The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)
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Bivalvia | Unionoida | Unionoidea | Unionidae | Unioninae | Anodontini subtribe ALASMIDONTINA Rafinesque, 1820: cladomics
17 genera Cladomics Summarynumber of species = 49 (53 total)
number of subtaxa (rank: genus) = 17
analyzed in 209 cladograms in 126 publications
maximum number of terminals in an analysis =
33 (mean = 4.29, median = 3)
maximum number of subtaxa (genus) =
15 (mean = 3.15, median = 2) Basic Monophylynumber of cladograms that test basic monophyly
(i.e., more than 1 included species) = 174
maximum number of included terminals = 33, (mean =
4.95 median = 4)
maximum number of subtaxa (genus) =
15 (mean = 3.58, median = 3)
proportion of cladograms that support basic monophyly
= 0.84 Rigorous Monophylynumber of cladograms that test rigorous monophyly = 127
(Rigorous monophyly refers to cladograms with terminal species representing
more than one subtaxon (genus) taxon and at least one rigorous outgroup (family).
maximum number of included terminals = 33 (mean = 5.32, median = 4)
maximum number of subtaxa (genus) =
15 (mean = 3.9, median = 3)
proportion of cladograms that support rigorous monophyly = 0.8
Pairwise Cladogram ClusteringThe graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.
Perkins et al., 2025 [Publ 4447], fig. 2_mtDNA-ITS ** Top Monophyly Cladogram ** iT| tT | bi | riS | oT | roT | roS | iD | | 33 | 33 | 34 | 15 | 3 | 3 | 1 | 0 | +----------------------- Anodonta
|
| +-------------------- Pseudanodonta
| |
--+ | +-------------- Utterbackiana **
| | +--+
| | | | +----------- Alasminota **
| | | +--+
+--+ | | +-------- Pseudodontoideus **
| | +--+
| | | +----- Lasmigona **
| | +--+
| | | +-- Pterosyna **
| | +--+
+--+ +-- Arcidens **
|
| +-- Prolasmidonta **
| +--+
| | +-- Platynaias **
| +-----+
| | | +-- Ligodonta **
| | +--+
| | +-- Alasmidonta **
+-----+
| +-- Anodontoides **
| +-----+
| | +-- Strophitus **
+--+
| +----- Sulcularia **
+--+
| +-- Pyganodon **
+--+
+-- Utterbackia **
Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4 ** Top Sister-Group Cladogram ** iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 1 | 5 | 117 | 28 | 2 | 36 | +----------- Cristariina
|
--+ +-------- Pseudanodonta
| |
| +-------- Anodonta
+--+
| +----- Pyganodon **
| |
| +----- Simpsonaias **
+--+
+----- Strophitus **
|
| +-- Alasmidonta **
+--+
+-- Lasmigona **
Whelan et al., 2023 [Publ 4287], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 33 | 33 | 34 | 14 | 8 | 8 | 2 | 27 | +-------------------------- Cristariina
|
--+ +----------------------- Anodonta_cygnea
| |
| | +-- Anodonta_nuttalliana
+--+ +-----------------+
| | +-- Pseudanodonta
| |
+--+ +----------------- Anodonta_anatina
| |
| | +-- Utterbackia **
| | +-----------+
+--+ | +-- Pyganodon **
| |
| | +-------- Utterbackiana **
| | |
+--+ +--+ +-- Lasmigona **
| | | +--+
| | | | +-- Pseudodontoideus **
| | +--+
| | | +-- Pterosyna **
| | +--+
+--+ +-- Arcidens **
|
| +----- Sulcularia **
| +--+
| | | +-- Anodontoides **
| | +--+
| | +-- Strophitus **
+--+
| +-- Prolasmidonta **
| +--+
| | +-- Platynaias **
+--+
| +-- Alasminota **
+--+
+-- Alasmidonta **
Whelan et al., 2023 [Publ 4287], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 27 | 27 | 26 | 12 | 1 | 1 | 1 | 23 | +----------------------- Anodontina
|
--+ +-------------------- Utterbackiana **
| |
| | +-- Utterbackia **
+--+ +--------------+
| | +-- Pyganodon **
| |
+--+ +----- Alasmidonta **
| +--------+
| | | +-- Arcidens **
| | +--+
+--+ +-- Pseudodontoideus **
|
| +----------- Alasminota **
| |
+--+ +-- Anodontoides **
| +-----+
| | +-- Strophitus **
+--+
| +----- Sulcularia **
+--+
| +-- Platynaias **
+--+
+-- Prolasmidonta **
Whelan et al., 2023 [Publ 4287], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 25 | 25 | 23 | 11 | 2 | 2 | 1 | 32 | +----------------------- Anodontina
|
--+ +-------------------- Sulcularia **
| |
| | +-------- Utterbackiana **
+--+ +--------+
| | | +----- Alasminota **
| | +--+
| | | +-- Arcidens **
+--+ +--+
| +-- Pseudodontoideus **
|
| +-------------- Alasmidonta **
+--+
| +----------- Platynaias **
+--+
| +-------- Strophitus **
+--+
| +----- Pyganodon **
+--+
| +-- Utterbackia **
+--+
+-- Anodontoides **
Smith et al., 2018 [Publ 3790], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 17 | 17 | 16 | 8 | 1 | 1 | 1 | 28 | +-------------- Cristariina
|
| +-- Utterbackia **
--+ +--------+
| | +-- Pyganodon **
| |
+--+ +-- Anodontoides **
| +-----+
| | +-- Strophitus **
| |
+--+ +-- Alasmidonta **
| +--+
| | +-- Alasminota **
+--+
| +-- Pseudodontoideus **
+--+
+-- Utterbackiana **
Inoue et al., 2018 [Publ 3858], fig. 1_3a-c iT| tT | bi | riS | oT | roT | roS | iD | | 16 | 20 | 0 | 10 | 88 | 4 | 2 | 42 | +-- Alasmidonta_marginata **
|
+-- Prolasmidonta **
|
+-- Alasmidonta_undulata **
|
+-- Cristariina
|
+-- Anodonta_kennerlyi
|
+-- Anodonta_nuttalliana
|
+-- Anodonta_oregonensis
|
+-- Arcidens_confragosus **
|
+-- Arcidens_wheeleri **
|
+-- Pterosyna **
|
--+-- Platynaias_compressa **
|
+-- Lasmigona **
|
+-- Platynaias_decorata **
|
+-- Platynaias_subviridis **
|
+-- Pyganodon_grandis **
|
+-- Pyganodon_lacustris **
|
+-- Pseudodontoideus **
|
+-- Strophitus **
|
+-- Utterbackia_imbecillis **
|
+-- Utterbackia_peninsularis__1 **
|
+-- Utterbackia_peninsularis__2 **
Zanatta et al., 2007 [Publ 2698], fig. 1A iT| tT | bi | riS | oT | roT | roS | iD | | 14 | 16 | 10 | 6 | 2 | 2 | 1 | 42 | +-- Platynaias **
|
+--------+-- Lasmigona **
| |
| +-- Pterosyna **
--+
| +-------- Anodonta_cygnea
| |
+--+ +-- Anodonta_kennerlyi
| +--+
| | +-- Utterbackiana **
+--+
| +-- Utterbackia **
+--+
+-- Pyganodon **
Zanatta et al., 2007 [Publ 2698], fig. 1B iT| tT | bi | riS | oT | roT | roS | iD | | 14 | 16 | 13 | 6 | 2 | 2 | 1 | 44 | +-- Lasmigona **
