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Page last updated
Thu 3 September 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Search the database pages:

Bivalvia | Unionida | Unionoidea | Unionidae | Unioninae | Unionini

subtribe ACULAMPROTULINA Huang & Wu in Huang et al., 2019: cladomics
     1 genus

Cladomics Summary

number of species = 9 (9 total)
number of subtaxa (rank: genus) = 1
analyzed in 114 cladograms in 64 publications
maximum number of terminals in an analysis = 6 (mean = 2.68, median = 2)
maximum number of subtaxa (genus) = 1 (mean = 1, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 67
maximum number of included terminals = 6, (mean = 3.87 median = 4)
maximum number of subtaxa (genus) = 1 (mean = 1, median = 1)
proportion of cladograms that support basic monophyly = 1


Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.


PCoA Ordination


NJ Tree


Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
66516256140
  +----------- Globunionina      
--+                              
  |  +-------- Scriptolamprotula ** 
  +--+                           
     |  +----- Middendorffinaiina
     +--+                        
        |  +-- Unionina          
        +--+                     
           +-- Oxynaiina         

Wu et al., 2020 [Publ 4121], fig. 3

iTtTbiriSoTroTroSiD
553144924
  +----- Scriptolamprotula **
--+                       
  |  +-- Unionina         
  +--+                    
     +-- Oxynaiina        

Wu et al., 2018 [Publ 3836], fig. 2

iTtTbiriSoTroTroSiD
553148926
  +----- Scriptolamprotula **
--+                       
  |  +-- Unionina         
  +--+                    
     +-- Oxynaiina        

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

iTtTbiriSoTroTroSiD
55311364244
  +----------- Scriptolamprotula ** 
--+                              
  |  +-------- Globunionina      
  +--+                           
     |  +----- Oxynaiina         
     +--+                        
        |  +-- Middendorffinaiina
        +--+                     
           +-- Unionina          

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iTtTbiriSoTroTroSiD
5531911824
  +----- Scriptolamprotula **
--+                       
  |  +-- Unionina         
  +--+                    
     +-- Oxynaiina        

Wu et al., 2021 [Publ 4366], fig. 3-BI

iTtTbiriSoTroTroSiD
5531691124
  +----- Scriptolamprotula **
--+                       
  |  +-- Unionina         
  +--+                    
     +-- Oxynaiina        

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iTtTbiriSoTroTroSiD
5531871824
  +----- Scriptolamprotula **
--+                       
  |  +-- Unionina         
  +--+                    
     +-- Oxynaiina        

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
55211951625
  +----- Scriptolamprotula **
--+                       
  |  +-- Unionina         
  +--+                    
     +-- Oxynaiina        

Wu et al., 2026 [Publ 4811], fig. 4

iTtTbiriSoTroTroSiD
5531902844
  +----------- Globunionina      
--+                              
  |  +-------- Scriptolamprotula ** 
  +--+                           
     |  +----- Unionina          
     +--+                        
        |  +-- Middendorffinaiina
        +--+                     
           +-- Oxynaiina         

Lopes-Lima et al., 2020 [Publ 4080], fig. 5

iTtTbiriSoTroTroSiD
55314212
  +-- Oxynaiina        
--+                    
  +-- Scriptolamprotula **

Wu et al., 2022 [Publ 4238], fig. 3

iTtTbiriSoTroTroSiD
5531691624
  +----- Scriptolamprotula **
--+                       
  |  +-- Unionina         
  +--+                    
     +-- Oxynaiina        

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
55311931524
  +----- Scriptolamprotula **
--+                       
  |  +-- Unionina         
  +--+                    
     +-- Oxynaiina        

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iTtTbiriSoTroTroSiD
55311172336
  +-------- Scriptolamprotula ** 
--+                           
  |  +----- Middendorffinaiina
  +--+                        
     |  +-- Unionina          
     +--+                     
        +-- Oxynaiina         

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iTtTbiriSoTroTroSiD
5531911824
  +----- Scriptolamprotula **
--+                       
  |  +-- Unionina         
  +--+                    
     +-- Oxynaiina        

