The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)
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Bivalvia | Unionida | Unionoidea | Unionidae | Unioninae | Unionini subtribe ACULAMPROTULINA Huang & Wu in Huang et al., 2019: cladomics 1 genus Cladomics Summarynumber of species = 9 (9 total) number of subtaxa (rank: genus) = 1 analyzed in 114 cladograms in 64 publications maximum number of terminals in an analysis =
6 (mean = 2.68, median = 2) maximum number of subtaxa (genus) =
1 (mean = 1, median = 1) Basic Monophylynumber of cladograms that test basic monophyly
(i.e., more than 1 included species) = 67 maximum number of included terminals = 6, (mean =
3.87 median = 4) maximum number of subtaxa (genus) =
1 (mean = 1, median = 1) proportion of cladograms that support basic monophyly
= 1
Pairwise Cladogram ClusteringThe graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.
Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14 ** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram ** | iT | tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 5 | 1 | 625 | 61 | 4 | 0 |
+----------- Globunionina
--+
| +-------- Scriptolamprotula **
+--+
| +----- Middendorffinaiina
+--+
| +-- Unionina
+--+
+-- Oxynaiina
Wu et al., 2020 [Publ 4121], fig. 3 | iT | tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 1 | 44 | 9 | 2 | 4 |
+----- Scriptolamprotula **
--+
| +-- Unionina
+--+
+-- Oxynaiina
Wu et al., 2018 [Publ 3836], fig. 2 | iT | tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 1 | 48 | 9 | 2 | 6 |
+----- Scriptolamprotula **
--+
| +-- Unionina
+--+
+-- Oxynaiina
Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5 | iT | tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 1 | 136 | 42 | 4 | 4 |
+----------- Scriptolamprotula **
--+
| +-------- Globunionina
+--+
| +----- Oxynaiina
+--+
| +-- Middendorffinaiina
+--+
+-- Unionina
Wu et al., 2023 [Publ 4339], fig. 4_S5-aa | iT | tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 1 | 91 | 18 | 2 | 4 |
+----- Scriptolamprotula **
--+
| +-- Unionina
+--+
+-- Oxynaiina
Wu et al., 2021 [Publ 4366], fig. 3-BI | iT | tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 1 | 69 | 11 | 2 | 4 |
+----- Scriptolamprotula **
--+
| +-- Unionina
+--+
+-- Oxynaiina
Wu et al., 2024 [Publ 4350], fig. 8b_S2 | iT | tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 1 | 87 | 18 | 2 | 4 |
+----- Scriptolamprotula **
--+
| +-- Unionina
+--+
+-- Oxynaiina
Huang et al., 2019 [Publ 3843], fig. 2 | iT | tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 2 | 1 | 195 | 16 | 2 | 5 |
+----- Scriptolamprotula **
--+
| +-- Unionina
+--+
+-- Oxynaiina
Wu et al., 2026 [Publ 4811], fig. 4 | iT | tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 1 | 90 | 28 | 4 | 4 |
+----------- Globunionina
--+
| +-------- Scriptolamprotula **
+--+
| +----- Unionina
+--+
| +-- Middendorffinaiina
+--+
+-- Oxynaiina
Lopes-Lima et al., 2020 [Publ 4080], fig. 5 | iT | tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 1 | 4 | 2 | 1 | 2 |
+-- Oxynaiina
--+
+-- Scriptolamprotula **
Wu et al., 2022 [Publ 4238], fig. 3 | iT | tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 1 | 69 | 16 | 2 | 4 |
+----- Scriptolamprotula **
--+
| +-- Unionina
+--+
+-- Oxynaiina
Huang et al., 2019 [Publ 3843], fig. S3 | iT | tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 1 | 193 | 15 | 2 | 4 |
+----- Scriptolamprotula **
--+
| +-- Unionina
+--+
+-- Oxynaiina
Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4 | iT | tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 1 | 117 | 23 | 3 | 6 |
+-------- Scriptolamprotula **
--+
| +----- Middendorffinaiina
+--+
| +-- Unionina
+--+
+-- Oxynaiina
