The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)
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Bivalvia | Unionoida | Unionoidea | Unionidae | Unioninae | Unionini genus Tchangsinaia Starobogatov, 1970: cladomics
1 Recent valid species (1 total species) Cladomics Summarynumber of species = 1 (1 total)
analyzed in 81 cladograms in 51 publications
maximum number of terminals in an analysis =
1 (mean = 1, median = 1)
Dai et al., 2025 [Publ 4440], fig. 1 ** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram ** iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 44 | 21 | 10 | -- | +----------------------- Globunio
|
--+ +-------------------- Aculamprotula
| |
+--+ +----------------- Middendorffinaia
| |
+--+ +-------------- Nodularia
| |
+--+ +----------- Cuneopsis
| |
| | +-- Diaurora
+--+--------+
| +-- Schistodesmus
|
| +-------- Pseudocuneopsis
+--+
| +----- Unio
+--+
| +-- Tchangsinaia_piscicula **
+--+
+-- Inversiunio
Wang et al., 2018 [Publ 3846], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 30 | 7 | 4 | -- | +----------- Aculamprotula
--+
| +-------- Unio
+--+
| +----- Nodularia
+--+
| +-- Tchangsinaia_piscicula **
+--+
+-- Cuneopsis
Wu et al., 2020 [Publ 4121], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 48 | 13 | 6 | -- | +----------------- Aculamprotula
--+
| +-------------- Unio
+--+
| +----------- Pseudocuneopsis
+--+
| +-------- Nodularia
+--+
| +----- Tchangsinaia_piscicula **
+--+
| +-- Cuneopsis
+--+
+-- Schistodesmus
Dai et al., 2025 [Publ 4440], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 44 | 21 | 10 | -- | +-------------------- Globunio
|
--+ +----------------- Aculamprotula
| |
| | +-- Middendorffinaia
+--+ +-----------+
| | +-- Nodularia
| |
+--+ +----------- Cuneopsis
| |
| | +-- Diaurora
+--+--------+
| +-- Schistodesmus
|
| +-------- Pseudocuneopsis
+--+
| +----- Inversiunio
+--+
| +-- Tchangsinaia_piscicula **
+--+
+-- Unio
Bolotov et al., 2017 [Publ 3761], fig. S1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 138 | 9 | 5 | -- | +-------------- Unio
--+
| +----------- Aculamprotula
+--+
| +-------- Nodularia
+--+
| +----- Cuneopsis
+--+
| +-- Tchangsinaia_piscicula **
+--+
+-- Schistodesmus
Froufe et al., 2019 [Publ 3936], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 43 | 9 | 4 | -- | +----------- Aculamprotula
--+
| +-------- Unio
+--+
| +----- Nodularia
+--+
| +-- Tchangsinaia_piscicula **
+--+
+-- Schistodesmus
Wu et al., 2021 [Publ 4366], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 90 | 10 | 5 | -- | +-------------- Aculamprotula
--+
| +----------- Unio
+--+
| +-------- Nodularia
+--+
| +----- Cuneopsis
+--+
| +-- Tchangsinaia_piscicula **
+--+
+-- Schistodesmus
Mabuchi et al., 2021 [Publ 4233], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 4 | 4 | 1 | -- | +-- Tchangsinaia_piscicula **
--+
+-- Nodularia
Cha et al., 2018 [Publ 3916], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 37 | 9 | 4 | -- | +----------- Aculamprotula
--+
| +-------- Unio
+--+
| +----- Schistodesmus
+--+
| +-- Nodularia
+--+
+-- Tchangsinaia_piscicula **
Pavan-Kumar et al., 2022 [Publ 4263], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 35 | 5 | 4 | -- | +----------- Aculamprotula
--+
| +-------- Unio
+--+
| +----- Schistodesmus
