MUSSELpdb | tribe Acuticostini Starobogatov, 1967 cladomics

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Page last updated
Thu 3 September 2025

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Bivalvia | Unionida | Unionoidea | Unionidae | Unioninae

tribe ACUTICOSTINI Starobogatov, 1967: cladomics
1 genus

Cladomics Summary

number of species = 2 (2 total)
number of subtaxa (rank: genus) = 1
analyzed in 63 cladograms in 38 publications
maximum number of terminals in an analysis = 2 (mean = 1.11, median = 1)
maximum number of subtaxa (genus) = 1 (mean = 1, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 7
maximum number of included terminals = 2, (mean = 2 median = 2)
maximum number of subtaxa (genus) = 1 (mean = 1, median = 1)
proportion of cladograms that support basic monophyly = 1

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

tT	bi	riS	oT	roT	roS	iD
1	2	0	1	630	156	4	--

  +----------------- Lepidodesmini               
  |                                              
  |     +----------- Lanceolariini               
  |  +--+                                        
--+  |  |  +-------- Pseudoacuticosta_retiaria__2
  |  |  +--+                                     
  |  |     |  +----- Acuticosta_chinensis__2 **     
  |  |     +--+                                  
  +--+        |  +-- Acuticosta_chinensis__1 **     
     |        +--+                               
     |           +-- Pseudoacuticosta_retiaria__1
     |                                           
     |  +----------- Globunio                    
     +--+                                        
        |  +-------- Aculamprotulina             
        +--+                                     
           |  +----- Middendorffinaiina          
           +--+                                  
              |  +-- Unionina                    
              +--+                               
                 +-- Oxynaiina

Huang et al., 2019 [Publ 3843], fig. S1

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	45	31	4	--

  +-------------- Anemina      
  |                            
--+  +----------- Sinanodonta  
  |  |                         
  +--+  +-------- Acuticosta **   
     |  |                      
     +--+     +-- Lepidodesmini
        |  +--+                
        |  |  +-- Cristaria    
        +--+                   
           |  +-- Lanceolariini
           +--+                
              +-- Unionini

Zhou et al., 2007 [Publ 2658], fig. 1

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	19	11	3	--

     +----- Aculamprotulina
  +--+                     
  |  |  +-- Tchangsinaia   
  |  +--+                  
--+     +-- Nodularia      
  |                        
  |  +----- Schistodesmus  
  +--+                     
     |  +-- Lanceolariini  
     +--+                  
        +-- Acuticosta **

Wang et al., 2025 [Publ 4488], fig. 3B

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	63	61	3	--

  +-------- Lepidodesmini
--+                      
  |  +----- Unionini     
  +--+                   
     |  +-- Acuticosta **   
     +--+                
        +-- Anodontini

Wu et al., 2026 [Publ 4811], fig. 3

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	15	8	3	--

  +----------- Unionina     
--+                         
  |  +-------- Lanceolariini
  +--+                      
     |  +----- Globunionina 
     +--+                   
        |  +-- Acuticosta **   
        +--+                
           +-- Anodontini

Ouyang et al., 2015 [Publ 4493], fig. 1

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	29	14	3	--

  +-------- Unionini     
--+                      
  |  +----- Lanceolariini
  +--+                   
     |  +-- Acuticosta **   
     +--+                
        +-- Anodontini

Ouyang et al., 2015 [Publ 4493], fig. 2

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	29	14	3	--

  +-------- Lanceolariini  
  |                        
--+     +-- Aculamprotulina
  |  +--+                  
  |  |  +-- Acuticosta **     
  +--+                     
     |  +-- Anodontini     
     +--+                  
        +-- Oxynaiina

Wu et al., 2026 [Publ 4811], fig. 4

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	93	63	4	--

  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticosta **   
        +--+                
           +-- Lanceolariini

Dai et al., 2024 [Publ 4386], fig. 5-ML

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	21	16	4	--

  +----------- Anodontina   
  |                         
--+        +-- Acuticosta **   
  |  +-----+                
  |  |     +-- Cristariina  
  +--+                      
     |  +----- Lepidodesmini
     +--+                   
        |  +-- Lanceolariini
        +--+                
           +-- Unionini

Dai et al., 2024 [Publ 4386], fig. 5-BI

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	21	16	4	--

  +----------- Acuticosta **        
  |                              
  +----------- Cristariina       
  |                              
--+----------- Anodontina        
  |                              
  |  +-------- Lanceolariini     
  |  |                           
  |  |     +-- Aculamprotulina   
  +--+-----+                     
     |     +-- Lepidodesmini     
     |                           
     |  +----- Unionina          
     +--+                        
        |  +-- Middendorffinaiina
        +--+                     
           +-- Oxynaiina