Wu et al., 2023 [Publ 4339], fig. 4_S5-nt | iT | tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 1 | 91 | 18 | 2 | 4 |
+----- Scriptolamprotula **
--+
| +-- Unionina
+--+
+-- Oxynaiina
Dai et al., 2025 [Publ 4440], fig. 1 | iT | tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 2 | 1 | 40 | 17 | 4 | 3 |
+----------------------- Globunionina
|
--+ +-------------------- Scriptolamprotula **
| |
+--+ +----------------- Middendorffinaiina
| |
+--+ +-------------- Nodularia
| |
+--+ +----------- Cuneopsis
| |
| | +-- Diaurora
+--+--------+
| +-- Schistodesmus
|
| +-------- Pseudocuneopsis
+--+
| +----- Unionina
+--+
| +-- Tchangsinaia
+--+
+-- Inversiunio
Hou et al., 2025 [Publ 4445], fig. 5 | iT | tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 1 | 48 | 28 | 2 | 4 |
+----- Scriptolamprotula **
--+
| +-- Unionina
+--+
+-- Oxynaiina
Huang et al., 2019 [Publ 3843], fig. S1 | iT | tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 1 | 42 | 12 | 1 | 2 |
+-- Scriptolamprotula **
--+
+-- Oxynaiina
Huang et al., 2019 [Publ 3843], fig. S4 | iT | tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 1 | 195 | 16 | 2 | 4 |
Wu et al., 2021 [Publ 4366], fig. 3-ML | iT | tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 1 | 69 | 11 | 2 | 4 |
+----- Scriptolamprotula **
--+
| +-- Unionina
+--+
+-- Oxynaiina
Hou et al., 2024 [Publ 4400], fig. 4 | iT | tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 1 | 71 | 44 | 3 | 4 |
+-------- Scriptolamprotula **
--+
| +----- Unionina
+--+
| +-- Middendorffinaiina
+--+
+-- Oxynaiina
Wu et al., 2024 [Publ 4350], fig. 8a_S1 | iT | tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 1 | 87 | 18 | 2 | 4 |
+----- Scriptolamprotula **
--+
| +-- Unionina
+--+
+-- Oxynaiina
Dai et al., 2024 [Publ 4386], fig. 6 | iT | tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 1 | 37 | 17 | 2 | 4 |
+----- Scriptolamprotula **
--+
| +-- Unionina
+--+
+-- Oxynaiina
Dai et al., 2025 [Publ 4440], fig. 2 | iT | tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 1 | 40 | 17 | 4 | 4 |
+-------------------- Globunionina
|
--+ +----------------- Scriptolamprotula **
| |
| | +-- Middendorffinaiina
+--+ +-----------+
| | +-- Nodularia
| |
+--+ +----------- Cuneopsis
| |
| | +-- Diaurora
+--+--------+
| +-- Schistodesmus
|
| +-------- Pseudocuneopsis
+--+
| +----- Inversiunio
+--+
| +-- Tchangsinaia
+--+
+-- Unionina
Wu et al., 2021 [Publ 4366], fig. 4 | iT | tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 1 | 64 | 11 | 2 | 4 |
+----- Scriptolamprotula **
--+
| +-- Unionina
+--+
+-- Oxynaiina
Wu et al., 2024 [Publ 4379], fig. 5 | iT | tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 1 | 49 | 25 | 2 | 4 |
+----- Scriptolamprotula **
--+
| +-- Unionina
+--+
+-- Oxynaiina
Dai et al., 2025 [Publ 4440], fig. 3 | iT | tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 1 | 83 | 23 | 3 | 4 |
+-------- Globunionina
--+
| +----- Scriptolamprotula **
+--+
| +-- Unionina
+--+
+-- Oxynaiina
Chen et al., 2025 [Publ 4448], fig. 3 | iT | tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 1 | 81 | 23 | 2 | 4 |
+----- Scriptolamprotula **
--+
| +-- Unionina
+--+
+-- Oxynaiina
Wang et al., 2025 [Publ 4488], fig. 3B | iT | tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 1 | 60 | 28 | 3 | 4 |
+-------- Scriptolamprotula **
--+
| +----- Unionina
+--+
| +-- Middendorffinaiina
+--+
+-- Oxynaiina
Huang et al., 2019 [Publ 3843], fig. 1 | iT | tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 1 | 40 | 12 | 1 | 6 |
Wu et al., 2022 [Publ 4238], fig. 2d | iT | tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 1 | 34 | 19 | 2 | 4 |
+----- Scriptolamprotula **
--+
| +-- Unionina
+--+
+-- Oxynaiina
Konopleva et al., 2023 [Publ 4269], fig. S1_4 | iT | tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 1 | 52 | 8 | 3 | 5 |
