The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)
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Bivalvia | Unionoida | Unionoidea | Unionidae | Unioninae tribe ACUTICOSTINI Starobogatov, 1967: cladomics
1 genus Cladomics Summarynumber of species = 5 (5 total)
number of subtaxa (rank: genus) = 1
analyzed in 53 cladograms in 32 publications
maximum number of terminals in an analysis =
2 (mean = 1.04, median = 1)
maximum number of subtaxa (genus) =
1 (mean = 1, median = 1) Basic Monophylynumber of cladograms that test basic monophyly
(i.e., more than 1 included species) = 2
maximum number of included terminals = 2, (mean =
2 median = 2)
maximum number of subtaxa (genus) =
1 (mean = 1, median = 1)
proportion of cladograms that support basic monophyly
= 1
Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4 ** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram ** iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 120 | 69 | 4 | -- | +----- Acuticosta **
|
+----- Lepidodesmini
--+
+----- Unionini
|
| +-- Lanceolariini
+--+
+-- Anodontini
Huang et al., 2019 [Publ 3843], fig. S1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 43 | 29 | 4 | -- | +-------------- Anemina
|
--+ +----------- Sinanodonta
| |
+--+ +-------- Acuticosta **
| |
+--+ +-- Lepidodesmini
| +--+
| | +-- Cristaria
+--+
| +-- Lanceolariini
+--+
+-- Unionini
Zhou et al., 2007 [Publ 2658], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 19 | 11 | 3 | -- | +----- Aculamprotula
+--+
| | +-- Tchangsinaia
| +--+
--+ +-- Nodularia
|
| +----- Schistodesmus
+--+
| +-- Lanceolariini
+--+
+-- Acuticosta **
Wu et al., 2021 [Publ 4366], fig. 2_I iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 90 | 25 | 4 | -- | +-------- Acuticosta **
|
--+ +-- Lepidodesmini
| +--+
| | +-- Unionini
+--+
| +-- Lanceolariini
+--+
+-- Anodontini
Wu et al., 2021 [Publ 4366], fig. 3-BI iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 72 | 32 | 4 | -- | +----------- Lepidodesmini
--+
| +-------- Unionini
+--+
| +----- Anodontini
+--+
| +-- Acuticosta **
+--+
+-- Lanceolariini
Liu et al., 2023 [Publ 4310], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 23 | 21 | 3 | -- | +----------- Acuticosta **
|
--+ +-- Lanceolaria_lanceolata
| +-----+
| | +-- Lanceolaria_grayii
+--+
| +----- Lanceolaria_gladiola
+--+
| +-- Anodontini
+--+
+-- Unionini
Huang et al., 2019 [Publ 3843], fig. S4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 198 | 44 | 4 | -- | +-------- Acuticosta **
|
+-------- Alasmidonta
|
+-------- Anodonta_anatina
|
+-------- Anemina
|
+-------- Anodonta_cygnea
|
+-------- Cristaria
|
+-------- Anodonta_nuttalliana
|
+-------- Lasmigona
|
+-------- Lepidodesmini
|
+-------- Pseudanodonta
|
+-------- Pyganodon
|
--+-------- Simpsonaias
|
+-------- Strophitus
|
+-------- Schistodesmus
|
+-------- Utterbackia
|
+-------- Lanceolariini
|
+-------- Unio
|
+-------- Aculamprotula
|
| +-- Pletholophus
+-----+
| +-- Sinanodonta
|
| +----- Cuneopsis_celtiformis
| |
| +----- Tchangsinaia
| |
+--+----- Cuneopsis_rufescens
|
+----- Nodularia
|
| +-- Cuneopsis_szechenyii
+--+
+-- Cuneopsis_heudei