|
+--------+-- Platynaias **
| |
| +-- Pterosyna **
--+
| +-- Utterbackia **
| +-----+
| | +-- Pyganodon **
+--+
| +----- Anodonta_cygnea
+--+
| +-- Anodonta_kennerlyi
+--+
+-- Utterbackiana **
Inoue et al., 2014 [Publ 3635], fig. 3A iT| tT | bi | riS | oT | roT | roS | iD | | 14 | 14 | 13 | 9 | 6 | 0 | 0 | 29 | +-------- Pseudodontoideus **
+--+
| | +----- Lasmigona **
| +--+
| | +-- Pterosyna **
--+ +--+
| +-- Arcidens **
|
| +-------- Strophitus **
| |
+--+ +-- Pyganodon **
| +--+
| | +-- Utterbackia **
+--+
| +-- Platynaias **
+--+
+-- Alasmidonta **
Hoeh, 1990 [Publ 2717], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 13 | 15 | 13 | 6 | 2 | 2 | 1 | 44 | +-- Pterosyna **
|
+--------+-- Platynaias **
| |
| +-- Lasmigona **
--+
| +-- Utterbackia **
| +-----+
| | +-- Pyganodon **
+--+
| +----- Anodonta_cygnea
+--+
| +-- Anodonta_kennerlyi
+--+
+-- Utterbackiana **
Hoeh, 1990 [Publ 2717], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 13 | 15 | 13 | 6 | 2 | 2 | 1 | 44 | +-- Pterosyna **
|
+--------+-- Platynaias **
| |
| +-- Lasmigona **
--+
| +-- Utterbackia **
| +-----+
| | +-- Pyganodon **
+--+
| +----- Anodonta_cygnea
+--+
| +-- Anodonta_kennerlyi
+--+
+-- Utterbackiana **
Hoeh, 1990 [Publ 2717], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 13 | 15 | 2 | 6 | 2 | 2 | 1 | 45 | +----- Pyganodon_fragilis **
|
+----- Utterbackia_imbecillis **
|
+----- Utterbackia_peggyae **
|
+----- Anodonta_cygnea
|
+----- Anodonta_kennerlyi
|
+----- Utterbackiana_couperiana__1 **
|
--+----- Utterbackiana_couperiana__2 **
|
+----- Utterbackiana_suborbiculata **
|
| +-- Lasmigona **
+--+
| +-- Utterbackiana_implicata **
|
| +-- Pterosyna **
| |
| +-- Platynaias **
| |
| +-- Pyganodon_cataracta **
+--+
+-- Pyganodon_gibbosa **
|
+-- Pyganodon_grandis **
|
+-- Pyganodon_lacustris **
Breton et al., 2011 [Publ 3376], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 11 | 11 | 10 | 7 | 26 | 1 | 1 | 34 | +-------------- Cristariina
|
--+ +-- Pterosyna **
| +--------+
| | +-- Lasmigona **
+--+
| +-------- Strophitus **
| |
+--+ +-- Platynaias **
| +--+
| | +-- Alasmidonta **
+--+
| +-- Pyganodon **
+--+
+-- Utterbackia **
Chase et al., 2018 [Publ 3847], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 11 | 11 | 10 | 7 | 27 | 3 | 2 | 38 | +----------------- Cristariina
|
--+ +-------------- Anodontina
| |
| | +-- Pyganodon **
+--+ +--------+
| | +-- Utterbackia **
| |
+--+ +-- Lasmigona **
| +-----+
| | +-- Pterosyna **
+--+
| +----- Alasmidonta **
+--+
| +-- Strophitus **
+--+
+-- Platynaias **
Stewart et al., 2021 [Publ 4451], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 10 | 10 | 9 | 6 | 6 | 0 | 0 | 39 | +-- Alasmidonta **
+-----+
| +-- Pterosyna **
|
--+ +-- Strophitus **
| +--+
| | +-- Platynaias **
+--+
| +-- Pyganodon **
+--+
+-- Utterbackia **
Boyer et al., 2011 [Publ 2718], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 9 | 9 | 6 | 9 | 29 | 0 | 0 | 39 | +-------------- Strophitus **
|
| +----- Pterosyna **
--+ +-----+
| | | +-- Lasmigona **
| | +--+
+--+ +-- Arcidens **
|
| +-------- Utterbackia **
| |
+--+-------- Alasmidonta **
|
| +----- Pyganodon **
+--+
| +-- Anodontoides **
+--+
+-- Platynaias **
Krebs et al., 2003 [Publ 2691], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 9 | 10 | 5 | 8 | 24 | 1 | 1 | 32 | +----------- Utterbackiana **
|
+----------- Platynaias **
|
+----------- Alasmidonta **
--+
| +-- Pterosyna **
+--------+
| +-- Lasmigona **
|
| +-------- Strophitus **
+--+
| +----- Utterbackia **
+--+
| +-- Pyganodon **
+--+
+-- Anodontina
King et al., 1999 [Publ 2699], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 9 | 9 | 7 | 6 | 0 | 0 | 0 | 35 | +----------- Pyganodon **
|
--+ +-- Lasmigona **
| +-----+
| | +-- Pterosyna **
+--+
| +----- Strophitus **
+--+
| +-- Prolasmidonta **
+--+
+-- Platynaias **
Chong et al., 2008 [Publ 2668], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 8 | 18 | 10 | 8 | 10 | 9 | 2 | 46 | +----------------- Anodonta_cygnea
|
+----------------- Pyganodon **
|
+----------------- Alasminota **
|
+----------------- Anodonta_anatina
|
+----------------- Unio
|
+----------------- Cristariina
|
| +-- Anodonta_nuttalliana
--+--------------+
| +-- Anodonta_californiensis
|
| +-- Pseudodontoideus **
| |
+--------------+-- Alasmidonta **
| |
| +-- Pterosyna **
|
| +-------------- Pseudanodonta
| |
| | +----- Strophitus **
| +--------+
+--+ | +-- Prolasmidonta **
| +--+
| +-- Platynaias **
|
| +----------- Anodonta_oregonensis__4
+--+
| +-------- Anodonta_oregonensis__3
+--+
| +----- Anodonta_oregonensis__2
+--+
| +-- Anodonta_oregonensis__1
+--+
+-- Anodonta_kennerlyi
Wu et al., 2024 [Publ 4379], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 7 | 5 | 6 | 46 | 11 | 2 | 35 | +-------------------- Cristariina
|
| +----------------- Anodonta_nuttalliana
--+ |
| | +-------- Pseudanodonta
| | +-----+
+--+ | | +----- Anodonta_anatina
| | +--+
| | | +-- Anodonta_cygnea
| | +--+
+--+ +-- Anodonta_exulcerata
|
| +-- Pyganodon **
| +--------+
| | +-- Utterbackia **
+--+
| +-------- Pterosyna **
+--+
| +----- Prolasmidonta **
+--+
| +-- Alasmidonta **
+--+
+-- Platynaias **
Wu et al., 2024 [Publ 4350], fig. 8b_S2 iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 7 | 5 | 6 | 85 | 11 | 2 | 35 | +-------------------- Cristariina
|
| +----------------- Anodonta_nuttalliana
--+ |
| | +-------- Pseudanodonta
| | +-----+
+--+ | | +----- Anodonta_anatina
| | +--+
| | | +-- Anodonta_cygnea
| | +--+
+--+ +-- Anodonta_exulcerata
|
| +-- Pyganodon **
| +--------+
| | +-- Utterbackia **
+--+
| +-------- Pterosyna **
+--+
| +----- Prolasmidonta **
+--+
| +-- Alasmidonta **
+--+
+-- Platynaias **
Hoeh et al., 1995 [Publ 3391], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 7 | 4 | 2 | 0 | 0 | 0 | 39 | +-- Utterbackia **