+-------- Middendorffinaiina
--+
| +----- Scriptolamprotula **
+--+
| +-- Unionina
+--+
+-- Oxynaiina
Wu et al., 2020 [Publ 4121], fig. 2 | iT | tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 1 | 30 | 1 | 1 | 5 |
+-- Unionina
--+
+-- Scriptolamprotula **
Wu et al., 2019 [Publ 3864], fig. 3 | iT | tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 1 | 42 | 9 | 2 | 5 |
+----- Scriptolamprotula **
--+
| +-- Unionina
+--+
+-- Oxynaiina
Wu et al., 2020 [Publ 4121], fig. S1 | iT | tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 1 | 30 | 1 | 1 | 5 |
+-- Unionina
--+
+-- Scriptolamprotula **
Huang et al., 2019 [Publ 3843], fig. 3 | iT | tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 1 | 52 | 7 | 2 | 5 |
+----- Scriptolamprotula **
--+
| +-- Oxynaiina
+--+
+-- Unionina
Wu et al., 2020 [Publ 4147], fig. 3 | iT | tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 1 | 26 | 7 | 2 | 5 |
+----- Scriptolamprotula **
--+
| +-- Unionina
+--+
+-- Oxynaiina
Wu et al., 2018 [Publ 4143], fig. 2 | iT | tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 1 | 18 | 2 | 2 | 5 |
+----- Scriptolamprotula **
--+
| +-- Unionina
+--+
+-- Oxynaiina
Wu et al., 2020 [Publ 4147], fig. 4 | iT | tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 1 | 10 | 1 | 1 | 5 |
+-- Unionina
--+
+-- Scriptolamprotula **
Froufe et al., 2019 [Publ 3936], fig. 3 | iT | tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 41 | 7 | 2 | 4 |
+----- Scriptolamprotula **
--+
| +-- Oxynaiina
+--+
+-- Unionina
Sano et al., 2020 [Publ 4138], fig. 3 | iT | tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 65 | 10 | 2 | 4 |
+----- Scriptolamprotula **
--+
| +-- Unionina
+--+
+-- Oxynaiina
Soroka & Burzynski, 2017 [Publ 3837], fig. 1 | iT | tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 20 | 7 | 2 | 4 |
+----- Scriptolamprotula **
--+
| +-- Oxynaiina
+--+
+-- Unionina
Sano et al., 2020 [Publ 4138], fig. 2 | iT | tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 65 | 10 | 2 | 4 |
+----- Scriptolamprotula **
--+
| +-- Unionina
+--+
+-- Oxynaiina
Froufe et al., 2019 [Publ 3936], fig. 6 | iT | tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 41 | 7 | 2 | 4 |
+----- Scriptolamprotula **
--+
| +-- Oxynaiina
+--+
+-- Unionina
Zhou et al., 2007 [Publ 2658], fig. 2 | iT | tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 15 | 2 | 1 | 4 |
Fukata & Iigo, 2020 [Publ 4106], fig. 1 | iT | tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 51 | 10 | 2 | 4 |
+----- Scriptolamprotula **
--+
| +-- Unionina
+--+
+-- Oxynaiina
Wu et al., 2021 [Publ 4366], fig. 6 | iT | tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 88 | 8 | 2 | 4 |
+----- Scriptolamprotula **
--+
| +-- Unionina
+--+
+-- Oxynaiina
Cha et al., 2018 [Publ 3916], fig. 1 | iT | tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 35 | 7 | 2 | 4 |
+----- Scriptolamprotula **
--+
| +-- Oxynaiina
+--+
+-- Unionina
Aunins et al., 2018 [Publ 3863], fig. 1 | iT | tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 37 | 8 | 2 | 4 |
+----- Scriptolamprotula **
--+
| +-- Oxynaiina
+--+
+-- Unionina
Huang et al., 2018 [Publ 3828], fig. 6 | iT | tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 39 | 6 | 2 | 4 |
+----- Scriptolamprotula **
--+
| +-- Oxynaiina
+--+
+-- Unionina
Sano et al., 2020 [Publ 4138], fig. 4 | iT | tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 65 | 10 | 2 | 4 |
+----- Scriptolamprotula **
--+
| +-- Unionina
+--+
+-- Oxynaiina
Pfeiffer & Graf, 2013 [Publ 3531], fig. 2 | iT | tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 28 | 2 | 2 | 4 |
+----- Scriptolamprotula **
--+
| +-- Unionina
+--+
+-- Oxynaiina