Dai et al., 2025 [Publ 4440], fig. 1

iTtTbiriSoTroTroSiD
5521401743
  +----------------------- Globunionina      
  |                                          
--+  +-------------------- Scriptolamprotula ** 
  |  |                                       
  +--+  +----------------- Middendorffinaiina
     |  |                                    
     +--+  +-------------- Nodularia         
        |  |                                 
        +--+  +----------- Cuneopsis         
           |  |                              
           |  |        +-- Diaurora          
           +--+--------+                     
              |        +-- Schistodesmus     
              |                              
              |  +-------- Pseudocuneopsis   
              +--+                           
                 |  +----- Unionina          
                 +--+                        
                    |  +-- Tchangsinaia      
                    +--+                     
                       +-- Inversiunio       

Hou et al., 2025 [Publ 4445], fig. 5

iTtTbiriSoTroTroSiD
5531482824
  +----- Scriptolamprotula **
--+                       
  |  +-- Unionina         
  +--+                    
     +-- Oxynaiina        

Huang et al., 2019 [Publ 3843], fig. S1

iTtTbiriSoTroTroSiD
5531421212
  +-- Scriptolamprotula **
--+                    
  +-- Oxynaiina        

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
55311951624
1

Wu et al., 2021 [Publ 4366], fig. 3-ML

iTtTbiriSoTroTroSiD
5531691124
  +----- Scriptolamprotula **
--+                       
  |  +-- Unionina         
  +--+                    
     +-- Oxynaiina        

Hou et al., 2024 [Publ 4400], fig. 4

iTtTbiriSoTroTroSiD
5531714434
  +-------- Scriptolamprotula ** 
--+                           
  |  +----- Unionina          
  +--+                        
     |  +-- Middendorffinaiina
     +--+                     
        +-- Oxynaiina         

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iTtTbiriSoTroTroSiD
5531871824
  +----- Scriptolamprotula **
--+                       
  |  +-- Unionina         
  +--+                    
     +-- Oxynaiina        

Dai et al., 2024 [Publ 4386], fig. 6

iTtTbiriSoTroTroSiD
5531371724
  +----- Scriptolamprotula **
--+                       
  |  +-- Unionina         
  +--+                    
     +-- Oxynaiina        

Dai et al., 2025 [Publ 4440], fig. 2

iTtTbiriSoTroTroSiD
5531401744
  +-------------------- Globunionina      
  |                                       
--+  +----------------- Scriptolamprotula ** 
  |  |                                    
  |  |              +-- Middendorffinaiina
  +--+  +-----------+                     
     |  |           +-- Nodularia         
     |  |                                 
     +--+  +----------- Cuneopsis         
        |  |                              
        |  |        +-- Diaurora          
        +--+--------+                     
           |        +-- Schistodesmus     
           |                              
           |  +-------- Pseudocuneopsis   
           +--+                           
              |  +----- Inversiunio       
              +--+                        
                 |  +-- Tchangsinaia      
                 +--+                     
                    +-- Unionina          

Wu et al., 2021 [Publ 4366], fig. 4

iTtTbiriSoTroTroSiD
5531641124
  +----- Scriptolamprotula **
--+                       
  |  +-- Unionina         
  +--+                    
     +-- Oxynaiina        

Wu et al., 2024 [Publ 4379], fig. 5

iTtTbiriSoTroTroSiD
5531492524
  +----- Scriptolamprotula **
--+                       
  |  +-- Unionina         
  +--+                    
     +-- Oxynaiina        

Dai et al., 2025 [Publ 4440], fig. 3

iTtTbiriSoTroTroSiD
5531832334
  +-------- Globunionina     
--+                          
  |  +----- Scriptolamprotula **
  +--+                       
     |  +-- Unionina         
     +--+                    
        +-- Oxynaiina        

Chen et al., 2025 [Publ 4448], fig. 3

iTtTbiriSoTroTroSiD
5531812324
  +----- Scriptolamprotula **
--+                       
  |  +-- Unionina         
  +--+                    
     +-- Oxynaiina        