--+
+-- Pyganodon **
Smith et al., 2023 [Publ 4291], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 7 | 5 | 2 | 0 | 0 | 0 | 29 | +-- Anodontoides **
--+
+-- Strophitus **
Wu et al., 2023 [Publ 4339], fig. 4_S5-aa iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 7 | 5 | 6 | 89 | 11 | 2 | 37 | +-------------- Cristariina
|
--+ +----------- Anodontina
| |
| | +-- Pyganodon **
+--+ +-----+
| | +-- Utterbackia **
| |
+--+ +-- Alasmidonta **
| +--+
| | +-- Platynaias **
+--+
| +-- Prolasmidonta **
+--+
+-- Pterosyna **
Wu et al., 2023 [Publ 4339], fig. 4_S5-nt iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 7 | 5 | 6 | 89 | 11 | 2 | 35 | +-------------------- Cristariina
|
| +----------------- Anodonta_nuttalliana
--+ |
| +----------------- Pseudanodonta
| |
+--+ +-------------- Anodonta_cygnea
| |
| | +-- Anodonta_exulcerata
| +-----------+
+--+ +-- Anodonta_anatina
|
| +-- Pyganodon **
| +--------+
| | +-- Utterbackia **
+--+
| +-------- Pterosyna **
+--+
| +----- Prolasmidonta **
+--+
| +-- Alasmidonta **
+--+
+-- Platynaias **
Wu et al., 2021 [Publ 4366], fig. 3-BI iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 7 | 5 | 5 | 67 | 6 | 2 | 33 | +----------------- Cristariina
|
--+ +-------------- Anodonta_cygnea
| |
+--+ +----------- Anodonta_anatina
| |
+--+ +-- Pyganodon **
| +-----+
| | +-- Utterbackia **
+--+
| +----- Prolasmidonta **
+--+
| +-- Alasmidonta **
+--+
+-- Platynaias **
Huang et al., 2019 [Publ 3843], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 11 | 4 | 6 | 194 | 13 | 2 | 39 | +-------- Cristariina
|
| +----- Alasmidonta **
| |
--+ +----- Lasmigona **
| |
| +----- Pseudanodonta
| |
+--+----- Simpsonaias **
|
+----- Strophitus **
|
+----- Anodonta
|
| +-- Pyganodon **
+--+
+-- Utterbackia **
Huang et al., 2019 [Publ 3843], fig. S4 iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 47 | 24 | 6 | 194 | 13 | 2 | 74 | +-------- Acuticosta
|
+-------- Alasmidonta **
|
+-------- Anodonta_anatina
|
+-------- Anemina
|
+-------- Anodonta_cygnea
|
+-------- Cristaria
|
+-------- Anodonta_nuttalliana
|
+-------- Lasmigona **
|
+-------- Lepidodesma
|
+-------- Pseudanodonta
|
+-------- Pyganodon **
|
--+-------- Simpsonaias **
|
+-------- Strophitus **
|
+-------- Utterbackia **
|
+-------- Lanceolaria
|
+-------- Schistodesmus
|
+-------- Unio
|
+-------- Aculamprotula
|
| +-- Pletholophus
+-----+
| +-- Sinanodonta
|
| +----- Cuneopsis_celtiformis
| |
| +----- Tchangsinaia
| |
+--+----- Cuneopsis_rufescens
|
+----- Nodularia
|
| +-- Cuneopsis_szechenyii
+--+
+-- Cuneopsis_heudei
Wu et al., 2021 [Publ 4366], fig. 3-ML iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 7 | 5 | 5 | 67 | 6 | 2 | 33 | +----------------- Cristariina
|
--+ +-------------- Anodonta_cygnea
| |
+--+ +----------- Anodonta_anatina
| |
+--+ +-- Pyganodon **
| +-----+
| | +-- Utterbackia **
+--+
| +----- Prolasmidonta **
+--+
| +-- Alasmidonta **
+--+
+-- Platynaias **
Hou et al., 2025 [Publ 4445], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 7 | 5 | 6 | 46 | 11 | 2 | 35 | +-------------------- Cristariina
|
| +----------------- Anodonta_nuttalliana
--+ |
| | +-------- Pseudanodonta
| | +-----+
+--+ | | +----- Anodonta_anatina
| | +--+
| | | +-- Anodonta_cygnea
| | +--+
+--+ +-- Anodonta_exulcerata
|
| +-- Pyganodon **
| +--------+
| | +-- Utterbackia **
+--+
| +-------- Pterosyna **
+--+
| +----- Prolasmidonta **
+--+
| +-- Platynaias **
+--+
+-- Alasmidonta **
Huang et al., 2019 [Publ 3843], fig. S3 iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 11 | 8 | 6 | 192 | 13 | 2 | 39 | +-------------- Cristariina
|
| +----------- Anodonta
--+ |
| | +-- Alasmidonta **
| +--------+
+--+ +-- Lasmigona **
|
| +-- Pyganodon **
| +-----+
| | +-- Utterbackia **
+--+
| +----- Pseudanodonta
+--+
| +-- Simpsonaias **
+--+
+-- Strophitus **
Hou et al., 2024 [Publ 4400], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 7 | 5 | 6 | 69 | 11 | 2 | 35 | +-------------------- Cristariina
|
| +-------- Pseudanodonta
--+ +--------+
| | | +----- Anodonta_anatina
| | +--+
| | | +-- Anodonta_cygnea
+--+ +--+
| +-- Anodonta_exulcerata
|
| +-------------- Anodonta_nuttalliana
| |
+--+ +-- Pyganodon **
| +--------+
| | +-- Utterbackia **
+--+
| +-------- Pterosyna **
+--+
| +----- Prolasmidonta **
+--+
| +-- Alasmidonta **
+--+
+-- Platynaias **
Inoue et al., 2014 [Publ 3635], fig. 3B iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 7 | 2 | 3 | 1 | 0 | 0 | 37 | +----- Pyganodon_grandis **
|
+----- Pyganodon_cataracta **
|
--+----- Pyganodon_fragilis **
|
+----- Pyganodon **
|
| +-- Platynaias **
+--+
+-- Arcidens **
Wu et al., 2024 [Publ 4350], fig. 8a_S1 iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 7 | 5 | 6 | 85 | 11 | 2 | 35 | +-------------------- Cristariina
|
| +-------- Pseudanodonta
--+ +--------+
| | | +----- Anodonta_anatina
| | +--+
| | | +-- Anodonta_cygnea
+--+ +--+
| +-- Anodonta_exulcerata
|
| +-------------- Anodonta_nuttalliana
| |
+--+ +-- Pyganodon **
| +--------+
| | +-- Utterbackia **
+--+
| +-------- Pterosyna **
+--+
| +----- Prolasmidonta **
+--+
| +-- Alasmidonta **
+--+
+-- Platynaias **
King et al., 1999 [Publ 2699], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 3 | 5 | 0 | 0 | 0 | 32 | +-- Pterosyna **
+--+
| +-- Lasmigona **
--+
| +-- Prolasmidonta **
| |
+--+-- Strophitus **
|
+-- Platynaias **
Hewitt et al., 2018 [Publ 3758], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 3 | 6 | 3 | 0 | 0 | 33 | +-------- Pyganodon **
|
| +-- Platynaias **
--+-----+
| +-- Strophitus **
|
| +----- Lasmigona **
+--+
| +-- Arcidens **
+--+
+-- Pterosyna **
Boyer et al., 2011 [Publ 2718], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 2 | 6 | 29 | 0 | 0 | 34 | +----- Alasmidonta **
|