+--+
| +-- Nodularia
+--+
+-- Tchangsinaia_piscicula **
Sano et al., 2020 [Publ 4138], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 67 | 12 | 5 | -- | +-------------- Aculamprotula
--+
| +----------- Unio
+--+
| +-------- Inversiunio
+--+
| +----- Nodularia
+--+
| +-- Schistodesmus
+--+
+-- Tchangsinaia_piscicula **
Aunins et al., 2018 [Publ 3863], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 39 | 10 | 5 | -- | +-------------- Aculamprotula
--+
| +----------- Unio
+--+
| +-------- Nodularia
+--+
| +----- Tchangsinaia_piscicula **
+--+
| +-- Cuneopsis
+--+
+-- Schistodesmus
Dai et al., 2023 [Publ 4285], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 52 | 6 | 6 | -- | +-------------- Aculamprotula
|
--+ +----------- Unio
| |
+--+ +-- Pseudocuneopsis
| +-----+
| | +-- Nodularia
+--+
| +----- Schistodesmus
+--+
| +-- Cuneopsis
+--+
+-- Tchangsinaia_piscicula **
Wu et al., 2018 [Publ 3836], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 52 | 13 | 5 | -- | +----------- Aculamprotula
|
--+ +-------- Unio
| |
+--+ +-- Nodularia
| +--+
| | +-- Schistodesmus
+--+
| +-- Tchangsinaia_piscicula **
+--+
+-- Cuneopsis
Chen et al., 2023 [Publ 4302], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 39 | 16 | 9 | -- | +----------------- Aculamprotula
|
| +-- Middendorffinaia
--+ +-----------+
| | +-- Nodularia
| |
+--+ +----------- Cuneopsis
| |
| | +-- Diaurora
+--+--------+
| +-- Schistodesmus
|
| +-------- Pseudocuneopsis
+--+
| +----- Inversiunio
+--+
| +-- Tchangsinaia_piscicula **
+--+
+-- Unio
Hou et al., 2025 [Publ 4432], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 10 | 9 | 3 | -- | +-------- Tchangsinaia_piscicula **
--+
| +----- Pseudocuneopsis
+--+
| +-- Middendorffinaia
+--+
+-- Cuneopsis
Huang et al., 2018 [Publ 3828], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 41 | 8 | 3 | -- | +-------- Aculamprotula
--+
| +----- Unio
+--+
| +-- Tchangsinaia_piscicula **
+--+
+-- Nodularia
Bolotov et al., 2017 [Publ 3761], fig. S2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 138 | 9 | 5 | -- | +----------- Unio
|
--+ +-------- Aculamprotula
| |
| +-------- Schistodesmus
+--+
+-------- Nodularia
|
| +----- Cuneopsis_szechenyii
+--+
| +-- Tchangsinaia_piscicula **
+--+
+-- Cuneopsis_rufescens
Wu et al., 2019 [Publ 3864], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 45 | 12 | 6 | -- | +----------------- Aculamprotula
--+
| +-------------- Unio
+--+
| +----------- Pseudocuneopsis
+--+
| +-------- Nodularia
+--+
| +----- Tchangsinaia_piscicula **
+--+
| +-- Cuneopsis
+--+
+-- Schistodesmus
Chen et al., 2023 [Publ 4302], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 39 | 16 | 9 | -- | +----------------- Aculamprotula
|
| +-- Middendorffinaia
--+ +-----------+
| | +-- Nodularia
| |
+--+ +----------- Cuneopsis
| |
| | +-- Diaurora
+--+--------+
| +-- Schistodesmus
|
| +-------- Pseudocuneopsis
+--+
| +----- Inversiunio
+--+
| +-- Tchangsinaia_piscicula **
+--+
+-- Unio
Wu et al., 2022 [Publ 4238], fig. 2a_S1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 11 | 11 | 6 | -- | +----------------- Aculamprotula
--+
| +-------------- Nodularia
+--+
| +----------- Pseudocuneopsis
+--+
| +-------- Schistodesmus
+--+
| +----- Cuneopsis
+--+
| +-- Tchangsinaia_piscicula **