Wang et al., 2025 [Publ 4488], fig. 3B

iTtTbiriSoTroTroSiD
5531602834
  +-------- Scriptolamprotula ** 
--+                           
  |  +----- Unionina          
  +--+                        
     |  +-- Middendorffinaiina
     +--+                     
        +-- Oxynaiina         

Huang et al., 2019 [Publ 3843], fig. 1

iTtTbiriSoTroTroSiD
5531401216
1

Wu et al., 2022 [Publ 4238], fig. 2d

iTtTbiriSoTroTroSiD
5531341924
  +----- Scriptolamprotula **
--+                       
  |  +-- Unionina         
  +--+                    
     +-- Oxynaiina        

Konopleva et al., 2023 [Publ 4269], fig. S1_4

iTtTbiriSoTroTroSiD
442152835
  +-------- Middendorffinaiina
--+                           
  |  +----- Scriptolamprotula ** 
  +--+                        
     |  +-- Unionina          
     +--+                     
        +-- Oxynaiina         

Wu et al., 2020 [Publ 4121], fig. 2

iTtTbiriSoTroTroSiD
442130115
  +-- Unionina         
--+                    
  +-- Scriptolamprotula **

Wu et al., 2019 [Publ 3864], fig. 3

iTtTbiriSoTroTroSiD
442142925
  +----- Scriptolamprotula **
--+                       
  |  +-- Unionina         
  +--+                    
     +-- Oxynaiina        

Wu et al., 2020 [Publ 4121], fig. S1

iTtTbiriSoTroTroSiD
442130115
  +-- Unionina         
--+                    
  +-- Scriptolamprotula **

Huang et al., 2019 [Publ 3843], fig. 3

iTtTbiriSoTroTroSiD
442152725
  +----- Scriptolamprotula **
--+                       
  |  +-- Oxynaiina        
  +--+                    
     +-- Unionina         

Wu et al., 2020 [Publ 4147], fig. 3

iTtTbiriSoTroTroSiD
442126725
  +----- Scriptolamprotula **
--+                       
  |  +-- Unionina         
  +--+                    
     +-- Oxynaiina        

Wu et al., 2018 [Publ 4143], fig. 2

iTtTbiriSoTroTroSiD
442118225
  +----- Scriptolamprotula **
--+                       
  |  +-- Unionina         
  +--+                    
     +-- Oxynaiina        

Wu et al., 2020 [Publ 4147], fig. 4

iTtTbiriSoTroTroSiD
442110115
  +-- Unionina         
--+                    
  +-- Scriptolamprotula **

Froufe et al., 2019 [Publ 3936], fig. 3

iTtTbiriSoTroTroSiD
331141724
  +----- Scriptolamprotula **
--+                       
  |  +-- Oxynaiina        
  +--+                    
     +-- Unionina         

Sano et al., 2020 [Publ 4138], fig. 3

iTtTbiriSoTroTroSiD
3311651024
  +----- Scriptolamprotula **
--+                       
  |  +-- Unionina         
  +--+                    
     +-- Oxynaiina        

Soroka & Burzynski, 2017 [Publ 3837], fig. 1

iTtTbiriSoTroTroSiD
331120724
  +----- Scriptolamprotula **
--+                       
  |  +-- Oxynaiina        
  +--+                    
     +-- Unionina         

Sano et al., 2020 [Publ 4138], fig. 2

iTtTbiriSoTroTroSiD
3311651024
  +----- Scriptolamprotula **
--+                       
  |  +-- Unionina         
  +--+                    
     +-- Oxynaiina        

Froufe et al., 2019 [Publ 3936], fig. 6

iTtTbiriSoTroTroSiD
331141724
  +----- Scriptolamprotula **
--+                       
  |  +-- Oxynaiina        
  +--+                    
     +-- Unionina         

Zhou et al., 2007 [Publ 2658], fig. 2

iTtTbiriSoTroTroSiD
331115214
1

Fukata & Iigo, 2020 [Publ 4106], fig. 1

iTtTbiriSoTroTroSiD
3311511024
  +----- Scriptolamprotula **
--+                       
  |  +-- Unionina         
  +--+                    
     +-- Oxynaiina        

