MUSSELp
 
MUSSELpdb
  Introduction
Browse
d'basing
 
Taxonomy
  all genera
Unionidae
Margaritiferidae
Hyriidae
Etheriidae
Mycetopodidae
Iridinidae

Species & Genus Richness by Taxonomy
 
Geography
  Summary
North America
Central America
South America
Afrotropics
North Eurasia
East Asia
Australasia

Species Richness & Endemism by Geography
 
 

Page last updated
Thu 17 July 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

East Asia

East Asian SubregionsThe East Asia Region extends from the Amur River in Russia south through Japan, Korea, China, Indochina and Indonesia, and west to include India. East Asia is the most species-rich region on Earth, with a total of 361 species (91 genera) in three families: Unionidae, Margaritiferidae and Etheriidae.

The East Region is subdivided into seven subregions, each with a characteristic freshwater mussel assemblage. The low-richness basins north of the Amur draining into the Bering Sea and Sea of Okhotsk comprise Beringia (AS1; 3 gen., 3 spp., 0 endemic). The Amur-Korea Subregion (AS2; 17 gen., 29 spp., 16 endemic ) includes the Amur Basin and rivers draining to the Sea of Japan and the Yellow Sea, as far south as the Liao He Basin. Japan, Sakhalin Island, and the Kuril Islands are group as the Japan-Sakhalin Subregion (AS3; 14 gen., 28 spp., 19 endemic). The Yangtze-Huang Subregion (AS4; 29 gen., 86 spp., 62 endemic) covers the basins from the Pei River south through the Qiantang, as well as Taiwan. The area from southern China to the western Thailand, including the Mekong, is the Indochina Subregion (AS5; 45 gen., 149 spp., 126 endemic). The larger islands of Indonesia and the Philippines we group as the Sunda-Philippines Subregion (AS6; 16 gen., 31 spp., 24 endemic). The India-Myanmar Subregion (AS7; 20 gen., 82 spp., 77 endemic) extends from the Salween River in the east to the Indus on its western frontier. New Guinea is part of the Australasian Region, and Sulawesi does not seem to have any freshwater mussels.

Endemic species are marked with an asterisk (*).