Wu et al., 2020 [Publ 4121], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	48	26	4	--

  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticosta **   
        +--+                
           +-- Lanceolariini

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	38	16	4	--

  +----------- Anodontini     
  |                           
  |        +-- Acuticosta **     
--+  +-----+                  
  |  |     +-- Lanceolariini  
  |  |                        
  +--+     +-- Lepidodesmini  
     |  +--+                  
     |  |  +-- Aculamprotulina
     +--+                     
        |  +-- Unionina       
        +--+                  
           +-- Oxynaiina

Kim et al., 2021 [Publ 4197], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	17	16	3	--

  +-------- Aculamprotulina
  |                        
--+     +-- Acuticosta **     
  |  +--+                  
  |  |  +-- Lanceolariini  
  +--+                     
     |  +-- Unionina       
     +--+                  
        +-- Oxynaiina

Dai et al., 2025 [Publ 4440], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	87	42	4	--

  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticosta **   
        +--+                
           +-- Lanceolariini

Huang et al., 2019 [Publ 3843], fig. S4

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	199	46	4	--

  +-------- Acuticosta **           
  |                              
  +-------- Alasmidonta          
  |                              
  +-------- Anodonta_anatina     
  |                              
  +-------- Anemina              
  |                              
  +-------- Anodonta_cygnea      
  |                              
  +-------- Cristaria            
  |                              
  +-------- Brachyanodon         
  |                              
  +-------- Lasmigona            
  |                              
  +-------- Lepidodesmini        
  |                              
  +-------- Pseudanodonta        
  |                              
  +-------- Pyganodon            
  |                              
--+-------- Simpsonaias          
  |                              
  +-------- Strophitus           
  |                              
  +-------- Utterbackia          
  |                              
  +-------- Lanceolariini        
  |                              
  +-------- Schistodesmus        
  |                              
  +-------- Unionina             
  |                              
  +-------- Aculamprotulina      
  |                              
  |     +-- Pletholophus         
  +-----+                        
  |     +-- Sinanodonta          
  |                              
  |  +----- Cuneopsis_celtiformis
  |  |                           
  |  +----- Tchangsinaia         
  |  |                           
  +--+----- Cuneopsis_rufescens  
     |                           
     +----- Nodularia            
     |                           
     |  +-- Cuneopsis_szechenyii 
     +--+                        
        +-- Cuneopsis_heudei

Wu et al., 2021 [Publ 4366], fig. 3-ML

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	73	33	4	--

  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticosta **   
        +--+                
           +-- Lanceolariini

Wu et al., 2018 [Publ 3836], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	52	32	4	--

  +-------- Lepidodesmini
  |                      
--+-------- Unionini     
  |                      
  |  +----- Anodontini   
  +--+                   
     |  +-- Acuticosta **   
     +--+                
        +-- Lanceolariini

Chen et al., 2023 [Publ 4302], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	40	31	4	--

           +-- Anemina               
  +--------+                         
  |        +-- Amuranodonta          
  |                                  
  |     +----- Pletholophus          
--+  +--+                            
  |  |  |  +-- Sinanodonta           
  |  |  +--+                         
  |  |     +-- Beringiana            
  |  |                               
  +--+  +----- Lanceolaria_gladiola  
     |  |                            
     |  +----- Acuticosta **            
     |  |                            
     |  +----- Lepidodesmini         
     |  |                            
     +--+----- Lanceolaria_oxyrhyncha
        |                            
        +----- Cristaria             
        |                            
        +----- Unionini              
        |                            
        |  +-- Alasmidontina         
        +--+                         
           +-- Anodontina

Chen et al., 2025 [Publ 4487], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	40	31	4	--

  +-------------- Globunionina   
  |                              
  |        +----- Aculamprotulina
--+  +-----+                     
  |  |     |  +-- Unionina       
  |  |     +--+                  
  +--+        +-- Oxynaiina      
     |                           
     |  +-------- Acuticosta **     
     +--+                        
        |  +----- Lepidodesmini  
        +--+                     
           |  +-- Lanceolariini  
           +--+                  
              +-- Anodontini

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	140	68	4	--

  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Lanceolariini
     +--+                   
        |  +-- Acuticosta **   
        +--+                
           +-- Anodontini