Wu et al., 2021 [Publ 4366], fig. 6

iTtTbiriSoTroTroSiD
331188824
  +----- Scriptolamprotula **
--+                       
  |  +-- Unionina         
  +--+                    
     +-- Oxynaiina        

Cha et al., 2018 [Publ 3916], fig. 1

iTtTbiriSoTroTroSiD
331135724
  +----- Scriptolamprotula **
--+                       
  |  +-- Oxynaiina        
  +--+                    
     +-- Unionina         

Aunins et al., 2018 [Publ 3863], fig. 1

iTtTbiriSoTroTroSiD
331137824
  +----- Scriptolamprotula **
--+                       
  |  +-- Oxynaiina        
  +--+                    
     +-- Unionina         

Huang et al., 2018 [Publ 3828], fig. 6

iTtTbiriSoTroTroSiD
331139624
  +----- Scriptolamprotula **
--+                       
  |  +-- Oxynaiina        
  +--+                    
     +-- Unionina         

Sano et al., 2020 [Publ 4138], fig. 4

iTtTbiriSoTroTroSiD
3311651024
  +----- Scriptolamprotula **
--+                       
  |  +-- Unionina         
  +--+                    
     +-- Oxynaiina        

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

iTtTbiriSoTroTroSiD
331128224
  +----- Scriptolamprotula **
--+                       
  |  +-- Unionina         
  +--+                    
     +-- Oxynaiina        

Wu et al., 2016 [Publ 3711], fig. 1

iTtTbiriSoTroTroSiD
331118224
  +----- Scriptolamprotula **
--+                       
  |  +-- Unionina         
  +--+                    
     +-- Oxynaiina        

Zhou et al., 2007 [Publ 2658], fig. 1

iTtTbiriSoTroTroSiD
331118414
  +-- Oxynaiina        
--+                    
  +-- Scriptolamprotula **

Wu et al., 2024 [Publ 4379], fig. 4_6

iTtTbiriSoTroTroSiD
3311201214
1

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

iTtTbiriSoTroTroSiD
331145724
  +----- Scriptolamprotula **
--+                       
  |  +-- Oxynaiina        
  +--+                    
     +-- Unionina         

Wu et al., 2021 [Publ 4366], fig. 2_I

iTtTbiriSoTroTroSiD
331188824
  +----- Scriptolamprotula **
--+                       
  |  +-- Unionina         
  +--+                    
     +-- Oxynaiina        

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

iTtTbiriSoTroTroSiD
22013752--
1

Liu et al., 2023 [Publ 4310], fig. 5

iTtTbiriSoTroTroSiD
220122122--
  +-- Unionina         
  |                    
--+-- Scriptolamprotula **
  |                    
  +-- Oxynaiina        

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

iTtTbiriSoTroTroSiD
22013552--
  +----- Scriptolamprotula **
--+                       
  |  +-- Unionina         
  +--+                    
     +-- Oxynaiina        

Lopes-Lima et al., 2017 [Publ 3792], fig. 2

iTtTbiriSoTroTroSiD
22013052--
  +----- Scriptolamprotula **
--+                       
  |  +-- Unionina         
  +--+                    
     +-- Oxynaiina        

Chen et al., 2020 [Publ 4170], fig. 1

iTtTbiriSoTroTroSiD
22013211--
1

Liu et al., 2023 [Publ 4310], fig. 4

iTtTbiriSoTroTroSiD
220122122--
  +----- Unionina         
--+                       
  |  +-- Scriptolamprotula **
  +--+                    
     +-- Oxynaiina        

Zhou et al., 2017 [Publ 3788], fig. 2

iTtTbiriSoTroTroSiD
22011421--
  +-- Oxynaiina        
--+                    
  +-- Scriptolamprotula **

Wen et al., 2017 [Publ 3795], fig. 3

iTtTbiriSoTroTroSiD
22011411--
1

Dai et al., 2024 [Publ 4358], fig. 4

iTtTbiriSoTroTroSiD
220151162--
  +----- Scriptolamprotula **
--+                       
  |  +-- Unionina         
  +--+                    
     +-- Oxynaiina        