Wu et al., 2020 [Publ 4121], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 48 | 26 | 4 | -- | +----------- Lepidodesmini
--+
| +-------- Unionini
+--+
| +----- Anodontini
+--+
| +-- Acuticosta **
+--+
+-- Lanceolariini
Dai et al., 2025 [Publ 4440], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 44 | 35 | 4 | -- | +-- Anemina
+-----------+
| +-- Amuranodonta
|
| +----- Pletholophus
--+ +-----+
| | | +-- Sinanodonta
| | +--+
| | +-- Beringiana
+--+
| +-- Alasmidontina
| +-----+
| | +-- Anodontina
| |
+--+ +----- Lanceolaria_gladiola
| |
| +----- Lanceolaria_oxyrhyncha
| |
+--+----- Cristaria
|
| +-- Acuticosta **
| |
+--+-- Lepidodesmini
|
+-- Unionini
Pfeiffer & Graf, 2013 [Publ 3531], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 30 | 10 | 2 | -- | +----- Anodontini
--+
| +-- Acuticosta **
+--+
+-- Unionini
Dai et al., 2025 [Publ 4440], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 87 | 43 | 4 | -- | +----------- Lepidodesmini
--+
| +-------- Unionini
+--+
| +----- Anodontini
+--+
| +-- Acuticosta **
+--+
+-- Lanceolariini
Dai et al., 2024 [Publ 4386], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 39 | 25 | 4 | -- | +----------- Lepidodesmini
--+
| +-------- Unionini
+--+
| +----- Lanceolariini
+--+
| +-- Acuticosta **
+--+
+-- Anodontini
Dai et al., 2024 [Publ 4358], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 51 | 26 | 4 | -- | +----------- Lepidodesmini
--+
| +-------- Unionini
+--+
| +----- Lanceolariini
+--+
| +-- Acuticosta **
+--+
+-- Anodontini
Wu et al., 2022 [Publ 4238], fig. 2d iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 37 | 37 | 4 | -- | +----------- Lepidodesmini
--+
| +-------- Unionini
+--+
| +----- Lanceolariini
+--+
| +-- Acuticosta **
+--+
+-- Anodontini
Huang et al., 2019 [Publ 3843], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 198 | 44 | 4 | -- | +-------- Acuticosta **
|
--+ +----- Lepidodesmini
| |
+--+----- Unionini
|
| +-- Lanceolariini
+--+
+-- Anodontini
Wu et al., 2021 [Publ 4366], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 90 | 25 | 4 | -- | +----------- Acuticosta **
--+
| +-------- Unionini
+--+
| +----- Lanceolariini
+--+
| +-- Lepidodesmini
+--+
+-- Anodontini
Ouyang et al., 2011 [Publ 3408], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 12 | 8 | 3 | -- | +----------- Anodontini
|
--+ +-- Nodularia
| +-----+
| | +-- Cuneopsis
+--+
| +----- Lanceolaria_grayii
| |
+--+----- Acuticosta **
|
| +-- Lanceolaria_gladiola
+--+
+-- Aculamprotula
Dai et al., 2025 [Publ 4440], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 44 | 35 | 4 | -- | +----- Pletholophus
+--------+
| | +-- Sinanodonta
| +--+
| +-- Beringiana
--+
| +----------- Cristaria
| |
| | +-- Alasmidontina
+--+--------+
| +-- Anodontina
|
| +-------- Lanceolariini
+--+
| +----- Lepidodesmini
+--+
| +-- Acuticosta **
+--+
+-- Unionini
Huang et al., 2002 [Publ 2657], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 33 | 13 | 4 | -- | +-------- Strophitus
|
+-------- Schistodesmus
|
+-------- Lepidodesmini
|
+-------- Acuticosta **
|
+-------- Cristariina
--+
+-------- Lanceolariini
|
| +-- Nodularia
+-----+
| +-- Tchangsinaia
|
| +----- Utterbackia
| |
+--+----- Utterbackiana
|
| +-- Alasmidonta