Huang et al., 2019 [Publ 3843], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	55	25	4	--

  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Lanceolariini
     +--+                   
        |  +-- Acuticosta **   
        +--+                
           +-- Anodontini

Liu et al., 2017 [Publ 3939], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	14	13	3	--

-- Acuticosta_chinensis **

Chen et al., 2023 [Publ 4302], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	40	31	4	--

  +-------- Acuticosta **   
  |                      
--+  +----- Lepidodesmini
  |  |                   
  +--+----- Unionini     
     |                   
     |  +-- Lanceolariini
     +--+                
        +-- Anodontini

Liu et al., 2023 [Publ 4310], fig. 5

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	23	21	3	--

  +-------- Acuticosta **            
  |                               
--+  +----- Lanceolaria_grayii    
  |  |                            
  |  +----- Lanceolaria_gladiola  
  +--+                            
     +----- Lanceolaria_lanceolata
     |                            
     |  +-- Anodontini            
     +--+                         
        +-- Unionini

Zhou et al., 2007 [Publ 2658], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	17	7	2	--

  +----- Aculamprotulina
--+                     
  |  +-- Acuticosta **     
  +--+                  
     +-- Oxynaiina

Huang et al., 2019 [Publ 3843], fig. S3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	197	45	4	--

  +----------- Acuticosta **   
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Lepidodesmini
     +--+                   
        |  +-- Lanceolariini
        +--+                
           +-- Anodontini

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	95	46	4	--

  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticosta **   
        +--+                
           +-- Lanceolariini

Wu et al., 2021 [Publ 4215], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	23	3	1	--

  +-- Acuticosta **
--+             
  +-- Unionini

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	121	70	4	--

  +----- Acuticosta **   
  |                   
  +----- Lepidodesmini
--+                   
  +----- Unionini     
  |                   
  |  +-- Lanceolariini
  +--+                
     +-- Anodontini

Fukata & Iigo, 2020 [Publ 4106], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	53	27	4	--

  +----------- Lepidodesmini
--+                         
  |  +-------- Lanceolariini
  +--+                      
     |  +----- Unionini     
     +--+                   
        |  +-- Acuticosta **   
        +--+                
           +-- Anodontini

Zhang et al., 2004 [Publ 2735], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	13	11	3	--

  +-------- Anodontini   
--+                      
  |  +----- Unionini     
  +--+                   
     |  +-- Acuticosta **   
     +--+                
        +-- Lanceolariini

Konopleva et al., 2023 [Publ 4269], fig. S1_4

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	55	32	4	--

  +-------- Unionini     
  |                      
--+     +-- Lepidodesmini
  |  +--+                
  |  |  +-- Lanceolariini
  +--+                   
     |  +-- Acuticosta **   
     +--+                
        +-- Anodontini

Fukata & Iigo, 2019 [Publ 4108], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	16	16	2	--

        +-- Unionina       
  +-----+                  
  |     +-- Oxynaiina      
--+                        
  |  +----- Aculamprotulina
  +--+                     
     |  +-- Acuticosta **     
     +--+                  
        +-- Lanceolariini

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	95	46	4	--

  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticosta **   
        +--+                
           +-- Lanceolariini

Liu et al., 2023 [Publ 4310], fig. 4

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	23	21	3	--

  +----------- Acuticosta **            
  |                                  
--+        +-- Lanceolaria_lanceolata
  |  +-----+                         
  |  |     +-- Lanceolaria_grayii    
  +--+                               
     |  +----- Lanceolaria_gladiola  
     +--+                            
        |  +-- Anodontini            
        +--+                         
           +-- Unionini

Wu et al., 2024 [Publ 4350], fig. 8a_S1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	91	46	4	--

  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticosta **   
        +--+                
           +-- Lanceolariini

Chen et al., 2025 [Publ 4448], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	85	43	4	--

  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Lanceolariini
     +--+                   
        |  +-- Acuticosta **   
        +--+                
           +-- Anodontini

Dai et al., 2024 [Publ 4358], fig. 4

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	52	27	4	--

  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Lanceolariini
     +--+                   
        |  +-- Acuticosta **   
        +--+                
           +-- Anodontini

Dai et al., 2023 [Publ 4285], fig. 5

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	52	16	4	--

  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticosta **   
        +--+                
           +-- Lanceolariini

Dai et al., 2024 [Publ 4386], fig. 6

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	41	27	4	--

  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Lanceolariini
     +--+                   
        |  +-- Acuticosta **   
        +--+                
           +-- Anodontini