+----- Pyganodon **
|
--+ +-- Pterosyna **
+--+
| +-- Lasmigona **
|
| +-- Platynaias **
+--+
+-- Strophitus **
Lydeard et al., 1996 [Publ 2656], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 4 | 5 | 21 | 0 | 0 | 39 | +-- Utterbackia **
+-----+
| +-- Utterbackiana **
--+
| +----- Strophitus **
+--+
| +-- Lasmigona **
+--+
+-- Alasmidonta **
Sano et al., 2020 [Publ 4138], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 4 | 5 | 62 | 13 | 2 | 32 | +----------- Anodontina
|
--+ +-- Pyganodon **
| +-----+
| | +-- Utterbackia **
+--+
| +----- Alasmidonta **
+--+
| +-- Prolasmidonta **
+--+
+-- Platynaias **
Chen et al., 2020 [Publ 4170], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 34 | 32 | 5 | 28 | 5 | 2 | 63 | +----- Prolasmidonta **
+--+
| | +-- Alasmidonta **
| +--+
+-----------------+ +-- Anemina
| |
| | +----- Pseudodontina
| +--+
| | +-- Lamprotula_caveata
| +--+
| +-- Chamberlainia
|
| +----- Schistodesmus
| +-----+
| | | +-- Lepidodesma
--+ | +--+
| | +-- Aculamprotula
| +-----------+
| | | +----- Platynaias **
| | | +--+
| | | | | +-- Utterbackia_imbecillis **
| | | | +--+
| | +--+ +-- Pyganodon **
| | |
| | | +----- Sinanodonta
| | +--+
| | | +-- Cristaria
+--+ +--+
| +-- Anodontina
|
| +-- Utterbackia_peninsularis **
| +--------+
| | +-- Potamilus_alatus
| +--------+
| | | +-------- Lamprotula_leaii
| | +--+
| | | +----- Sinohyriopsis
| | +--+
+--+ | +-- Parvasolenaia
| +--+
| +-- Sinosolenaia
|
| +----------------- Gibbosula
+--+
| +-------------- Quadrulina
+--+
| +----------- Toxolasma
+--+
| +-------- Lampsilis_ornata
+--+
| +----- Potamilus_leptodon
+--+
| +-- Lampsilis_siliquoidea
+--+
+-- Lampsilis_powellii
Sano et al., 2020 [Publ 4138], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 4 | 5 | 62 | 13 | 2 | 32 | +----------- Anodontina
|
--+ +-- Pyganodon **
| +-----+
| | +-- Utterbackia **
+--+
| +----- Alasmidonta **
+--+
| +-- Prolasmidonta **
+--+
+-- Platynaias **
Wu et al., 2021 [Publ 4366], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 4 | 5 | 63 | 6 | 2 | 34 | +----------------- Cristariina
|
--+ +-------------- Anodonta_cygnea
| |
+--+ +----------- Anodonta_anatina
| |
+--+ +-- Pyganodon **
| +-----+
| | +-- Utterbackia **
+--+
| +----- Prolasmidonta **
+--+
| +-- Platynaias **
+--+
+-- Alasmidonta **
Sano et al., 2020 [Publ 4138], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 4 | 5 | 62 | 13 | 2 | 32 | +----------- Anodontina
|
--+ +-- Pyganodon **
| +-----+
| | +-- Utterbackia **
+--+
| +----- Alasmidonta **
+--+
| +-- Platynaias **
+--+
+-- Prolasmidonta **
Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 4 | 25 | 1 | 1 | 35 | +-------- Anodontina
|
--+ +-- Alasmidonta **
| +--+
| | +-- Lasmigona **
+--+
| +-- Strophitus **
+--+
+-- Pyganodon **
Guerra et al., 2017 [Publ 3772], fig. 2a iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 3 | 20 | 3 | 2 | 33 | +-------------- Sinanodonta
--+
| +----------- Cristaria
+--+
| +-------- Anodontina
+--+
| +----- Platynaias **
+--+
| +-- Pyganodon **
+--+
+-- Utterbackia **
Amaro et al., 2017 [Publ 3791], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 4 | 11 | 0 | 0 | 35 | +-- Lasmigona **
+--+
| +-- Platynaias **
--+
| +-- Pyganodon **
+--+
+-- Utterbackia **
Lydeard et al., 1996 [Publ 2656], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 6 | 4 | 5 | 23 | 1 | 1 | 39 | +----- Utterbackia **
+--+
| | +-- Utterbackiana **
| +--+
--+ +-- Anodontina
|
| +----- Strophitus **
+--+
| +-- Lasmigona **
+--+
+-- Alasmidonta **
Huang et al., 2002 [Publ 2657], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 5 | 29 | 2 | 1 | 38 | +-------------- Cristariina
--+
| +----------- Utterbackiana **
+--+
| +-------- Utterbackia **
+--+
| +----- Strophitus **
+--+
| +-- Alasmidonta **
+--+
+-- Lasmigona **
Wu et al., 2018 [Publ 3836], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 2 | 4 | 48 | 8 | 2 | 36 | +-------------- Cristariina
|
--+ +----------- Anodonta_cygnea
| |
+--+ +-------- Anodonta_anatina
| |
+--+ +----- Strophitus **
| |
+--+ +-- Platynaias **
| |
+--+-- Alasmidonta **
|
+-- Pyganodon **
Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 5 | 71 | 9 | 2 | 37 | +-------------- Cristariina
|
--+ +----------- Anodontina
| |
+--+ +-- Pyganodon **
| +-----+
| | +-- Simpsonaias **
+--+
| +----- Strophitus **
+--+
| +-- Alasmidonta **
+--+
+-- Lasmigona **
Bolotov et al., 2017 [Publ 3761], fig. S1 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 5 | 134 | 10 | 2 | 35 | +----------------- Cristariina
|
--+ +-------------- Anodonta_anatina
| |
| | +----- Pseudanodonta
+--+ +-----+
| | | +-- Anodonta_cygnea
| | +--+
| | +-- Anodonta_nuttalliana
+--+
| +-- Lasmigona **
| +-----+
| | +-- Alasmidonta **
+--+
| +----- Simpsonaias **
+--+
| +-- Pyganodon **
+--+
+-- Strophitus **
Guerra et al., 2017 [Publ 3772], fig. 2b iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 3 | 20 | 3 | 2 | 33 | +----------- Cristariina
--+
| +-------- Anodontina
+--+
| +----- Platynaias **
+--+
| +-- Pyganodon **
+--+
+-- Utterbackia **
Aunins et al., 2018 [Publ 3863], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 4 | 35 | 6 | 2 | 33 | +----------- Cristariina
|
--+ +-------- Anodontina
| |
+--+ +-- Prolasmidonta **
| +--+
| | +-- Alasmidonta **
+--+
| +-- Pyganodon **
+--+
+-- Utterbackia **
Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 4 | 25 | 1 | 1 | 35 | +-------- Anodontina
|
--+ +-- Alasmidonta **
| +--+
| | +-- Lasmigona **
+--+
| +-- Strophitus **
+--+
+-- Pyganodon **
Guerra et al., 2019 [Publ 4168], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 3 | 36 | 2 | 2 | 33 | +----------- Cristariina
--+
| +-------- Anodontina
+--+
| +----- Platynaias **
+--+
| +-- Pyganodon **
+--+
+-- Utterbackia **