+--+
+-- Lasmigona
Konopleva et al., 2023 [Publ 4269], fig. S1_4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 55 | 32 | 4 | -- | +-------- Unionini
|
--+ +-- Lepidodesmini
| +--+
| | +-- Lanceolariini
+--+
| +-- Acuticosta **
+--+
+-- Anodontini
Liu et al., 2023 [Publ 4310], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 23 | 21 | 3 | -- | +-------- Acuticosta **
|
--+ +----- Lanceolaria_grayii
| |
| +----- Lanceolaria_gladiola
+--+
+----- Lanceolaria_lanceolata
|
| +-- Anodontini
+--+
+-- Unionini
Huang et al., 2019 [Publ 3843], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 55 | 25 | 4 | -- | +----------- Lepidodesmini
--+
| +-------- Unionini
+--+
| +----- Lanceolariini
+--+
| +-- Acuticosta **
+--+
+-- Anodontini
Huang et al., 2002 [Publ 2657], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 33 | 13 | 4 | -- | +----------- Anodontini
|
| +----- Acuticosta **
--+ +--+
| | | +-- Nodularia
| | +--+
+--+ +-- Tchangsinaia
|
| +----- Lanceolariini
+--+
| +-- Schistodesmus
+--+
+-- Lepidodesmini
Huang et al., 2019 [Publ 3843], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 42 | 28 | 4 | -- | +----------------------------------- Anemina
|
--+ +-------------------------------- Acuticosta **
| |
+--+ +----------------------------- Sinanodonta
| |
+--+ +-------------------------- Aculamprotula
| |
+--+ +-- Cristaria
| +--------------------+
| | +-- Lanceolariini
+--+
| +-------------------- Lepidodesmini
+--+
| +----------------- Schistodesmus
+--+
| +-------------- Nodularia
+--+
| +----------- Cuneopsis_rufescens
+--+
| +-------- Tchangsinaia
+--+
| +----- Cuneopsis_celtiformis
+--+
| +-- Cuneopsis_szechenyii
+--+
+-- Cuneopsis_heudei
Pavan-Kumar et al., 2022 [Publ 4263], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 35 | 14 | 4 | -- | +----------- Lepidodesmini
--+
| +-------- Unionini
+--+
| +----- Lanceolariini
+--+
| +-- Acuticosta **
+--+
+-- Anodontini
Sano et al., 2017 [Publ 3799], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 32 | 20 | 4 | -- | +-------------- Anodontini
|
| +-- Tchangsinaia
--+ +--------+
| | +-- Nodularia
| |
+--+ +-- Schistodesmus
| +-----+
| | +-- Acuticosta **
+--+
| +----- Inversiunio
+--+
| +-- Lepidodesmini
+--+
+-- Lanceolariini
Wu et al., 2023 [Publ 4339], fig. 4_S5-aa iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 94 | 45 | 4 | -- | +----------- Lepidodesmini
--+
| +-------- Unionini
+--+
| +----- Anodontini
+--+
| +-- Acuticosta **
+--+
+-- Lanceolariini
Wang et al., 2018 [Publ 3865], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 30 | 16 | 3 | -- | +-------- Unionini
--+
| +----- Anodontini
+--+
| +-- Acuticosta **
+--+
+-- Lanceolariini
Wu et al., 2021 [Publ 4215], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 23 | 3 | 1 | -- | +-- Acuticosta **
--+
+-- Unionini
Wu et al., 2023 [Publ 4296], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 22 | 20 | 3 | -- | +-------- Acuticosta **
--+
| +----- Lanceolariini
+--+
| +-- Anodontini
+--+
+-- Unionini
Wu et al., 2021 [Publ 4366], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 67 | 31 | 4 | -- | +----------- Lepidodesmini
--+
| +-------- Unionini
+--+