+--+
+-- Inversiunio
Wu et al., 2024 [Publ 4379], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 52 | 28 | 6 | -- | +----------------- Aculamprotula
--+
| +-------------- Unio
+--+
| +----------- Pseudocuneopsis
+--+
| +-------- Nodularia
+--+
| +----- Schistodesmus
+--+
| +-- Tchangsinaia_piscicula **
+--+
+-- Cuneopsis
Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 75 | 9 | 5 | -- | +----------------- Unio
--+
| +-------------- Aculamprotula
+--+
| +----------- Nodularia
+--+
| +-------- Schistodesmus
+--+
| +----- Cuneopsis_szechenyii
+--+
| +-- Tchangsinaia_piscicula **
+--+
+-- Cuneopsis_rufescens
Wu et al., 2022 [Publ 4238], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 72 | 19 | 6 | -- | +----------------- Aculamprotula
--+
| +-------------- Unio
+--+
| +----------- Pseudocuneopsis
+--+
| +-------- Nodularia
+--+
| +----- Schistodesmus
+--+
| +-- Tchangsinaia_piscicula **
+--+
+-- Cuneopsis
Fukata & Iigo, 2019 [Publ 4108], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 15 | 11 | 6 | -- | +----------- Unio
|
--+ +-------- Pseudocuneopsis
| |
+--+ +-- Schistodesmus
| +--+
| | +-- Cuneopsis
+--+
| +-- Tchangsinaia_piscicula **
+--+
+-- Nodularia
Liu et al., 2023 [Publ 4310], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 23 | 13 | 7 | -- | +----------------- Unio
|
--+ +-------------- Aculamprotula
| |
+--+ +----------- Cuneopsis
| |
+--+ +-- Nodularia
| +-----+
| | +-- Schistodesmus
+--+
| +----- Tchangsinaia_piscicula **
+--+
| +-- Inversiunio
+--+
+-- Pseudocuneopsis
Hou et al., 2024 [Publ 4400], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 75 | 48 | 7 | -- | +-------------------- Aculamprotula
--+
| +----------------- Unio
+--+
| +-------------- Middendorffinaia
+--+
| +----------- Pseudocuneopsis
+--+
| +-------- Nodularia
+--+
| +----- Schistodesmus
+--+
| +-- Tchangsinaia_piscicula **
+--+
+-- Cuneopsis
Huang et al., 2019 [Publ 3843], fig. S4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 198 | 18 | 5 | -- | +----- Cuneopsis_celtiformis
|
+----- Tchangsinaia_piscicula **
|
--+----- Cuneopsis_rufescens
|
+----- Nodularia
|
| +-- Cuneopsis_szechenyii
+--+
+-- Cuneopsis_heudei
Sano et al., 2020 [Publ 4138], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 67 | 12 | 5 | -- | +-------------- Aculamprotula
--+
| +----------- Unio
+--+
| +-------- Nodularia
+--+
| +----- Inversiunio
+--+
| +-- Schistodesmus
+--+
+-- Tchangsinaia_piscicula **
Dai et al., 2024 [Publ 4386], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 39 | 19 | 6 | -- | +----------------- Aculamprotula
--+
| +-------------- Unio
+--+
| +----------- Pseudocuneopsis
+--+
| +-------- Nodularia
+--+
| +----- Schistodesmus
+--+
| +-- Tchangsinaia_piscicula **
+--+
+-- Cuneopsis
Pfeiffer et al., 2019 [Publ 3937], fig. 4-5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 56 | 2 | 2 | -- | +----- Unio
--+
| +-- Nodularia
+--+
+-- Tchangsinaia_piscicula **
Huang et al., 2019 [Publ 3843], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 198 | 18 | 5 | -- | +-------------- Aculamprotula
--+
| +----------- Unio
+--+
| +-------- Schistodesmus
+--+
| +----- Nodularia
+--+
| +-- Tchangsinaia_piscicula **
+--+
+-- Cuneopsis