Wu et al., 2016 [Publ 3711], fig. 1 | iT | tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 18 | 2 | 2 | 4 |
+----- Scriptolamprotula **
--+
| +-- Unionina
+--+
+-- Oxynaiina
Zhou et al., 2007 [Publ 2658], fig. 1 | iT | tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 18 | 4 | 1 | 4 |
+-- Oxynaiina
--+
+-- Scriptolamprotula **
Wu et al., 2024 [Publ 4379], fig. 4_6 | iT | tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 20 | 12 | 1 | 4 |
Zieritz et al., 2021 [Publ 4231], fig. 3-F_5 | iT | tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 45 | 7 | 2 | 4 |
+----- Scriptolamprotula **
--+
| +-- Oxynaiina
+--+
+-- Unionina
Wu et al., 2021 [Publ 4366], fig. 2_I | iT | tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 88 | 8 | 2 | 4 |
+----- Scriptolamprotula **
--+
| +-- Unionina
+--+
+-- Oxynaiina
Pavan-Kumar et al., 2022 [Publ 4263], fig. S5 | iT | tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 37 | 5 | 2 | -- |
Liu et al., 2023 [Publ 4310], fig. 5 | iT | tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 22 | 12 | 2 | -- |
+-- Unionina
|
--+-- Scriptolamprotula **
|
+-- Oxynaiina
Pavan-Kumar et al., 2022 [Publ 4263], fig. 4 | iT | tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 35 | 5 | 2 | -- |
+----- Scriptolamprotula **
--+
| +-- Unionina
+--+
+-- Oxynaiina
Lopes-Lima et al., 2017 [Publ 3792], fig. 2 | iT | tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 30 | 5 | 2 | -- |
+----- Scriptolamprotula **
--+
| +-- Unionina
+--+
+-- Oxynaiina
Chen et al., 2020 [Publ 4170], fig. 1 | iT | tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 32 | 1 | 1 | -- |
Liu et al., 2023 [Publ 4310], fig. 4 | iT | tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 22 | 12 | 2 | -- |
+----- Unionina
--+
| +-- Scriptolamprotula **
+--+
+-- Oxynaiina
Zhou et al., 2017 [Publ 3788], fig. 2 | iT | tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 14 | 2 | 1 | -- |
+-- Oxynaiina
--+
+-- Scriptolamprotula **
Wen et al., 2017 [Publ 3795], fig. 3 | iT | tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 14 | 1 | 1 | -- |
Dai et al., 2024 [Publ 4358], fig. 4 | iT | tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 51 | 16 | 2 | -- |
+----- Scriptolamprotula **
--+
| +-- Unionina
+--+
+-- Oxynaiina
Wu et al., 2023 [Publ 4296], fig. 4 | iT | tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 21 | 11 | 2 | -- |
+----- Unionina
--+
| +-- Scriptolamprotula **
+--+
+-- Oxynaiina
Fonseca et al., 2016 [Publ 3775], fig. 1 | iT | tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 29 | 5 | 2 | -- |
+----- Scriptolamprotula **
--+
| +-- Unionina
+--+
+-- Oxynaiina
Dai et al., 2024 [Publ 4386], fig. 5-BI | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 21 | 11 | 3 | -- |
-- Scriptolamprotula_scripta **
Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 75 | 9 | 2 | -- |
+----- Unionina
--+
| +-- Scriptolamprotula **
+--+
+-- Oxynaiina
Chen et al., 2025 [Publ 4487], fig. 2 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 40 | 17 | 3 | -- |
+----- Scriptolamprotula **
--+
| +-- Unionina
+--+
+-- Oxynaiina
Chen et al., 2023 [Publ 4302], fig. 2 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 40 | 17 | 3 | -- |
+----------------- Scriptolamprotula **
|
| +-- Middendorffinaiina
--+ +-----------+
| | +-- Nodularia
| |
+--+ +----------- Cuneopsis
| |
| | +-- Diaurora
+--+--------+
| +-- Schistodesmus
|
| +-------- Pseudocuneopsis
+--+
| +----- Inversiunio
+--+
| +-- Tchangsinaia
+--+
+-- Unionina
Xue et al., 2016 [Publ 3659], fig. 1 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 14 | 3 | 2 | -- |
+-------- Scriptolamprotula **
--+
| +----- Schistodesmus
+--+
| +-- Nodularia
+--+
+-- Unionina
Sonowal et al., 2021 [Publ 4221], fig. 3a | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 14 | 2 | 1 | -- |