Wu et al., 2021 [Publ 4366], fig. 3-BI

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	73	33	4	--

  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticosta **   
        +--+                
           +-- Lanceolariini

Dai et al., 2025 [Publ 4440], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	44	35	4	--

           +----- Pletholophus 
  +--------+                   
  |        |  +-- Sinanodonta  
  |        +--+                
  |           +-- Beringiana   
--+                            
  |  +----------- Cristaria    
  |  |                         
  |  |        +-- Alasmidontina
  +--+--------+                
     |        +-- Anodontina   
     |                         
     |  +-------- Lanceolariini
     +--+                      
        |  +----- Lepidodesmini
        +--+                   
           |  +-- Acuticosta **   
           +--+                
              +-- Unionini

Dai et al., 2025 [Publ 4440], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	44	35	4	--

              +-- Anemina               
  +-----------+                         
  |           +-- Amuranodonta          
  |                                     
  |        +----- Pletholophus          
--+  +-----+                            
  |  |     |  +-- Sinanodonta           
  |  |     +--+                         
  |  |        +-- Beringiana            
  +--+                                  
     |        +-- Alasmidontina         
     |  +-----+                         
     |  |     +-- Anodontina            
     |  |                               
     +--+  +----- Lanceolaria_gladiola  
        |  |                            
        |  +----- Lanceolaria_oxyrhyncha
        |  |                            
        +--+----- Cristaria             
           |                            
           |  +-- Acuticosta **            
           |  |                         
           +--+-- Lepidodesmini         
              |                         
              +-- Unionini

Ouyang et al., 2011 [Publ 3408], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	12	8	3	--

  +----------- Anodontini          
  |                                
--+  +-------- Oxynaiina           
  |  |                             
  +--+  +----- Lanceolaria_grayii  
     |  |                          
     +--+----- Acuticosta **          
        |                          
        |  +-- Lanceolaria_gladiola
        +--+                       
           +-- Aculamprotulina

Wu et al., 2021 [Publ 4366], fig. 6

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	90	25	4	--

  +----------- Acuticosta **   
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Lanceolariini
     +--+                   
        |  +-- Lepidodesmini
        +--+                
           +-- Anodontini

Wu et al., 2024 [Publ 4350], fig. 8b_S2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	91	46	4	--

  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticosta **   
        +--+                
           +-- Lanceolariini

Chen et al., 2025 [Publ 4487], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	40	31	4	--

  +-------------------- Unionini     
  |                                  
--+  +----------------- Acuticosta **   
  |  |                               
  |  |              +-- Alasmidontina
  +--+  +-----------+                
     |  |           +-- Anodontina   
     |  |                            
     |  |        +----- Lepidodesmini
     +--+  +-----+                   
        |  |     |  +-- Cristaria    
        |  |     +--+                
        |  |        +-- Lanceolariini
        +--+                         
           |        +-- Anemina      
           |  +-----+                
           |  |     +-- Amuranodonta 
           +--+                      
              |  +----- Pletholophus 
              +--+                   
                 |  +-- Sinanodonta  
                 +--+                
                    +-- Beringiana

Huang et al., 2002 [Publ 2657], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	33	13	4	--

  +----------- Anodontini   
  |                         
  |     +----- Acuticosta **   
--+  +--+                   
  |  |  |  +-- Nodularia    
  |  |  +--+                
  +--+     +-- Tchangsinaia 
     |                      
     |  +----- Lanceolariini
     +--+                   
        |  +-- Schistodesmus
        +--+                
           +-- Lepidodesmini

Wu et al., 2019 [Publ 3864], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	45	25	4	--

  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticosta **   
        +--+                
           +-- Lanceolariini

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	30	10	2	--

  +----- Anodontini
--+                
  |  +-- Acuticosta **
  +--+             
     +-- Unionini

Huang et al., 2019 [Publ 3843], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	44	30	4	--

  +----------------- Anemina        
  |                                 
--+  +-------------- Acuticosta **     
  |  |                              
  +--+  +----------- Sinanodonta    
     |  |                           
     +--+  +-------- Aculamprotulina
        |  |                        
        +--+     +-- Cristaria      
           |  +--+                  
           |  |  +-- Lanceolariini  
           +--+                     
              |  +-- Lepidodesmini  
              +--+                  
                 +-- Oxynaiina

Huang et al., 2019 [Publ 3843], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	199	46	4	--