Huang et al., 2019 [Publ 3843], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 55 | 10 | 4 | -- | +----------- Aculamprotula
--+
| +-------- Unio
+--+
| +----- Nodularia
+--+
| +-- Tchangsinaia_piscicula **
+--+
+-- Schistodesmus
Wu et al., 2020 [Publ 4147], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 28 | 9 | 6 | -- | +----------------- Aculamprotula
--+
| +-------------- Unio
+--+
| +----------- Pseudocuneopsis
+--+
| +-------- Nodularia
+--+
| +----- Schistodesmus
+--+
| +-- Cuneopsis
+--+
+-- Tchangsinaia_piscicula **
Wu et al., 2024 [Publ 4388], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 53 | 8 | 6 | -- | +----------------- Aculamprotula
--+
| +-------------- Unio
+--+
| +----------- Pseudocuneopsis
+--+
| +-------- Nodularia
+--+
| +----- Tchangsinaia_piscicula **
+--+
| +-- Cuneopsis
+--+
+-- Schistodesmus
Sano et al., 2020 [Publ 4138], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 67 | 12 | 5 | -- | +-------------- Aculamprotula
--+
| +----------- Unio
+--+
| +-------- Inversiunio
+--+
| +----- Nodularia
+--+
| +-- Tchangsinaia_piscicula **
+--+
+-- Schistodesmus
Wu et al., 2023 [Publ 4339], fig. 4_S5-aa iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 94 | 21 | 6 | -- | +-------------- Aculamprotula
|
--+ +----------- Unio
| |
+--+ +-- Nodularia
| +-----+
| | +-- Pseudocuneopsis
+--+
| +----- Tchangsinaia_piscicula **
+--+
| +-- Schistodesmus
+--+
+-- Cuneopsis
Hou et al., 2024 [Publ 4400], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 14 | 12 | 3 | -- | +----- Cuneopsis_rufescens
+--------+
| | +-- Middendorffinaia
| +--+
--+ +-- Pseudocuneopsis
|
| +----------- Tchangsinaia_piscicula **
| |
+--+ +-- Cuneopsis_kiangsiensis
| +-----+
| | +-- Cuneopsis_celtiformis
+--+
| +----- Cuneopsis_szechenyii
+--+
| +-- Cuneopsis_heudei
+--+
+-- Cuneopsis_demangei
Wu et al., 2021 [Publ 4366], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 67 | 14 | 6 | -- | +----------------- Aculamprotula
--+
| +-------------- Unio
+--+
| +----------- Pseudocuneopsis
+--+
| +-------- Nodularia
+--+
| +----- Schistodesmus
+--+
| +-- Tchangsinaia_piscicula **
+--+
+-- Cuneopsis
Hou et al., 2025 [Publ 4432], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 27 | 25 | 5 | -- | +-------------- Middendorffinaia
--+
| +----------- Pseudocuneopsis
+--+
| +-------- Nodularia
+--+
| +----- Schistodesmus
+--+
| +-- Tchangsinaia_piscicula **
+--+
+-- Cuneopsis
Lopes-Lima et al., 2024 [Publ 4376], fig. 4-5_MtG iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 21 | 18 | 2 | -- | +----- Unio
--+
| +-- Tchangsinaia_piscicula **
+--+
+-- Nodularia
Wu et al., 2021 [Publ 4366], fig. 3-ML iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 72 | 14 | 6 | -- | +----------------- Aculamprotula
--+
| +-------------- Unio
+--+
| +----------- Pseudocuneopsis
+--+
| +-------- Nodularia
+--+
| +----- Tchangsinaia_piscicula **
+--+
| +-- Schistodesmus
+--+
+-- Cuneopsis
Wu et al., 2024 [Publ 4350], fig. 8a_S1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 90 | 21 | 6 | -- | +-------------- Aculamprotula
|
--+ +----------- Unio
| |
+--+ +-------- Nodularia
| |
+--+-------- Pseudocuneopsis
|
| +----- Schistodesmus
+--+
| +-- Tchangsinaia_piscicula **
+--+
+-- Cuneopsis
Bolotov et al., 2017 [Publ 3764], fig. S1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 124 | 9 | 5 | -- | +-- Unio_crassus