-- Scriptolamprotula_tientsinensis **
Guerra et al., 2019 [Publ 4168], fig. 1-M | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 21 | 4 | 2 | -- |
+----- Oxynaiina
--+
| +-- Scriptolamprotula **
+--+
+-- Unionina
Chen et al., 2025 [Publ 4487], fig. 3 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 40 | 17 | 3 | -- |
+-------- Globunionina
--+
| +----- Scriptolamprotula **
+--+
| +-- Unionina
+--+
+-- Oxynaiina
Bolotov et al., 2017 [Publ 3761], fig. S2 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 137 | 9 | 2 | -- |
+----------- Unionina
|
--+ +-------- Scriptolamprotula **
| |
| +-------- Schistodesmus
+--+
+-------- Nodularia
|
| +----- Cuneopsis_szechenyii
+--+
| +-- Tchangsinaia
+--+
+-- Cuneopsis_rufescens
Bolotov et al., 2017 [Publ 3764], fig. S1 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 123 | 9 | 2 | -- |
+-- Unio_crassus
|
+-- Unio_pictorum
|
+-- Unio_tumidus
|
+-- Scriptolamprotula **
|
+-- Cuneopsis_szechenyii
--+
+-- Tchangsinaia
|
+-- Cuneopsis_rufescens
|
+-- Nodularia_douglasiae
|
+-- Nodularia_nuxpersicae
|
+-- Schistodesmus
Sonowal et al., 2021 [Publ 4221], fig. 3b | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 14 | 2 | 1 | -- |
-- Scriptolamprotula_tientsinensis **
Soroka & Burzynski, 2018 [Publ 3762], fig. 3 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 6 | 1 | 1 | -- |
+-- Unionina
--+
+-- Scriptolamprotula **
Wu et al., 2022 [Publ 4238], fig. 2e | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 17 | 12 | 2 | -- |
+----- Scriptolamprotula **
--+
| +-- Unionina
+--+
+-- Oxynaiina
Burzynski & Soroka, 2018 [Publ 3862], fig. 5a | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 19 | 4 | 1 | -- |
+-- Scriptolamprotula **
--+
+-- Unionina
Guerra et al., 2019 [Publ 4168], fig. 2 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 40 | 4 | 2 | -- |
-- Scriptolamprotula_tortuosa **
Wu et al., 2022 [Publ 4238], fig. 2a_S1 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 11 | 11 | 1 | -- |
+-- Scriptolamprotula **
--+
+-- Oxynaiina
Dai et al., 2024 [Publ 4386], fig. 3 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 13 | 10 | 3 | -- |
-- Scriptolamprotula_scripta **
Burzynski et al., 2017 [Publ 3774], fig. 1-top | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 12 | 2 | 1 | -- |
+-- Scriptolamprotula **
--+
+-- Unionina
Guerra et al., 2017 [Publ 3772], fig. 2b | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 24 | 1 | 1 | -- |
-- Scriptolamprotula_tortuosa **
Wang et al., 2018 [Publ 3865], fig. 2 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 30 | 6 | 2 | -- |
+----- Scriptolamprotula **
--+
| +-- Oxynaiina
+--+
+-- Unionina
Burzynski et al., 2017 [Publ 3774], fig. 1-bottom | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 12 | 2 | 1 | -- |
+-- Scriptolamprotula **
--+
+-- Unionina
Araujo et al., 2017 [Publ 3801], fig. 2 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 13 | 2 | 2 | -- |
+-- Oxynaiina
--+
+-- Scriptolamprotula **
Dai et al., 2024 [Publ 4386], fig. 5-ML | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 21 | 11 | 3 | -- |
+-------- Scriptolamprotula **
--+
| +----- Unionina
+--+
| +-- Middendorffinaiina
+--+
+-- Oxynaiina
Kim et al., 2021 [Publ 4197], fig. 1 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 17 | 11 | 2 | -- |
-- Scriptolamprotula_tientsinensis **
Guerra et al., 2019 [Publ 4168], fig. 2-M | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 21 | 4 | 2 | -- |
+----- Scriptolamprotula **
--+
| +-- Oxynaiina
+--+
+-- Unionina
Chen et al., 2023 [Publ 4302], fig. 1 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 40 | 17 | 3 | -- |
+----------------- Scriptolamprotula **
|
| +-- Middendorffinaiina