| +----- Anodontini
+--+
| +-- Acuticosta **
+--+
+-- Lanceolariini
Aunins et al., 2018 [Publ 3863], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 39 | 25 | 4 | -- | +----------- Lepidodesmini
--+
| +-------- Unionini
+--+
| +----- Anodontini
+--+
| +-- Acuticosta **
+--+
+-- Lanceolariini
Sano et al., 2020 [Publ 4138], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 33 | 21 | 4 | -- | +----- Lanceolaria_grayii__1
|
+----- Lanceolaria_grayii__2
|
+----- Cristaria_plicata__1
|
+----- Cristaria_plicata__2
|
+----- Sinanodonta_lauta
|
+----- Beringiana_japonica
|
+----- Beringiana_fukuharai__1
|
+----- Beringiana_beringiana
|
+----- Beringiana_fukuharai__2
|
+----- Beringiana_fukuharai__3
|
+----- Buldowskia_iwakawai__1
|
+----- Buldowskia_iwakawai__2
|
+----- Sinanodonta_tumens__1
--+
+----- Sinanodonta_tumens__2
|
+----- Pletholophus
|
+----- Anemina
|
+----- Lanceolaria_lanceolata
|
+----- Cristaria_plicata__3
|
+----- Lanceolaria_gladiola
|
+----- Sinanodonta_angula
|
+----- Lepidodesmini
|
| +-- Aculamprotula
+--+
| +-- Acuticosta **
|
| +-- Cuneopsis
+--+
| +-- Nodularia
|
| +-- Schistodesmus
+--+
+-- Inversiunio
Dai et al., 2024 [Publ 4386], fig. 5-BI iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 20 | 15 | 4 | -- | +----------------- Acuticosta **
|
+----------------- Cristariina
|
+----------------- Anodontina
--+
| +-------------- Lanceolariini
| |
| | +-- Aculamprotula
| +-----------+
+--+ +-- Lepidodesmini
|
| +----------- Unio
| |
+--+ +-------- Middendorffinaia
| |
+--+ +----- Cuneopsis
| |
+--+----- Nodularia
|
| +-- Inversiunio
+--+
+-- Schistodesmus
Wu et al., 2024 [Publ 4350], fig. 8b_S2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 90 | 45 | 4 | -- | +----------- Lepidodesmini
--+
| +-------- Unionini
+--+
| +----- Anodontini
+--+
| +-- Acuticosta **
+--+
+-- Lanceolariini
Dai et al., 2023 [Publ 4285], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 52 | 16 | 4 | -- | +----------- Lepidodesmini
--+
| +-------- Unionini
+--+
| +----- Anodontini
+--+
| +-- Acuticosta **
+--+
+-- Lanceolariini
Wu et al., 2021 [Publ 4366], fig. 3-ML iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 72 | 32 | 4 | -- | +----------- Lepidodesmini
--+
| +-------- Unionini
+--+
| +----- Anodontini
+--+
| +-- Acuticosta **
+--+
+-- Lanceolariini
Wu et al., 2024 [Publ 4350], fig. 8a_S1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 90 | 45 | 4 | -- | +----------- Lepidodesmini
--+
| +-------- Unionini
+--+
| +----- Anodontini
+--+
| +-- Acuticosta **
+--+
+-- Lanceolariini
Wu et al., 2018 [Publ 3836], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 52 | 32 | 4 | -- | +-------- Lepidodesmini
|
--+-------- Unionini
|
| +----- Anodontini
+--+
| +-- Acuticosta **
+--+
+-- Lanceolariini
Liu et al., 2017 [Publ 3939], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 14 | 13 | 3 | -- | -- Acuticosta_chinensis **
Kim et al., 2021 [Publ 4197], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 16 | 15 | 3 | -- | +-------------- Aculamprotula
|
--+ +-- Acuticosta **
| +--------+
| | +-- Lanceolariini
+--+
| +-------- Unio
+--+
| +----- Nodularia
+--+
| +-- Schistodesmus
+--+
+-- Cuneopsis