Wu et al., 2023 [Publ 4296], fig. 4

iTtTbiriSoTroTroSiD
220121112--
  +----- Unionina         
--+                       
  |  +-- Scriptolamprotula **
  +--+                    
     +-- Oxynaiina        

Fonseca et al., 2016 [Publ 3775], fig. 1

iTtTbiriSoTroTroSiD
22012952--
  +----- Scriptolamprotula **
--+                       
  |  +-- Unionina         
  +--+                    
     +-- Oxynaiina        

Dai et al., 2024 [Publ 4386], fig. 5-BI

iTtTbiriSoTroTroSiD
110121113--
-- Scriptolamprotula_scripta **

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

iTtTbiriSoTroTroSiD
11017592--
  +----- Unionina         
--+                       
  |  +-- Scriptolamprotula **
  +--+                    
     +-- Oxynaiina        

Chen et al., 2025 [Publ 4487], fig. 2

iTtTbiriSoTroTroSiD
110140173--
  +----- Scriptolamprotula **
--+                       
  |  +-- Unionina         
  +--+                    
     +-- Oxynaiina        

Chen et al., 2023 [Publ 4302], fig. 2

iTtTbiriSoTroTroSiD
110140173--
  +----------------- Scriptolamprotula ** 
  |                                    
  |              +-- Middendorffinaiina
--+  +-----------+                     
  |  |           +-- Nodularia         
  |  |                                 
  +--+  +----------- Cuneopsis         
     |  |                              
     |  |        +-- Diaurora          
     +--+--------+                     
        |        +-- Schistodesmus     
        |                              
        |  +-------- Pseudocuneopsis   
        +--+                           
           |  +----- Inversiunio       
           +--+                        
              |  +-- Tchangsinaia      
              +--+                     
                 +-- Unionina          

Xue et al., 2016 [Publ 3659], fig. 1

iTtTbiriSoTroTroSiD
11011432--
  +-------- Scriptolamprotula **
--+                          
  |  +----- Schistodesmus    
  +--+                       
     |  +-- Nodularia        
     +--+                    
        +-- Unionina         

Sonowal et al., 2021 [Publ 4221], fig. 3a

iTtTbiriSoTroTroSiD
11011421--
-- Scriptolamprotula_tientsinensis **

Guerra et al., 2019 [Publ 4168], fig. 1-M

iTtTbiriSoTroTroSiD
11012142--
  +----- Oxynaiina        
--+                       
  |  +-- Scriptolamprotula **
  +--+                    
     +-- Unionina         

Chen et al., 2025 [Publ 4487], fig. 3

iTtTbiriSoTroTroSiD
110140173--
  +-------- Globunionina     
--+                          
  |  +----- Scriptolamprotula **
  +--+                       
     |  +-- Unionina         
     +--+                    
        +-- Oxynaiina        

Bolotov et al., 2017 [Publ 3761], fig. S2

iTtTbiriSoTroTroSiD
110113792--
  +----------- Unionina            
  |                                
--+  +-------- Scriptolamprotula **   
  |  |                             
  |  +-------- Schistodesmus       
  +--+                             
     +-------- Nodularia           
     |                             
     |  +----- Cuneopsis_szechenyii
     +--+                          
        |  +-- Tchangsinaia        
        +--+                       
           +-- Cuneopsis_rufescens 

Bolotov et al., 2017 [Publ 3764], fig. S1

iTtTbiriSoTroTroSiD
110112392--
  +-- Unio_crassus         
  |                        
  +-- Unio_pictorum        
  |                        
  +-- Unio_tumidus         
  |                        
  +-- Scriptolamprotula **    
  |                        
  +-- Cuneopsis_szechenyii 
--+                        
  +-- Tchangsinaia         
  |                        
  +-- Cuneopsis_rufescens  
  |                        
  +-- Nodularia_douglasiae 
  |                        
  +-- Nodularia_nuxpersicae
  |                        
  +-- Schistodesmus        

Sonowal et al., 2021 [Publ 4221], fig. 3b

iTtTbiriSoTroTroSiD
11011421--
-- Scriptolamprotula_tientsinensis **

Soroka & Burzynski, 2018 [Publ 3762], fig. 3

iTtTbiriSoTroTroSiD
1101611--
  +-- Unionina         
--+                    
  +-- Scriptolamprotula **