|
+-- Unio_pictorum
|
+-- Unio_tumidus
|
+-- Aculamprotula
|
+-- Cuneopsis_szechenyii
--+
+-- Tchangsinaia_piscicula **
|
+-- Cuneopsis_rufescens
|
+-- Nodularia_douglasiae
|
+-- Nodularia_nuxpersicae
|
+-- Schistodesmus
Zhou et al., 2007 [Publ 2658], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 20 | 6 | 3 | -- | +----- Aculamprotula
--+
| +-- Tchangsinaia_piscicula **
+--+
+-- Nodularia
Wu et al., 2024 [Publ 4379], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 11 | 11 | 3 | -- | +----- Nodularia
|
--+ +-- Cuneopsis
| |
+--+-- Tchangsinaia_piscicula **
|
+-- Pseudocuneopsis
Huang et al., 2019 [Publ 3843], fig. S3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 196 | 17 | 5 | -- | +-------------- Aculamprotula
--+
| +----------- Unio
+--+
| +-------- Nodularia
+--+
| +----- Schistodesmus
+--+
| +-- Tchangsinaia_piscicula **
+--+
+-- Cuneopsis
Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 75 | 9 | 5 | -- | +----------------- Unio
--+
| +-------------- Aculamprotula
+--+
| +----------- Nodularia
+--+
| +-------- Schistodesmus
+--+
| +----- Cuneopsis_szechenyii
+--+
| +-- Tchangsinaia_piscicula **
+--+
+-- Cuneopsis_rufescens
Feng et al., 2016 [Publ 3800], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 13 | 0 | 0 | -- | -- Tchangsinaia_piscicula **
Wu et al., 2021 [Publ 4366], fig. 3-BI iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 72 | 14 | 6 | -- | +----------------- Aculamprotula
--+
| +-------------- Unio
+--+
| +----------- Pseudocuneopsis
+--+
| +-------- Nodularia
+--+
| +----- Schistodesmus
+--+
| +-- Cuneopsis
+--+
+-- Tchangsinaia_piscicula **
Wu et al., 2022 [Publ 4238], fig. 2e iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 16 | 11 | 6 | -- | +----------------- Aculamprotula
--+
| +-------------- Unio
+--+
| +----------- Pseudocuneopsis
+--+
| +-------- Nodularia
+--+
| +----- Tchangsinaia_piscicula **
+--+
| +-- Cuneopsis
+--+
+-- Schistodesmus
Wu et al., 2022 [Publ 4238], fig. 2d iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 37 | 22 | 7 | -- | +-------------------- Aculamprotula
--+
| +----------------- Unio
+--+
| +-------------- Pseudocuneopsis
+--+
| +----------- Nodularia
+--+
| +-------- Schistodesmus
+--+
| +----- Cuneopsis
+--+
| +-- Tchangsinaia_piscicula **
+--+
+-- Inversiunio
Pfeiffer et al., 2019 [Publ 3937], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 56 | 2 | 2 | -- | +----- Unio
--+
| +-- Tchangsinaia_piscicula **
+--+
+-- Nodularia
Konopleva et al., 2023 [Publ 4269], fig. S1_4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 55 | 11 | 5 | -- | +-------------- Middendorffinaia
--+
| +----------- Aculamprotula
+--+
| +-------- Unio
+--+
| +----- Nodularia
+--+
| +-- Tchangsinaia_piscicula **
+--+
+-- Cuneopsis
Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 79 | 10 | 5 | -- | +-------------- Unio
--+
| +----------- Aculamprotula
+--+
| +-------- Nodularia
+--+
| +----- Schistodesmus
+--+
| +-- Tchangsinaia_piscicula **
+--+
+-- Cuneopsis
Huang et al., 2019 [Publ 3843], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 42 | 14 | 4 | -- | +----------------- Schistodesmus
--+
| +-------------- Nodularia
+--+
| +----------- Cuneopsis_rufescens
+--+