Chen et al., 2023 [Publ 4302], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 39 | 30 | 4 | -- | +-------- Acuticosta **
|
--+ +----- Lepidodesmini
| |
+--+----- Unionini
|
| +-- Lanceolariini
+--+
+-- Anodontini
Dai et al., 2024 [Publ 4386], fig. 5-ML iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 20 | 15 | 4 | -- | +----------- Anodontina
|
--+ +-- Acuticosta **
| +-----+
| | +-- Cristariina
+--+
| +----- Lepidodesmini
+--+
| +-- Lanceolariini
+--+
+-- Unionini
Zhou et al., 2007 [Publ 2658], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 17 | 7 | 2 | -- | +-------- Aculamprotula
--+
| +----- Acuticosta **
+--+
| +-- Nodularia
+--+
+-- Cuneopsis
Wu et al., 2023 [Publ 4339], fig. 4_S5-nt iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 94 | 45 | 4 | -- | +----------- Lepidodesmini
--+
| +-------- Unionini
+--+
| +----- Anodontini
+--+
| +-- Acuticosta **
+--+
+-- Lanceolariini
Wu et al., 2019 [Publ 3864], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 45 | 25 | 4 | -- | +----------- Lepidodesmini
--+
| +-------- Unionini
+--+
| +----- Anodontini
+--+
| +-- Acuticosta **
+--+
+-- Lanceolariini
Chen et al., 2023 [Publ 4302], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 39 | 30 | 4 | -- | +-- Anemina
+--------+
| +-- Amuranodonta
|
| +----- Pletholophus
--+ +--+
| | | +-- Sinanodonta
| | +--+
| | +-- Beringiana
| |
+--+ +----- Lanceolaria_gladiola
| |
| +----- Acuticosta **
| |
| +----- Lepidodesmini
| |
+--+----- Lanceolaria_oxyrhyncha
|
+----- Cristaria
|
+----- Unionini
|
| +-- Alasmidontina
+--+
+-- Anodontina
Huang et al., 2019 [Publ 3843], fig. S3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 196 | 43 | 4 | -- | +----------- Acuticosta **
--+
| +-------- Unionini
+--+
| +----- Lepidodesmini
+--+
| +-- Lanceolariini
+--+
+-- Anodontini
Pavan-Kumar et al., 2022 [Publ 4263], fig. S5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 37 | 15 | 4 | -- | +----------------- Anodontini
|
| +-- Acuticosta **
--+ +-----------+
| | +-- Lanceolariini
| |
+--+ +-- Lepidodesmini
| +--------+
| | +-- Aculamprotula
+--+
| +-------- Unio
+--+
| +----- Schistodesmus
+--+
| +-- Nodularia
+--+
+-- Tchangsinaia
Wu et al., 2022 [Publ 4238], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 72 | 34 | 4 | -- | +----------- Lepidodesmini
--+
| +-------- Unionini
+--+
| +----- Anodontini
+--+
| +-- Acuticosta **
+--+
+-- Lanceolariini
Fukata & Iigo, 2020 [Publ 4106], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 52 | 26 | 4 | -- | +----------- Lepidodesmini
--+
| +-------- Lanceolariini
+--+
| +----- Unionini
+--+
| +-- Acuticosta **
+--+
+-- Anodontini
Zhang et al., 2004 [Publ 2735], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 13 | 11 | 3 | -- | +-------- Anodontini
--+
| +----- Unionini
+--+
| +-- Acuticosta **
+--+
+-- Lanceolariini
Fukata & Iigo, 2019 [Publ 4108], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 15 | 15 | 2 | -- | +----- Aculamprotula
+--------+
| | +-- Acuticosta **
| +--+
--+ +-- Lanceolariini
|
| +----------- Unio
| |
+--+ +-------- Pseudocuneopsis
| |
+--+ +-- Schistodesmus
| +--+
| | +-- Cuneopsis
+--+
| +-- Tchangsinaia
+--+
+-- Nodularia
|