Bolotov et al., 2017 [Publ 3761], fig. S2 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 1 | 5 | 134 | 10 | 2 | 36 | +-------- Anodonta_anatina
|
+-------- Pseudanodonta
|
| +-- Anodonta_cygnea
--+-----+
| +-- Anodonta_nuttalliana
|
| +----- Pyganodon **
| |
| +----- Simpsonaias **
+--+
+----- Strophitus **
|
| +-- Alasmidonta **
+--+
+-- Lasmigona **
Bolotov et al., 2018 [Publ 3826], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 5 | 78 | 5 | 2 | 37 | +----------------- Cristariina
--+
| +-------------- Anodontina
+--+
| +----------- Pyganodon **
+--+
| +-------- Simpsonaias **
+--+
| +----- Strophitus **
+--+
| +-- Alasmidonta **
+--+
+-- Lasmigona **
Bolotov et al., 2017 [Publ 3764], fig. S1 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 17 | 0 | 5 | 120 | 10 | 2 | 47 | +-- Alasmidonta **
|
+-- Anodonta_anatina
|
+-- Anodonta_cygnea
|
+-- Anodonta_nuttalliana
|
+-- Lasmigona **
|
+-- Pseudanodonta
|
+-- Pyganodon **
|
+-- Simpsonaias **
|
+-- Strophitus **
--+
+-- Sinanodonta_lauta
|
+-- Cristaria_plicata
|
+-- Cristaria_radiata
|
+-- Pletholophus
|
+-- Sinanodonta_lucida
|
+-- Sinanodonta_jourdyi
|
+-- Lanceolaria_gladiola
|
+-- Lanceolaria_grayii__1
|
+-- Lanceolaria_grayii__2
Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 5 | 75 | 9 | 2 | 37 | +-------------- Cristariina
|
--+ +----------- Anodontina
| |
+--+ +-- Pyganodon **
| +-----+
| | +-- Simpsonaias **
+--+
| +----- Strophitus **
+--+
| +-- Alasmidonta **
+--+
+-- Lasmigona **
Guerra et al., 2019 [Publ 4168], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 3 | 36 | 2 | 2 | 33 | +----------- Cristariina
--+
| +-------- Anodontina
+--+
| +----- Platynaias **
+--+
| +-- Pyganodon **
+--+
+-- Utterbackia **
Huang et al., 2002 [Publ 2657], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 14 | 5 | 5 | 29 | 2 | 1 | 46 | +-------- Strophitus **
|
+-------- Schistodesmus
|
+-------- Lepidodesma
|
+-------- Acuticosta
|
+-------- Cristariina
--+
+-------- Lanceolaria
|
| +-- Nodularia
+-----+
| +-- Tchangsinaia
|
| +----- Utterbackia **
| |
+--+----- Utterbackiana **
|
| +-- Alasmidonta **
+--+
+-- Lasmigona **
Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 5 | 71 | 9 | 2 | 35 | +-------------- Cristariina
|
--+ +----------- Anodontina
| |
+--+ +-- Alasmidonta **
| +-----+
| | +-- Lasmigona **
+--+
| +----- Pyganodon **
+--+
| +-- Strophitus **
+--+
+-- Simpsonaias **
Wu et al., 2019 [Publ 3864], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 3 | 42 | 5 | 2 | 34 | +----------- Cristariina
--+
| +-------- Anodontina
+--+
| +----- Platynaias **
+--+
| +-- Pyganodon **
+--+
+-- Utterbackia **
Wu et al., 2022 [Publ 4238], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 3 | 70 | 6 | 2 | 34 | +----------- Cristariina
--+
| +-------- Anodontina
+--+
| +----- Platynaias **
+--+
| +-- Pyganodon **
+--+
+-- Utterbackia **
Hoeh et al., 2009 [Publ 2716], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 42 | 0 | 0 | 32 | +-------- Lasmigona **
--+
| +----- Alasmidonta **
+--+
| +-- Strophitus **
+--+
+-- Pyganodon **
Hoeh et al., 2009 [Publ 2716], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 42 | 0 | 0 | 32 | +-------- Lasmigona **
--+
| +----- Alasmidonta **
+--+
| +-- Strophitus **
+--+
+-- Pyganodon **
Huang et al., 2019 [Publ 3843], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 3 | 52 | 6 | 2 | 34 | +-------------- Cristariina
--+
| +----------- Anodonta_cygnea
+--+
| +-------- Anodonta_anatina
+--+
| +----- Lasmigona **
+--+
| +-- Pyganodon **
+--+
+-- Utterbackia **
Hoeh et al., 2009 [Publ 2716], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 43 | 0 | 0 | 32 | +-------- Lasmigona **
--+
| +----- Alasmidonta **
+--+
| +-- Strophitus **
+--+
+-- Pyganodon **
Soroka & Burzynski, 2017 [Publ 3837], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 3 | 19 | 6 | 2 | 34 | +----------- Cristariina
--+
| +-------- Anodontina
+--+
| +----- Platynaias **
+--+
| +-- Pyganodon **
+--+
+-- Utterbackia **
Riccardi et al., 2020 [Publ 4089], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 3 | 15 | 9 | 2 | 34 | +----------------- Cristariina
|
--+ +-------------- Anodonta_anatina
| |
| | +----- Pseudanodonta
+--+ +-----+
| | | +-- Anodonta_exulcerata
| | +--+
+--+ +-- Anodonta_cygnea
|
| +-------- Anodonta_nuttalliana
+--+
| +----- Platynaias **
+--+
| +-- Pyganodon **
+--+
+-- Utterbackia **
Cyr et al., 2007 [Publ 2745], fig. 2D iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 1 | 0 | 0 | 0 | 38 | Fonseca et al., 2016 [Publ 3775], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 3 | 27 | 5 | 2 | 34 | +----------- Cristariina
--+
| +-------- Anodontina
+--+
| +----- Platynaias **
+--+
| +-- Pyganodon **
+--+
+-- Utterbackia **
Zhou et al., 2016 [Publ 3710], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 3 | 16 | 2 | 1 | 34 | +----------- Sinanodonta
--+
| +-------- Cristaria
+--+
| +----- Platynaias **
+--+
| +-- Pyganodon **
+--+
+-- Utterbackia **
Fukata & Iigo, 2020 [Publ 4106], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 50 | 6 | 2 | 32 | +----------- Cristariina
|
--+ +-------- Anodontina
| |
+--+ +-- Pyganodon **
| +--+
| | +-- Utterbackia **
+--+
| +-- Alasmidonta **
+--+
+-- Prolasmidonta **
Amaro et al., 2017 [Publ 3791], fig. 3B iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 3 | 8 | 0 | 0 | 34 | +----- Platynaias **
--+
| +-- Pyganodon **
+--+
+-- Utterbackia **
Amaro et al., 2017 [Publ 3791], fig. 3A iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 3 | 8 | 0 | 0 | 34 | +----- Platynaias **
--+
| +-- Pyganodon **
+--+
+-- Utterbackia **
Cyr et al., 2007 [Publ 2745], fig. 2A iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 1 | 1 | 0 | 0 | 38 | Cha et al., 2018 [Publ 3916], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 3 | 34 | 5 | 2 | 34 | +----------- Cristariina
--+