Beringia (AS1)
  1. Beringiana beringiana
  2. Margaritifera middendorffi
  1. Nodularia douglasiae
Amur-Korea (AS2)
  1. Aculamprotula coreana*
  2. Amuranodonta kijaensis*
  3. Anemina arcaeformis
  4. Beringiana beringiana
  5. Buldowskia flavotincta*
  6. Buldowskia iwakawai
  7. Buldowskia shadini*
  8. Buldowskia suifunica*
  9. Cristaria beirensis*
  10. Cristaria plicata
  11. Inversidens pantoensis
  12. Inversiunio verrucosus*
  13. Koreosolenaia sitgyensis*
  14. Lamprotula leaii
  15. Lamprotula seomjinensis*
  1. Lanceolaria acrorrhyncha*
  2. Lanceolaria cylindrica*
  3. Lanceolaria grayii
  4. Margaritifera dahurica*
  5. Middendorffinaia mongolica*
  6. Nodularia breviconcha*
  7. Nodularia douglasiae
  8. Nodularia dualobtusa*
  9. Parvasolenaia triangularis
  10. Sinanodonta lauta
  11. Sinanodonta schrenkii*
  12. Sinanodonta woodiana
  13. Unio pictorum
  14. Unio tumidus
Japan-Sakhalin (AS3)
  1. Anemina arcaeformis
  2. Beringiana beringiana
  3. Beringiana fukuharai*
  4. Beringiana gosannensis*
  5. Beringiana japonica*
  6. Buldowskia iwakawai
  7. Buldowskia kamiyai*
  8. Cristaria clessini*
  9. Cristaria plicata
  10. Inversidens brandtii*
  11. Inversiunio jokohamensis*
  12. Inversiunio reinianus*
  13. Inversiunio yanagawensis*
  14. Lanceolaria kihirai*
  1. Lanceolaria oxyrhyncha*
  2. Margaritifera laevis*
  3. Margaritifera middendorffi
  4. Nodularia douglasiae
  5. Nodularia nipponensis*
  6. Obovalis omiensis*
  7. Pletholophus reinianus*
  8. Pronodularia japanensis*
  9. Sinanodonta calipygos*
  10. Sinanodonta lauta
  11. Sinanodonta pacifica
  12. Sinanodonta tumens*
  13. Sinohyriopsis cumingii
  14. Sinohyriopsis schlegelii*
Yangtze-Huang (AS4)
  1. Acudonta baitiaoensis*
  2. Aculamprotula fibrosa*
  3. Aculamprotula grandidens*
  4. Aculamprotula nodulosa
  5. Aculamprotula polysticta*
  6. Aculamprotula scripta*
  7. Aculamprotula tientsinensis*
  8. Aculamprotula tortuosa*
  9. Aculamprotula zonata*
  10. Acuticosta chinensis*
  11. Acuticosta ovata*
  12. Acuticosta retiaria*
  13. Acuticosta sichuanica*
  14. Acuticosta trisulcata*
  15. Amblema plicata
  16. Anemina arcaeformis
  17. Cristaria plicata
  18. Cristaria radiata*
  19. Cuneopsis celtiformis*
  20. Cuneopsis celtiosimilis*
  21. Cuneopsis heudei*
  22. Cuneopsis kiangsiensis*
  23. Cuneopsis rufescens*
  24. Cuneopsis szechenyii*
  25. Diaurora aurorea*
  26. Diaurora laeve*
  27. Gibbosula confragosa*
  28. Gibbosula rochechouartii*
  29. Globunio mirificus*
  30. Inversidens pantoensis
  31. Inversidens rentianensis*
  32. Lamprotula bazini
  33. Lamprotula caveata
  34. Lamprotula chiai*
  35. Lamprotula cornuumlunae*
  36. Lamprotula divergens*
  37. Lamprotula elongata*
  38. Lamprotula kouangensis*
  39. Lamprotula leaii
  40. Lamprotula linae*
  41. Lamprotula microsticta*
  42. Lamprotula paschalis*
  43. Lamprotula triclava*
  1. Lanceolaria eucylindrica*
  2. Lanceolaria gladiola
  3. Lanceolaria grayii
  4. Lanceolaria lanceolata*
  5. Lanceolaria triformis*
  6. Lanceolaria yueyingae*
  7. Lepidodesma aligera*
  8. Lepidodesma languilati*
  9. Nodularia douglasiae
  10. Nodularia fusiformans*
  11. Nodularia hanensis*
  12. Nodularia huana*
  13. Nodularia persculpta*
  14. Parvasolenaia neotriangularis*
  15. Parvasolenaia rivularis*
  16. Parvasolenaia triangularis
  17. Pletholophus tenuis
  18. Potamilus alatus
  19. Pseudobaphia aurea*
  20. Pseudobaphia biesiana*
  21. Pseudocuneopsis capitata*
  22. Pseudocuneopsis sichuanensis*
  23. Pseudocuneopsis yemaoi*
  24. Pseudopostolata angula*
  25. Ptychorhynchus liuovata
  26. Ptychorhynchus murinum*
  27. Ptychorhynchus pfisteri
  28. Ptychorhynchus schomburgianum
  29. Reginaia ebenus
  30. Schistodesmus lampreyanus*
  31. Schistodesmus spinosus*
  32. Simpsonasus nankingensis*
  33. Sinanodonta angula*