| +-------- Tchangsinaia_piscicula **
+--+
| +----- Cuneopsis_celtiformis
+--+
| +-- Cuneopsis_szechenyii
+--+
+-- Cuneopsis_heudei
Wu et al., 2023 [Publ 4296], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 22 | 12 | 7 | -- | +----------------- Unio
|
--+ +-------------- Aculamprotula
| |
+--+ +----------- Cuneopsis
| |
+--+ +-- Nodularia
| +-----+
| | +-- Schistodesmus
+--+
| +----- Tchangsinaia_piscicula **
+--+
| +-- Inversiunio
+--+
+-- Pseudocuneopsis
Lopes-Lima et al., 2017 [Publ 3792], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 31 | 6 | 4 | -- | +----------- Aculamprotula
--+
| +-------- Unio
+--+
| +----- Schistodesmus
+--+
| +-- Tchangsinaia_piscicula **
+--+
+-- Nodularia
Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 120 | 26 | 7 | -- | +----------------- Aculamprotula
|
--+ +-------------- Middendorffinaia
| |
+--+ +----------- Unio
| |
+--+ +-------- Nodularia
| |
+--+ +----- Schistodesmus
| |
+--+----- Cuneopsis
|
| +-- Tchangsinaia_piscicula **
+--+
+-- Inversiunio
Huang et al., 2019 [Publ 3843], fig. S1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 44 | 14 | 4 | -- | +----------- Aculamprotula
--+
| +-------- Nodularia
+--+
| +----- Tchangsinaia_piscicula **
+--+
| +-- Schistodesmus
+--+
+-- Cuneopsis
Zhang et al., 2004 [Publ 2735], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 13 | 6 | 4 | -- | +-------------- Tchangsinaia_piscicula **
--+
| +----------- Cuneopsis_heudei
+--+
| +-------- Nodularia
+--+
| +----- Pseudocuneopsis
+--+
| +-- Cuneopsis_rufescens
+--+
+-- Schistodesmus
Wu et al., 2021 [Publ 4366], fig. 2_I iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 90 | 10 | 5 | -- | +-------------- Aculamprotula
--+
| +----------- Unio
+--+
| +-------- Nodularia
+--+
| +----- Cuneopsis
+--+
| +-- Tchangsinaia_piscicula **
+--+
+-- Schistodesmus
Dai et al., 2025 [Publ 4440], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 87 | 28 | 8 | -- | +----------------- Globunio
|
--+ +-------------- Aculamprotula
| |
+--+ +----------- Unio
| |
+--+ +-- Nodularia
| +-----+
| | +-- Pseudocuneopsis
+--+
| +----- Schistodesmus
+--+
| +-- Tchangsinaia_piscicula **
+--+
+-- Cuneopsis
Dai et al., 2024 [Publ 4358], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 51 | 16 | 6 | -- | +----------------- Aculamprotula
--+
| +-------------- Unio
+--+
| +----------- Pseudocuneopsis
+--+
| +-------- Nodularia
+--+
| +----- Schistodesmus
+--+
| +-- Tchangsinaia_piscicula **
+--+
+-- Cuneopsis
Huang et al., 2002 [Publ 2657], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 33 | 2 | 2 | -- | +-- Nodularia
--+
+-- Tchangsinaia_piscicula **
Soroka & Burzynski, 2017 [Publ 3837], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 22 | 9 | 4 | -- | +----------- Aculamprotula
--+
| +-------- Unio
+--+
| +----- Schistodesmus
+--+
| +-- Tchangsinaia_piscicula **
+--+
+-- Nodularia
Liu et al., 2023 [Publ 4310], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 23 | 13 | 7 | -- | +-------- Unio
|
+-------- Aculamprotula
|
--+ +----- Cuneopsis
| |
| | +-- Nodularia
| +--+
+--+ +-- Schistodesmus
|
| +-- Tchangsinaia_piscicula **
| |
+--+-- Inversiunio
|
+-- Pseudocuneopsis
Huang et al., 2002 [Publ 2657], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 33 | 2 | 2 | -- | +-- Nodularia