| +-------- Anodontina
+--+
| +----- Platynaias **
+--+
| +-- Pyganodon **
+--+
+-- Utterbackia **
Zieritz et al., 2021 [Publ 4231], fig. 3-F_5 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 3 | 44 | 5 | 2 | 34 | +----------- Cristariina
--+
| +-------- Anodontina
+--+
| +----- Platynaias **
+--+
| +-- Pyganodon **
+--+
+-- Utterbackia **
Campbell et al., 2005 [Publ 2660], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 6 | 3 | 4 | 98 | 2 | 1 | 35 | +----- Pyganodon **
|
| +-- Lasmigona **
--+--+
| +-- Pseudodontoideus **
|
| +-- Alasminota **
+--+
+-- Anodontina
Froufe et al., 2019 [Publ 3936], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 3 | 40 | 5 | 2 | 34 | +----------- Cristariina
--+
| +-------- Anodontina
+--+
| +----- Pterosyna **
+--+
| +-- Pyganodon **
+--+
+-- Utterbackia **
Jadhav & Jamkhedkar, 2009 [Publ 3697], fig. 11 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 1 | 4 | 21 | 2 | 1 | 33 | +-------- Anodontina
|
--+ +----- Lasmigona **
| |
+--+ +-- Strophitus **
| |
+--+-- Alasmidonta **
|
+-- Pyganodon **
Dai et al., 2025 [Publ 4440], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 3 | 84 | 6 | 2 | 34 | +----------- Cristariina
--+
| +-------- Anodontina
+--+
| +----- Platynaias **
+--+
| +-- Pyganodon **
+--+
+-- Utterbackia **
Wu et al., 2022 [Publ 4238], fig. 2d iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 3 | 35 | 6 | 2 | 34 | +----------- Cristariina
--+
| +-------- Anodontina
+--+
| +----- Platynaias **
+--+
| +-- Pyganodon **
+--+
+-- Utterbackia **
Cyr et al., 2007 [Publ 2745], fig. 2B iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 3 | 1 | 1 | 1 | 1 | 38 | +-- Anodontina
--+
+-- Pyganodon **
Chen et al., 2025 [Publ 4448], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 3 | 82 | 7 | 2 | 34 | +----------- Cristariina
--+
| +-------- Anodontina
+--+
| +----- Platynaias **
+--+
| +-- Pyganodon **
+--+
+-- Utterbackia **
Lopes-Lima et al., 2017 [Publ 3792], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 3 | 28 | 5 | 2 | 34 | +----------- Cristariina
--+
| +-------- Anodontina
+--+
| +----- Platynaias **
+--+
| +-- Pyganodon **
+--+
+-- Utterbackia **
Huang et al., 2013 [Publ 3629], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 3 | 12 | 2 | 1 | 34 | +-------- Cristariina
--+
| +----- Platynaias **
+--+
| +-- Pyganodon **
+--+
+-- Utterbackia **
Huang et al., 2018 [Publ 3828], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 3 | 38 | 5 | 2 | 34 | +----------- Cristariina
--+
| +-------- Anodontina
+--+
| +----- Platynaias **
+--+
| +-- Pyganodon **
+--+
+-- Utterbackia **
Wu et al., 2020 [Publ 4121], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 3 | 45 | 5 | 2 | 34 | +----------- Cristariina
--+
| +-------- Anodontina
+--+
| +----- Platynaias **
+--+
| +-- Pyganodon **
+--+
+-- Utterbackia **
Froufe et al., 2019 [Publ 3936], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 3 | 40 | 5 | 2 | 34 | +----------- Cristariina
--+
| +-------- Anodontina
+--+
| +----- Pterosyna **
+--+
| +-- Pyganodon **
+--+
+-- Utterbackia **
Graf & Cummings, 2006 [Publ 1968], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 4 | 43 | 0 | 0 | 32 | +-------- Lasmigona **
--+
| +----- Alasmidonta **
+--+
| +-- Pyganodon **
+--+
+-- Strophitus **
Campbell et al., 2005 [Publ 2660], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 5 | 3 | 4 | 98 | 2 | 1 | 35 | +----------- Anodonta_oregonensis
|
--+ +-- Anodonta_cygnea
| +-----+
| | +-- Alasminota **
+--+
| +----- Pyganodon **
+--+
| +-- Lasmigona **
+--+
+-- Pseudodontoideus **
Cyr et al., 2007 [Publ 2745], fig. 2C iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 3 | 1 | 1 | 1 | 1 | 38 | +-- Anodontina
--+
+-- Pyganodon **
Wu et al., 2016 [Publ 3711], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 2 | 18 | 4 | 2 | 35 | +-------------- Sinanodonta
--+
| +----------- Anemina
+--+
| +-------- Cristaria
+--+
| +----- Anodontina
+--+
| +-- Pyganodon **
+--+
+-- Utterbackia **
Amaro et al., 2017 [Publ 3791], fig. 2-M iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 4 | 1 | 3 | 7 | 1 | 1 | 36 | +----- Pyganodon **
|
--+ +-- Lasmigona **
| |
+--+-- Cristariina
|
+-- Utterbackia **
Krebs et al., 2019 [Publ 4092], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 0 | 0 | 0 | 37 | Rosenberg et al., 1997 [Publ 2720], fig. 2B iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 17 | 0 | 2 | 19 | 1 | 1 | 50 | +----- Anodontina
|
| +-- Gonidea
| |
| +-- Pyganodon_grandis **
| |
| +-- Utterbackia **
| |
| +-- Pyganodon_cataracta **
--+ |
| +-- Lampsilis_straminea
| |
| +-- Lampsilis_teres
| |
| +-- Megalonaias
| |
| +-- Obliquaria
| |
+--+-- Theliderma
|
+-- Quadrula
|
+-- Uniomerus
|
+-- Amblema
|
+-- Elliptio
|
+-- Fusconaia
|
+-- Plectomerus
|
+-- Aculamprotula
|
+-- Unio
Dai et al., 2024 [Publ 4358], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 50 | 4 | 2 | 34 | +----------- Cristariina
--+
| +-------- Anodontina
+--+
| +----- Platynaias **
+--+
| +-- Pyganodon **
+--+
+-- Utterbackia **
Moyer & Díaz-Ferguson, 2012 [Publ 3375], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 2 | 3 | 0 | 0 | 33 | +-- Lasmigona **
--+
+-- Platynaias **
Wu et al., 2020 [Publ 4147], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 2 | 27 | 4 | 2 | 35 | +-------- Cristariina
--+
| +----- Anodontina
+--+
| +-- Pyganodon **
+--+
+-- Utterbackia **
Wu et al., 2021 [Publ 4366], fig. 2_I iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 6 | 4 | 3 | 88 | 7 | 2 | 36 | +-------------- Cristariina
--+
| +----------- Lasmigona **
+--+
| +-------- Anodonta
+--+
| +----- Pseudanodonta
+--+
| +-- Pyganodon **
+--+
+-- Strophitus **
Liu et al., 2023 [Publ 4310], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 21 | 1 | 1 | 33 | +-------- Anodontina
--+