Wu et al., 2022 [Publ 4238], fig. 2e

iTtTbiriSoTroTroSiD
110117122--
  +----- Scriptolamprotula **
--+                       
  |  +-- Unionina         
  +--+                    
     +-- Oxynaiina        

Burzynski & Soroka, 2018 [Publ 3862], fig. 5a

iTtTbiriSoTroTroSiD
11011941--
  +-- Scriptolamprotula **
--+                    
  +-- Unionina         

Guerra et al., 2019 [Publ 4168], fig. 2

iTtTbiriSoTroTroSiD
11014042--
-- Scriptolamprotula_tortuosa **

Wu et al., 2022 [Publ 4238], fig. 2a_S1

iTtTbiriSoTroTroSiD
110111111--
  +-- Scriptolamprotula **
--+                    
  +-- Oxynaiina        

Dai et al., 2024 [Publ 4386], fig. 3

iTtTbiriSoTroTroSiD
110113103--
-- Scriptolamprotula_scripta **

Burzynski et al., 2017 [Publ 3774], fig. 1-top

iTtTbiriSoTroTroSiD
11011221--
  +-- Scriptolamprotula **
--+                    
  +-- Unionina         

Guerra et al., 2017 [Publ 3772], fig. 2b

iTtTbiriSoTroTroSiD
11012411--
-- Scriptolamprotula_tortuosa **

Wang et al., 2018 [Publ 3865], fig. 2

iTtTbiriSoTroTroSiD
11013062--
  +----- Scriptolamprotula **
--+                       
  |  +-- Oxynaiina        
  +--+                    
     +-- Unionina         

Burzynski et al., 2017 [Publ 3774], fig. 1-bottom

iTtTbiriSoTroTroSiD
11011221--
  +-- Scriptolamprotula **
--+                    
  +-- Unionina         

Araujo et al., 2017 [Publ 3801], fig. 2

iTtTbiriSoTroTroSiD
11011322--
  +-- Oxynaiina        
--+                    
  +-- Scriptolamprotula **

Dai et al., 2024 [Publ 4386], fig. 5-ML

iTtTbiriSoTroTroSiD
110121113--
  +-------- Scriptolamprotula ** 
--+                           
  |  +----- Unionina          
  +--+                        
     |  +-- Middendorffinaiina
     +--+                     
        +-- Oxynaiina         

Kim et al., 2021 [Publ 4197], fig. 1

iTtTbiriSoTroTroSiD
110117112--
-- Scriptolamprotula_tientsinensis **

Guerra et al., 2019 [Publ 4168], fig. 2-M

iTtTbiriSoTroTroSiD
11012142--
  +----- Scriptolamprotula **
--+                       
  |  +-- Oxynaiina        
  +--+                    
     +-- Unionina         

Chen et al., 2023 [Publ 4302], fig. 1

iTtTbiriSoTroTroSiD
110140173--
  +----------------- Scriptolamprotula ** 
  |                                    
  |              +-- Middendorffinaiina
--+  +-----------+                     
  |  |           +-- Nodularia         
  |  |                                 
  +--+  +----------- Cuneopsis         
     |  |                              
     |  |        +-- Diaurora          
     +--+--------+                     
        |        +-- Schistodesmus     
        |                              
        |  +-------- Pseudocuneopsis   
        +--+                           
           |  +----- Inversiunio       
           +--+                        
              |  +-- Tchangsinaia      
              +--+                     
                 +-- Unionina          

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

iTtTbiriSoTroTroSiD
11017592--
  +----- Unionina         
--+                       
  |  +-- Scriptolamprotula **
  +--+                    
     +-- Oxynaiina        

Campbell & Lydeard, 2026 [Publ 4810], fig. 4

iTtTbiriSoTroTroSiD
110146132--
  +----- Scriptolamprotula **
--+                       
  |  +-- Unionina         
  +--+                    
     +-- Oxynaiina        

Huang et al., 2018 [Publ 3828], fig. 3_5

iTtTbiriSoTroTroSiD
11012811--
  +-- Unionina         
--+                    
  +-- Scriptolamprotula **