--+ +-----------+
| | +-- Nodularia
| |
+--+ +----------- Cuneopsis
| |
| | +-- Diaurora
+--+--------+
| +-- Schistodesmus
|
| +-------- Pseudocuneopsis
+--+
| +----- Inversiunio
+--+
| +-- Tchangsinaia
+--+
+-- Unionina
Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 75 | 9 | 2 | -- |
+----- Unionina
--+
| +-- Scriptolamprotula **
+--+
+-- Oxynaiina
Campbell & Lydeard, 2026 [Publ 4810], fig. 4 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 46 | 13 | 2 | -- |
+----- Scriptolamprotula **
--+
| +-- Unionina
+--+
+-- Oxynaiina
Huang et al., 2018 [Publ 3828], fig. 3_5 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 28 | 1 | 1 | -- |
+-- Unionina
--+
+-- Scriptolamprotula **
Fukata & Iigo, 2019 [Publ 4108], fig. 1 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 16 | 12 | 2 | -- |
-- Scriptolamprotula_tientsinensis **
Burzynski & Soroka, 2018 [Publ 3862], fig. 5b | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 9 | 4 | 1 | -- |
+-- Scriptolamprotula **
--+
+-- Unionina
Rosenberg et al., 1997 [Publ 2720], fig. 2B | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 21 | 1 | 1 | -- |
-- Scriptolamprotula_tortuosa **
Dai et al., 2023 [Publ 4285], fig. 5 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 52 | 6 | 2 | -- |
+----- Scriptolamprotula **
--+
| +-- Unionina
+--+
+-- Oxynaiina
Ouyang et al., 2015 [Publ 4493], fig. 1 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 30 | 7 | 1 | -- |
+-- Schistodesmus
+--------+
| +-- Scriptolamprotula **
|
--+ +-- Tchangsinaia
| +-----+
| | +-- Cuneopsis_rufescens
+--+
| +----- Nodularia
+--+
| +-- Cuneopsis_szechenyii
+--+
+-- Cuneopsis_celtiformis
Ouyang et al., 2011 [Publ 3408], fig. 1 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 12 | 2 | 1 | -- |
-- Scriptolamprotula_tortuosa **
Wu et al., 2024 [Publ 4388], fig. 6 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 52 | 8 | 2 | -- |
+----- Scriptolamprotula **
--+
| +-- Unionina
+--+
+-- Oxynaiina
Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 79 | 10 | 2 | -- |
+----- Unionina
--+
| +-- Scriptolamprotula **
+--+
+-- Oxynaiina
Sonowal et al., 2021 [Publ 4221], fig. 4 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 15 | 2 | 1 | -- |
-- Scriptolamprotula_tientsinensis **
Sano et al., 2020 [Publ 4138], fig. 1 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 33 | 6 | 1 | -- |
-- Scriptolamprotula_tortuosa **
Guerra et al., 2019 [Publ 4168], fig. 1 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 40 | 4 | 2 | -- |
+----- Scriptolamprotula **
--+
| +-- Oxynaiina
+--+
+-- Unionina
Bolotov et al., 2017 [Publ 3761], fig. S1 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 137 | 9 | 2 | -- |
+----- Unionina
--+
| +-- Scriptolamprotula **
+--+
+-- Oxynaiina
Soroka & Burzynski, 2018 [Publ 3762], fig. 3-M | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 6 | 1 | 1 | -- |
+-- Unionina
--+
+-- Scriptolamprotula **
Guerra et al., 2017 [Publ 3772], fig. 2a | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 24 | 1 | 1 | -- |
+-- Scriptolamprotula **
--+
+-- Unionina
Ouyang et al., 2015 [Publ 4493], fig. 2 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 30 | 7 | 1 | -- |
-- Scriptolamprotula_tortuosa **
Wang et al., 2018 [Publ 3846], fig. 2 | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 30 | 7 | 2 | -- |
+----- Scriptolamprotula **
--+
| +-- Oxynaiina
+--+
+-- Unionina
Huang et al., 2019 [Publ 3843], fig. 3-M | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 22 | 4 | 2 | -- |
+----- Oxynaiina
--+
| +-- Scriptolamprotula **
+--+
+-- Unionina
Huang et al., 2018 [Publ 3828], fig. 6-M | iT | tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 16 | 3 | 1 | -- |
+-- Scriptolamprotula **
--+
+-- Unionina
|