| +----- Lasmigona **
+--+
| +-- Strophitus **
+--+
+-- Pyganodon **
Liu et al., 2023 [Publ 4310], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 4 | 2 | 3 | 21 | 1 | 1 | 34 | +-------- Lasmigona **
--+
| +----- Anodontina
+--+
| +-- Pyganodon **
+--+
+-- Strophitus **
Wu et al., 2021 [Publ 4366], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 5 | 3 | 3 | 88 | 7 | 2 | 35 | +-------------- Cristariina
|
--+ +----------- Anodonta_cygnea
| |
+--+ +-- Lasmigona **
| +-----+
| | +-- Anodonta_anatina
+--+
| +----- Pseudanodonta
+--+
| +-- Pyganodon **
+--+
+-- Strophitus **
Rosenberg et al., 1997 [Publ 2720], fig. 2A iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 17 | 0 | 2 | 19 | 1 | 1 | 50 | +----- Anodontina
|
| +-- Gonidea
| |
| +-- Pyganodon_grandis **
| |
| +-- Utterbackia **
| |
| +-- Pyganodon_cataracta **
--+ |
| +-- Lampsilis_straminea
| |
| +-- Lampsilis_teres
| |
| +-- Megalonaias
| |
| +-- Obliquaria
| |
+--+-- Theliderma
|
+-- Quadrula
|
+-- Uniomerus
|
+-- Amblema
|
+-- Elliptio
|
+-- Fusconaia
|
+-- Plectomerus
|
+-- Pleurobema
|
+-- Unio
Wu et al., 2018 [Publ 4143], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 2 | 19 | 4 | 2 | 35 | +-------------- Sinanodonta
--+
| +----------- Anemina
+--+
| +-------- Cristaria
+--+
| +----- Anodontina
+--+
| +-- Pyganodon **
+--+
+-- Utterbackia **
Wu et al., 2023 [Publ 4296], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 20 | 1 | 1 | 33 | +-------- Anodontina
--+
| +----- Lasmigona **
+--+
| +-- Strophitus **
+--+
+-- Pyganodon **
Rosenberg et al., 1994 [Publ 2719], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 10 | 0 | 2 | 16 | 0 | 0 | 43 | +-- Pyganodon_cataracta **
|
+-- Pyganodon_grandis **
|
+-- Utterbackia **
|
+-- Lampsilis_teres
|
+-- Lampsilis_straminea
--+
+-- Megalonaias
|
+-- Obliquaria
|
+-- Theliderma
|
+-- Quadrula
|
+-- Uniomerus
Pfeiffer & Graf, 2013 [Publ 3531], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 28 | 2 | 1 | 35 | +-------- Cristariina
--+
| +----- Strophitus **
+--+
| +-- Platynaias **
+--+
+-- Pyganodon **
Dai et al., 2023 [Publ 4285], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 50 | 4 | 2 | 34 | +----------- Cristariina
--+
| +-------- Anodontina
+--+
| +----- Platynaias **
+--+
| +-- Pyganodon **
+--+
+-- Utterbackia **
Amaro et al., 2017 [Publ 3791], fig. 3B-M iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 5 | 0 | 0 | -- | +-- Utterbackia **
--+
+-- Pyganodon **
Wu et al., 2024 [Publ 4388], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 52 | 5 | 2 | -- | +-------- Cristariina
--+
| +----- Anodontina
+--+
| +-- Alasmidonta **
+--+
+-- Utterbackia **
Wu et al., 2022 [Publ 4238], fig. 2e iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 16 | 2 | 2 | -- | +-------- Cristariina
--+
| +----- Anodontina
+--+
| +-- Utterbackia **
+--+
+-- Pyganodon **
Huang et al., 2013 [Publ 3629], fig. 3-M iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 4 | 0 | 0 | -- | +-- Pyganodon **
--+
+-- Utterbackia **
Curole & Kocher, 2005 [Publ 2682], fig. 1-M iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 10 | 0 | 0 | -- | +-- Utterbackiana **
--+
+-- Pyganodon **
Araujo et al., 2018 [Publ 3779], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 16 | 1 | 1 | -- | +----- Anodontina
--+
| +-- Pyganodon **
+--+
+-- Alasmidonta **
Bogan & Hoeh, 2000 [Publ 2650], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 32 | 1 | 1 | -- | +----- Anodontina
--+
| +-- Strophitus **
+--+
+-- Pyganodon **
Burzynski & Soroka, 2018 [Publ 3862], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 2 | 2 | 2 | -- | +-------- Cristariina
--+
| +----- Anodontina
+--+
| +-- Utterbackia **
+--+
+-- Pyganodon **
Hoeh et al., 2002 [Publ 2653], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 29 | 1 | 1 | -- | +----- Anodontina
--+
| +-- Pyganodon **
+--+
+-- Strophitus **
Wang et al., 2018 [Publ 3846], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 29 | 5 | 2 | -- | +-------- Cristariina
--+
| +----- Anodontina
+--+
| +-- Pyganodon **
+--+
+-- Platynaias **
Dai et al., 2025 [Publ 4440], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 43 | 8 | 2 | -- | +----- Anodontina
--+
| +-- Alasmidonta **
+--+
+-- Lasmigona **
Soroka & Burzynski, 2018 [Publ 3762], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 5 | 1 | 1 | -- | +----- Anodontina
--+
| +-- Pyganodon **
+--+
+-- Utterbackia **
Walker et al., 2006 [Publ 2666], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 13 | 1 | 1 | -- | +-- Anodontina
--+
+-- Pyganodon **
Froufe et al., 2019 [Publ 3936], fig. 3-M iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 17 | 2 | 2 | -- | +-------- Cristariina
--+
| +----- Anodontina
+--+
| +-- Utterbackia **
+--+
+-- Pyganodon **
Manendo et al., 2008 [Publ 2669], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 11 | 3 | 1 | -- | +-------- Anodonta
--+
| +----- Pseudanodonta
+--+
| +-- Platynaias **
+--+
+-- Pyganodon **
Roe & Hoeh, 2003 [Publ 2655], fig. 4f3 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 29 | 1 | 1 | -- | +----- Anodontina
--+
| +-- Pyganodon **
+--+
+-- Strophitus **
Walker et al., 2006 [Publ 2666], fig. 6-M iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 12 | 0 | 0 | -- | Chen et al., 2023 [Publ 4302], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 39 | 8 | 2 | -- | +-------- Cristariina
--+
| +----- Anodontina
+--+
| +-- Alasmidonta **
+--+
+-- Lasmigona **
Wu et al., 2023 [Publ 4339], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 17 | 7 | 2 | -- | +----- Cristariina
--+
| +-- Platynaias **
+--+
+-- Anodontina
Graf, 2002 [Publ 2651], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 16 | 0 | 0 | -- | +-- Alasmidonta **
--+
+-- Pyganodon **