  34. Sinanodonta elliptica
  35. Sinanodonta lucida
  36. Sinanodonta pacifica
  37. Sinanodonta woodiana
  38. Sinohyriopsis cumingii
  39. Sinosolenaia carinata*
  40. Sinosolenaia iridinea*
  41. Sinosolenaia oleivora*
  42. Sinosolenaia recognita
  43. Tchangsinaia piscicula*
Indochina (AS5)
  1. Aculamprotula nodulosa
  2. Bineurus anodontinum*
  3. Bineurus exilis*
  4. Bineurus loeiensis*
  5. Bineurus mouhotii*
  6. Bineurus panhai*
  7. Chamberlainia duclerci*
  8. Chamberlainia hainesiana*
  9. Chamberlainia somsakpanhai*
  10. Cosmopseudodon resupinatus*
  11. Cosmopseudodon wenshanensis*
  12. Cristaria bellua*
  13. Cristaria plicata
  14. Cristaria truncata*
  15. Cuneopsis demangei*
  16. Ensidens dugasti*
  17. Ensidens ingallsianus*
  18. Ensidens jaculus*
  19. Ensidens sagittarius*
  20. Ensidens spiculus*
  21. Ensidens telus*
  22. Gibbosula crassa*
  23. Gibbosula laosensis*
  24. Guiunio rarus*
  25. Harmandia somboriensis*
  26. Hyriopsis bialata*
  27. Hyriopsis delaportei*
  28. Hyriopsis desowitzi*
  29. Hyriopsis khoratensis*
  30. Hyriopsis kratiensis*
  31. Hyriopsis myersiana*
  32. Hyriopsis panhai*
  33. Hyriopsis phuphaniensis*
  34. Hyriopsis sakhonensis*
  35. Indonaia gratiosa*
  36. Indonaia substriata*
  37. Isannaia fortunata*
  38. Isannaia occultata*
  39. Lamprotula bazini
  40. Lamprotula blaisei*
  41. Lamprotula caveata
  42. Lamprotula leaii
  43. Lamprotula nanningensis*
  44. Lamprotula plumbea*
  45. Lamprotula salaputium*
  46. Lanceolaria bilirata*
  47. Lanceolaria fruhstorferi*
  48. Lanceolaria gladiola
  49. Lanceolaria grayii
  50. Lanceolaria laevis*
  51. Lannanaia kokensis*
  52. Lens comptus*
  53. Lens contradens
  54. Lens eximius*
  55. Lens fultoni*
  56. Lens inornatus*
  57. Lens maenamensis*
  58. Lens micropterus*
  59. Lens misellus*
  60. Lens novoselovi*
  61. Lens pallegoixi*
  62. Lens rolfbrandti*
  63. Leoparreysia subcircularis*
  64. Leoparreysia superstes*
  65. Mimo guoliangi*
  66. Mimo jiaoyue*
  67. Modellnaia siamensis*
  68. Namkongnaia inkhavilayi*
  69. Namkongnaia lemeslei*
  70. Nodularia diespiter*
  71. Nodularia dorri*
  72. Nodularia douglasiae
  73. Nodularia gladiator*
  74. Nodularia guiensis*
  75. Nodularia micheloti*
  1. Nodularia nuxpersicae*
  2. Nyeinchanconcha nyeinchani*
  3. Physunio massini*
  4. Physunio modelli*
  5. Physunio pellucidus*
  6. Physunio superbus
  7. Pilsbryoconcha acuta*
  8. Pilsbryoconcha carinifera*
  9. Pilsbryoconcha exilis
  10. Pilsbryoconcha hoikaab*
  11. Pilsbryoconcha kittitati*
  12. Pilsbryoconcha linguaeformis*
  13. Pilsbryoconcha mekongiana*
  14. Pilsbryoconcha schomburgki*
  15. Pletholophus guangzhouensis*
  16. Pletholophus honglinhensis*
  17. Pletholophus tenuis
  18. Postolata guangxiensis*
  19. Postolata longjiangensis*
  20. Protunio messageri*
  21. Pseudobaphia banggiangensis*
  22. Pseudocuneopsis wuana*
  23. Pseudocuneopsis yangshuoensis*
  24. Pseudodon cambodjensis*
  25. Pseudodon mekongi*
  26. Pseudodon vagulus*
  27. Pseudodon vondembuschianus
  28. Ptychorhynchus liuovata
  29. Ptychorhynchus pfisteri
  30. Ptychorhynchus schoedei*
  31. Ptychorhynchus schomburgianum
  32. Rectidens sumatrensis
  33. Rhombuniopsis fultoni*
  34. Rhombuniopsis heres*
  35. Rhombuniopsis linan*
  36. Rhombuniopsis songmeng*
  37. Rhombuniopsis tauriformis*
  38. Scabiellus songkramensis*
  39. Scabies anceps*
  40. Scabies crispata
  41. Scabies humilis*
  42. Scabies mandarinus*
  43. Scabies nucleus*
  44. Scabies phaselus*
  45. Scabies pilata*
  46. Scabies scobinatus*
  47. Sinanodonta elliptica
  48. Sinanodonta jourdyi*
  49. Sinanodonta lucida
  50. Sinanodonta pacifica
  51. Sinohyriopsis cumingii
  52. Sinohyriopsis goliath*
  53. Sinosolenaia recognita
  54. Solenaia emarginata*
  55. Songkhlanaia moreleti*