Fukata & Iigo, 2019 [Publ 4108], fig. 1

iTtTbiriSoTroTroSiD
110116122--
-- Scriptolamprotula_tientsinensis **

Burzynski & Soroka, 2018 [Publ 3862], fig. 5b

iTtTbiriSoTroTroSiD
1101941--
  +-- Scriptolamprotula **
--+                    
  +-- Unionina         

Rosenberg et al., 1997 [Publ 2720], fig. 2B

iTtTbiriSoTroTroSiD
11012111--
-- Scriptolamprotula_tortuosa **

Dai et al., 2023 [Publ 4285], fig. 5

iTtTbiriSoTroTroSiD
11015262--
  +----- Scriptolamprotula **
--+                       
  |  +-- Unionina         
  +--+                    
     +-- Oxynaiina        

Ouyang et al., 2015 [Publ 4493], fig. 1

iTtTbiriSoTroTroSiD
11013071--
           +-- Schistodesmus        
  +--------+                        
  |        +-- Scriptolamprotula **    
  |                                 
--+        +-- Tchangsinaia         
  |  +-----+                        
  |  |     +-- Cuneopsis_rufescens  
  +--+                              
     |  +----- Nodularia            
     +--+                           
        |  +-- Cuneopsis_szechenyii 
        +--+                        
           +-- Cuneopsis_celtiformis

Ouyang et al., 2011 [Publ 3408], fig. 1

iTtTbiriSoTroTroSiD
11011221--
-- Scriptolamprotula_tortuosa **

Wu et al., 2024 [Publ 4388], fig. 6

iTtTbiriSoTroTroSiD
11015282--
  +----- Scriptolamprotula **
--+                       
  |  +-- Unionina         
  +--+                    
     +-- Oxynaiina        

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iTtTbiriSoTroTroSiD
110179102--
  +----- Unionina         
--+                       
  |  +-- Scriptolamprotula **
  +--+                    
     +-- Oxynaiina        

Sonowal et al., 2021 [Publ 4221], fig. 4

iTtTbiriSoTroTroSiD
11011521--
-- Scriptolamprotula_tientsinensis **

Sano et al., 2020 [Publ 4138], fig. 1

iTtTbiriSoTroTroSiD
11013361--
-- Scriptolamprotula_tortuosa **

Guerra et al., 2019 [Publ 4168], fig. 1

iTtTbiriSoTroTroSiD
11014042--
  +----- Scriptolamprotula **
--+                       
  |  +-- Oxynaiina        
  +--+                    
     +-- Unionina         

Bolotov et al., 2017 [Publ 3761], fig. S1

iTtTbiriSoTroTroSiD
110113792--
  +----- Unionina         
--+                       
  |  +-- Scriptolamprotula **
  +--+                    
     +-- Oxynaiina        

Soroka & Burzynski, 2018 [Publ 3762], fig. 3-M

iTtTbiriSoTroTroSiD
1101611--
  +-- Unionina         
--+                    
  +-- Scriptolamprotula **

Guerra et al., 2017 [Publ 3772], fig. 2a

iTtTbiriSoTroTroSiD
11012411--
  +-- Scriptolamprotula **
--+                    
  +-- Unionina         

Ouyang et al., 2015 [Publ 4493], fig. 2

iTtTbiriSoTroTroSiD
11013071--
-- Scriptolamprotula_tortuosa **

Wang et al., 2018 [Publ 3846], fig. 2

iTtTbiriSoTroTroSiD
11013072--
  +----- Scriptolamprotula **
--+                       
  |  +-- Oxynaiina        
  +--+                    
     +-- Unionina         

Huang et al., 2019 [Publ 3843], fig. 3-M

iTtTbiriSoTroTroSiD
11012242--
  +----- Oxynaiina        
--+                       
  |  +-- Scriptolamprotula **
  +--+                    
     +-- Unionina         

Huang et al., 2018 [Publ 3828], fig. 6-M

iTtTbiriSoTroTroSiD
11011631--
  +-- Scriptolamprotula **
--+                    
  +-- Unionina         
 
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