

iTtTbiriSoTroTroSiD
11012953--
     +----- Microdontia        
  +--+                         
  |  |  +-- Alathyria_pertexta 
  |  +--+                      
--+     +-- Lortiella_froggatti **
  |                            
  |  +----- Velesunio          
  +--+                         
     |  +-- Alathyria_jacksoni 
     +--+                      
        +-- Alathyria_profuga

Roe & Hoeh, 2003 [Publ 2655], fig. 4f2

iTtTbiriSoTroTroSiD
11013011--
-- Lortiella_rugata **

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
110119864--
     +----- Alathyria_jacksoni 
  +--+                         
  |  |  +-- Alathyria_profuga  
  |  +--+                      
--+     +-- Velesunio          
  |                            
  |  +----- Alathyria_pertexta 
  +--+                         
     |  +-- Lortiella_froggatti **
     +--+                      
        +-- Microdontia

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
110119864--
     +----- Velesunio          
  +--+                         
  |  |  +-- Alathyria_jacksoni 
  |  +--+                      
--+     +-- Alathyria_profuga  
  |                            
  |  +----- Alathyria_pertexta 
  +--+                         
     |  +-- Lortiella_froggatti **
     +--+                      
        +-- Microdontia

Graf & Cummings, 2006 [Publ 1968], fig. 2

iTtTbiriSoTroTroSiD
11014621--
  +----- Velesunio_ambiguus
--+                        
  |  +-- Velesunio_angasi  
  +--+                     
     +-- Lortiella_rugata **

Hoeh et al., 2001 [Publ 2652], fig. 14f2

iTtTbiriSoTroTroSiD
11013011--
-- Lortiella_rugata **

Marshall et al., 2014 [Publ 3593], fig. 1

iTtTbiriSoTroTroSiD
11011321--
  +----- Velesunio_ambiguus
--+                        
  |  +-- Velesunio         
  +--+                     
     +-- Lortiella **

Walker et al., 2006 [Publ 2666], fig. 6

iTtTbiriSoTroTroSiD
11013542--
-- 2889

Hoeh et al., 2009 [Publ 2716], fig. 3

iTtTbiriSoTroTroSiD
11014621--
  +----- Velesunio_ambiguus
--+                        
  |  +-- Velesunio_angasi  
  +--+                     
     +-- Lortiella_rugata **

Hoeh et al., 2001 [Publ 2652], fig. 14f4

iTtTbiriSoTroTroSiD
11013011--
  +-- Lortiella_rugata **
--+                   
  +-- Velesunio

Hoeh et al., 2009 [Publ 2716], fig. 5

iTtTbiriSoTroTroSiD
11014521--
  +----- Velesunio_ambiguus
--+                        
  |  +-- Velesunio_angasi  
  +--+                     
     +-- Lortiella_rugata **

Roe & Hoeh, 2003 [Publ 2655], fig. 4f3

iTtTbiriSoTroTroSiD
11013011--
  +-- Velesunio       
--+                   
  +-- Lortiella_rugata **

Bolotov et al., 2017 [Publ 3761], fig. S1

iTtTbiriSoTroTroSiD
110113853--
     +----- Alathyria_pertexta 
  +--+                         
  |  |  +-- Microdontia        
  |  +--+                      
--+     +-- Lortiella_froggatti **
  |                            
  |  +----- Velesunio          
  +--+                         
     |  +-- Alathyria_jacksoni 
     +--+                      
        +-- Alathyria_profuga

Santos-Neto et al., 2016 [Publ 3739], fig. 2a_b

iTtTbiriSoTroTroSiD
11011954--
  +----------- Velesunio_ambiguus
--+                              
  |  +-------- Microdontia       
  +--+                           
     |  +----- Lortiella **         
     +--+                        
        |  +-- Alathyria         
        +--+                     
           +-- Velesunio_angasi

Bolotov et al., 2016 [Publ 3743], fig. 2

iTtTbiriSoTroTroSiD
11012953--
     +----- Microdontia        
  +--+                         
  |  |  +-- Lortiella_froggatti **
  |  +--+                      
--+     +-- Alathyria_pertexta 
  |                            
  |  +----- Velesunio          
  +--+                         
     |  +-- Alathyria_jacksoni 
     +--+                      
        +-- Alathyria_profuga

Bogan & Hoeh, 2000 [Publ 2650], fig. 1

iTtTbiriSoTroTroSiD
11013311--
  +-- Velesunio       
--+                   
  +-- Lortiella_rugata **

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
110119664--
     +----- Velesunio          
  +--+                         
  |  |  +-- Alathyria_jacksoni 
  |  +--+                      
--+     +-- Alathyria_profuga  
  |                            
  |  +----- Alathyria_pertexta 
  +--+                         
     |  +-- Lortiella_froggatti **
     +--+                      
        +-- Microdontia

Hoeh et al., 2009 [Publ 2716], fig. 4

iTtTbiriSoTroTroSiD
11014521--
  +----- Velesunio_ambiguus
--+                        
  |  +-- Velesunio_angasi  
  +--+                     
     +-- Lortiella_rugata **

Hoeh et al., 2002 [Publ 2653], fig. 2

iTtTbiriSoTroTroSiD
11013011--
  +-- Velesunio
--+            
  +-- Lortiella **

Santos-Neto et al., 2016 [Publ 3739], fig. 4

iTtTbiriSoTroTroSiD
11011954--
  +----------- Velesunio_ambiguus
--+                              
  |  +-------- Alathyria         
  +--+                           
     |  +----- Velesunio_angasi  
     +--+                        
        |  +-- Lortiella **         
        +--+                     
           +-- Microdontia
 
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