  +-------- Acuticosta **   
  |                      
--+  +----- Lepidodesmini
  |  |                   
  +--+----- Unionini     
     |                   
     |  +-- Lanceolariini
     +--+                
        +-- Anodontini

Huang et al., 2002 [Publ 2657], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	33	13	4	--

  +-------- Strophitus   
  |                      
  +-------- Schistodesmus
  |                      
  +-------- Lepidodesmini
  |                      
  +-------- Acuticosta **   
  |                      
  +-------- Cristariina  
--+                      
  +-------- Lanceolariini
  |                      
  |     +-- Nodularia    
  +-----+                
  |     +-- Tchangsinaia 
  |                      
  |  +----- Utterbackia  
  |  |                   
  +--+----- Utterbackiana
     |                   
     |  +-- Alasmidonta  
     +--+                
        +-- Lasmigona

Wu et al., 2021 [Publ 4366], fig. 2_I

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	90	25	4	--

  +-------- Acuticosta **   
  |                      
--+     +-- Lepidodesmini
  |  +--+                
  |  |  +-- Unionini     
  +--+                   
     |  +-- Lanceolariini
     +--+                
        +-- Anodontini

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	36	15	4	--

  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Lanceolariini
     +--+                   
        |  +-- Acuticosta **   
        +--+                
           +-- Anodontini

Wu et al., 2023 [Publ 4296], fig. 4

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	22	20	3	--

  +-------- Acuticosta **   
--+                      
  |  +----- Lanceolariini
  +--+                   
     |  +-- Anodontini   
     +--+                
        +-- Unionini

Aunins et al., 2018 [Publ 3863], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	39	25	4	--

  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticosta **   
        +--+                
           +-- Lanceolariini

Wu et al., 2021 [Publ 4366], fig. 4

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	68	32	4	--

  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticosta **   
        +--+                
           +-- Lanceolariini

Sano et al., 2020 [Publ 4138], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	33	21	4	--

  +----- Lanceolaria_grayii__1  
  |                             
  +----- Lanceolaria_grayii__2  
  |                             
  +----- Cristaria_plicata__1   
  |                             
  +----- Cristaria_plicata__2   
  |                             
  +----- Sinanodonta_lauta      
  |                             
  +----- Beringiana_japonica    
  |                             
  +----- Beringiana_fukuharai__1
  |                             
  +----- Beringiana_beringiana  
  |                             
  +----- Beringiana_fukuharai__2
  |                             
  +----- Beringiana_fukuharai__3
  |                             
  +----- Buldowskia_iwakawai__1 
  |                             
  +----- Buldowskia_iwakawai__2 
  |                             
  +----- Sinanodonta_tumens__1  
--+                             
  +----- Sinanodonta_tumens__2  
  |                             
  +----- Pletholophus           
  |                             
  +----- Anemina                
  |                             
  +----- Lanceolaria_lanceolata 
  |                             
  +----- Cristaria_plicata__3   
  |                             
  +----- Lanceolaria_gladiola   
  |                             
  +----- Sinanodonta_angula     
  |                             
  +----- Lepidodesmini          
  |                             
  |  +-- Aculamprotulina        
  +--+                          
  |  +-- Acuticosta **             
  |                             
  |  +-- Cuneopsis              
  +--+                          
  |  +-- Nodularia              
  |                             
  |  +-- Schistodesmus          
  +--+                          
     +-- Inversiunio

Wu et al., 2022 [Publ 4238], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	73	35	4	--

  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticosta **   
        +--+                
           +-- Lanceolariini

Wu et al., 2022 [Publ 4238], fig. 2d

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	38	38	4	--

  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Lanceolariini
     +--+                   
        |  +-- Acuticosta **   
        +--+                
           +-- Anodontini

Wang et al., 2018 [Publ 3865], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	30	16	3	--

  +-------- Unionini     
--+                      
  |  +----- Anodontini   
  +--+                   
     |  +-- Acuticosta **   
     +--+                
        +-- Lanceolariini

Sano et al., 2017 [Publ 3799], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	32	20	4	--

  +-------------- Anodontini   
  |                            
  |           +-- Tchangsinaia 
--+  +--------+                
  |  |        +-- Nodularia    
  |  |                         
  +--+        +-- Schistodesmus
     |  +-----+                
     |  |     +-- Acuticosta **   
     +--+                      
        |  +----- Inversiunio  
        +--+                   
           |  +-- Lepidodesmini
           +--+                
              +-- Lanceolariini

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