--+
+-- Tchangsinaia_piscicula **
Zieritz et al., 2021 [Publ 4231], fig. 3-F_5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 47 | 9 | 4 | -- | +----------- Aculamprotula
--+
| +-------- Unio
+--+
| +----- Nodularia
+--+
| +-- Schistodesmus
+--+
+-- Tchangsinaia_piscicula **
Zhou et al., 2017 [Publ 3788], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 15 | 3 | 2 | -- | +----- Aculamprotula
--+
| +-- Nodularia
+--+
+-- Tchangsinaia_piscicula **
Sano et al., 2017 [Publ 3799], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 32 | 7 | 3 | -- | +-- Tchangsinaia_piscicula **
--+
+-- Nodularia
Wang et al., 2018 [Publ 3865], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 30 | 6 | 3 | -- | +-------- Aculamprotula
--+
| +----- Unio
+--+
| +-- Tchangsinaia_piscicula **
+--+
+-- Nodularia
Choi et al., 2020 [Publ 4169], fig. 2a iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 11 | 11 | 4 | -- | +----------- Nodularia
--+
| +-------- Schistodesmus
+--+
| +----- Cuneopsis
+--+
| +-- Unio
+--+
+-- Tchangsinaia_piscicula **
Fonseca et al., 2016 [Publ 3775], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 30 | 6 | 4 | -- | +----------- Aculamprotula
--+
| +-------- Unio
+--+
| +----- Schistodesmus
+--+
| +-- Nodularia
+--+
+-- Tchangsinaia_piscicula **
Wu et al., 2024 [Publ 4350], fig. 8b_S2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 90 | 21 | 6 | -- | +-------------- Aculamprotula
|
--+ +----------- Unio
| |
+--+ +-------- Nodularia
| |
+--+-------- Pseudocuneopsis
|
| +----- Schistodesmus
+--+
| +-- Tchangsinaia_piscicula **
+--+
+-- Cuneopsis
Hou et al., 2025 [Publ 4401], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 27 | 25 | 5 | -- | +-------------- Middendorffinaia
--+
| +----------- Pseudocuneopsis
+--+
| +-------- Nodularia
+--+
| +----- Schistodesmus
+--+
| +-- Tchangsinaia_piscicula **
+--+
+-- Cuneopsis
Froufe et al., 2019 [Publ 3936], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 43 | 9 | 4 | -- | +----------- Aculamprotula
--+
| +-------- Unio
+--+
| +----- Nodularia
+--+
| +-- Tchangsinaia_piscicula **
+--+
+-- Schistodesmus
Lopes-Lima et al., 2024 [Publ 4376], fig. 4-5_AHE iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 21 | 18 | 2 | -- | +----- Unio
--+
| +-- Tchangsinaia_piscicula **
+--+
+-- Nodularia
Wu et al., 2023 [Publ 4339], fig. 4_S5-nt iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 94 | 21 | 6 | -- | +-------------------- Aculamprotula
|
--+ +----------------- Unio
| |
+--+ +-------------- Pseudocuneopsis
| |
+--+ +----------- Nodularia
| |
+--+ +-- Schistodesmus_lampreyanus__1
| +-----+
| | +-- Cuneopsis
+--+
| +----- Tchangsinaia_piscicula **
+--+
| +-- Schistodesmus_lampreyanus__2
+--+
+-- Schistodesmus_spinosus
Pavan-Kumar et al., 2022 [Publ 4263], fig. S5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 37 | 5 | 4 | -- | +-------- Unio
--+
| +----- Schistodesmus
+--+
| +-- Nodularia
+--+
+-- Tchangsinaia_piscicula **
Fukata & Iigo, 2020 [Publ 4106], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 52 | 11 | 6 | -- | +-------------- Aculamprotula
|
--+ +----------- Unio
| |
+--+ +-- Schistodesmus
| +-----+
| | +-- Cuneopsis
+--+
| +----- Pseudocuneopsis
+--+
| +-- Nodularia
+--+
+-- Tchangsinaia_piscicula **
|