  56. Songkhlanaia songkhramensis*
  57. Songkhlanaia tamodienica*
  58. Sundadontina brandti*
  59. Sundadontina cumingii*
  60. Sundadontina harmandi*
  61. Sundadontina mabilli*
  62. Sundadontina ovalis*
  63. Sundadontina plugpomenica*
  64. Sundadontina ponderosa*
  65. Sundadontina sulcata*
  66. Sundadontina taskaevi*
  67. Sundadontina tumida*
  68. Thaiconcha callifera*
  69. Thaiconcha munelliptica*
  70. Thaiconcha thaiensis*
  71. Trapezoideus foliaceus*
  72. Trapezoideus subviridis*
  73. Trapezoideus thachiadensis*
  74. Unionetta fabagina*
Sunda-Philippines (AS6)
  1. Caudiculatus caudiculatus*
  2. Ctenodesma bersinara*
  3. Ctenodesma borneensis*
  4. Ctenodesma mawonae*
  5. Ctenodesma scheibeneri*
  6. Discomya radulosa*
  7. Elongaria orientalis*
  8. Elongaria trompi*
  9. Hyriopsis velthuizeni*
  10. Khairuloconcha lunbawangorum*
  11. Khairuloconcha sahanae*
  12. Lens contradens
  13. Lens lugens*
  14. Lens semmelinki*
  15. Physunio superbus
  16. Pilsbryoconcha exilis
  1. Pilsbryoconcha expressa*
  2. Pressidens exanthematicus*
  3. Prohyriopsis stolata*
  4. Pseudodon aeneolus*
  5. Pseudodon inoscularis*
  6. Pseudodon vondembuschianus
  7. Pseudodon walpolei*
  8. Rectidens lauris*
  9. Rectidens sumatrensis
  10. Schepmania nieuwenhuisi*
  11. Schepmania parcesculpta*
  12. Simpsonella burroughiana*
  13. Simpsonella gracilis*
  14. Sinanodonta lauta
  15. Sinanodonta pacifica
India-Myanmar (AS7)
  1. Arcidopsis footei*
  2. Assamnaia involuta*
  3. Balwantia soleniformis*
  4. Indochinella pugio*
  5. Indonaia andersoniana*
  6. Indonaia bonneaudii*
  7. Indonaia caerulea*
  8. Indonaia corbis*
  9. Indonaia cylindrica*
  10. Indonaia olivaria*
  11. Indonaia rugosa*
  12. Indonaia shurtleffiana*
  13. Indonaia subclathrata*
  14. Indonaia theobaldi*
  15. Indopseudodon bogani*
  16. Indopseudodon crebristriatus*
  17. Indopseudodon indawgyiensis*
  18. Indopseudodon kayinensis*
  19. Indopseudodon salwenianus*
  20. Lamellidens candaharicus*
  21. Lamellidens chindwinensis*
  22. Lamellidens corrianus*
  23. Lamellidens ferrugineus*
  24. Lamellidens friersoni*
  25. Lamellidens generosus*
  26. Lamellidens jenkinsianus*
  27. Lamellidens lamellatus*
  28. Lamellidens mainwaringi*
  29. Lamellidens marginalis*
  30. Lamellidens nongyangensis
  31. Lamellidens savadiensis*
  32. Lamellidens unioides*
  33. Leoparreysia burmana*
  34. Leoparreysia canefrii*
  35. Leoparreysia olivacea*
  36. Leoparreysia perconvexa*
  37. Leoparreysia tavoyensis*
  38. Leoparreysia vulcanus*
  39. Leoparreysia whitteni*
  40. Parreysia corrugata*
  41. Parreysia keralaensis*
  1. Parreysia marcens*
  2. Parreysia nagpoorensis*
  3. Parreysia plagiosoma*
  4. Parreysia rajahensis*
  5. Parreysia rakhinensis*
  6. Parreysia smaragdites*
  7. Pilsbryoconcha exilis
  8. Pseudodon lenyanensis*
  9. Pseudodon nicobaricus*
  10. Pseudomulleria dalyi*
  11. Pseudoscabies scobina*
  12. Radiatula chaudhurii*
  13. Radiatula chindwinensis*
  14. Radiatula crispisulcata*
  15. Radiatula kachinensis*
  16. Radiatula mouhoti*
  17. Radiatula myitkyinae*
  18. Radiatula myitthanensis*
  19. Scabies crispata
  20. Sinanodonta pacifica
  21. Sinanodonta woodiana
  22. Sundadontina tanintharyiensis*
  23. Trapezidens angustior*
  24. Trapezidens dolichorhynchus*
  25. Trapezidens exolescens*
  26. Trapezidens mogaungensis*
  27. Trapezidens scutum*
  28. Trapezidens yeti*
  29. Trapezoideus lenya*
  30. Trapezoideus mitanensis*
  31. Yaukthwa avaensis*
  32. Yaukthwa baniensis*
  33. Yaukthwa dalliana*
  34. Yaukthwa elongatula*
  35. Yaukthwa inlenensis*
  36. Yaukthwa nesemanni*
  37. Yaukthwa paiensis*
  38. Yaukthwa panhai*
  39. Yaukthwa peguensis*
  40. Yaukthwa rectangularis*
  41. Yaukthwa zayleymanensis*
 
NSF icon MUSSEL icon
"Making the world a better place, one mollusk at a time."