Walker et al., 2006 [Publ 2666], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 34 | 0 | 0 | -- | Guerra et al., 2017 [Publ 3772], fig. 2b-M iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 6 | 0 | 0 | -- | +-- Utterbackia **
--+
+-- Pyganodon **
Guerra et al., 2017 [Publ 3772], fig. 2a-M iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 6 | 0 | 0 | -- | +-- Utterbackia **
--+
+-- Pyganodon **
Chase et al., 2018 [Publ 3861], fig. 1-M iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 9 | 1 | 1 | -- | +----- Anodontina
--+
| +-- Utterbackia **
+--+
+-- Pyganodon **
Krebs, 2004 [Publ 2692], fig. 2-M iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 2 | 0 | 0 | -- | +-- Lasmigona **
--+
+-- Pyganodon **
Soroka & Burzynski, 2018 [Publ 3762], fig. 3-M iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 5 | 1 | 1 | -- | +----- Anodontina
--+
| +-- Pyganodon **
+--+
+-- Utterbackia **
Huang et al., 2019 [Publ 3843], fig. 3-M iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 21 | 2 | 2 | -- | +-------- Cristariina
--+
| +----- Anodontina
+--+
| +-- Pyganodon **
+--+
+-- Utterbackia **
Huang et al., 2018 [Publ 3828], fig. 6-M iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 15 | 2 | 2 | -- | +-------- Cristariina
--+
| +----- Anodontina
+--+
| +-- Pyganodon **
+--+
+-- Utterbackia **
Galal-Khallaf et al., 2024 [Publ 4459], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 26 | 11 | 2 | -- | +-------------- Cristariina
|
| +-- Anodonta_exulcerata
--+ +--------+
| | +-- Anodonta_cygnea
| |
+--+ +-- Anodonta_californiensis
| +-----+
| | +-- Anodonta_nuttalliana
| |
+--+ +-- Anodonta_kennerlyi
| +--+
| | +-- Anodonta_oregonensis
+--+
| +-- Anodonta_anatina
+--+
+-- Utterbackiana **
Krebs, 2004 [Publ 2692], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 3 | 0 | 0 | -- | +-- Lasmigona **
--+
+-- Pyganodon **
Dai et al., 2025 [Publ 4440], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 43 | 8 | 2 | -- | +----- Anodontina
--+
| +-- Alasmidonta **
+--+
+-- Lasmigona **
Chase et al., 2018 [Publ 3861], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 9 | 1 | 1 | -- | +----- Anodontina
--+
| +-- Utterbackia **
+--+
+-- Pyganodon **
Guerra et al., 2019 [Publ 4168], fig. 2-M iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 20 | 2 | 2 | -- | +-- Utterbackia **
+--+
| +-- Pyganodon **
--+
| +-- Anodontina
+--+
+-- Cristariina
Froufe et al., 2019 [Publ 3936], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 17 | 2 | 2 | -- | +-------- Cristariina
--+
| +----- Anodontina
+--+
| +-- Utterbackia **
+--+
+-- Pyganodon **
Burzynski & Soroka, 2018 [Publ 3862], fig. 5b iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 8 | 2 | 2 | -- | +-------- Cristariina
--+
| +----- Anodontina
+--+
| +-- Utterbackia **
+--+
+-- Pyganodon **
Amaro et al., 2017 [Publ 3791], fig. 3A-M iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 5 | 0 | 0 | -- | +-- Pyganodon **
--+
+-- Utterbackia **
Walker et al., 2006 [Publ 2666], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 13 | 1 | 1 | -- | +-- Anodontina
--+
+-- Pyganodon **
Feng et al., 2016 [Publ 3800], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 12 | 0 | 2 | 12 | 4 | 2 | 44 | +-- Anodontina
+-----------+
| +-- Utterbackia **
|
--+ +----- Tchangsinaia
| +-----+
| | | +-- Pyganodon **
| | +--+
+--+ +-- Cristariina
|
| +-------- Sinosolenaia
+--+
| +----- Quadrulina
+--+
| +-- Potamilus
+--+
+-- Lampsilis
Chen et al., 2023 [Publ 4302], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 39 | 8 | 2 | -- | +----- Anodontina
--+
| +-- Alasmidonta **
+--+
+-- Lasmigona **
Roe & Hoeh, 2003 [Publ 2655], fig. 4f2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 3 | 0 | 2 | 29 | 1 | 1 | 35 | +----- Pyganodon **
--+
| +-- Anodontina
+--+
+-- Strophitus **
Dai et al., 2024 [Publ 4386], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 39 | 1 | 1 | -- | +----- Cristariina
--+
| +-- Pyganodon **
+--+
+-- Utterbackia **
Walker et al., 2006 [Publ 2666], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 13 | 1 | 1 | -- | +-- Anodontina
--+
+-- Pyganodon **
Smith, 2021 [Publ 4331], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 18 | 1 | 1 | -- | +----- Cristariina
--+
| +-- Pyganodon **
+--+
+-- Utterbackia **
Zieritz et al., 2021 [Publ 4231], fig. 3-M_4 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 21 | 2 | 2 | -- | +-------- Cristariina
--+
| +----- Anodontina
+--+
| +-- Utterbackia **
+--+
+-- Pyganodon **
Lopes-Lima et al., 2017 [Publ 3792], fig. 2-M iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 8 | 1 | 1 | -- | +----- Anodontina
--+
| +-- Utterbackia **
+--+
+-- Pyganodon **
Hoeh et al., 1996 [Publ 2694], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 2 | 0 | 0 | -- | Burzynski & Soroka, 2018 [Publ 3862], fig. 5a iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 18 | 2 | 2 | -- | +----- Cristariina
|
--+----- Anodontina
|
| +-- Utterbackia **
+--+
+-- Pyganodon **
Pfeiffer & Graf, 2015 [Publ 3620], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 38 | 0 | 0 | -- | +-- Pyganodon **
--+
+-- Lasmigona **
Froufe et al., 2015 [Publ 3664], fig. 1-M iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 6 | 1 | 1 | -- | +----- Anodontina
--+
| +-- Utterbackia **
+--+
+-- Pyganodon **
Hoeh et al., 2001 [Publ 2652], fig. 14f2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 3 | 0 | 2 | 29 | 1 | 1 | 35 | +----- Pyganodon **
--+
| +-- Anodontina
+--+
+-- Strophitus **
Guerra et al., 2019 [Publ 4168], fig. 1-M iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 20 | 2 | 2 | -- | +-------- Cristariina
--+
| +----- Anodontina
+--+
| +-- Utterbackia **
+--+
+-- Pyganodon **
Hoeh et al., 2002 [Publ 2673], fig. 1-M iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 7 | 0 | 0 | -- | Burzynski et al., 2017 [Publ 3774], fig. 1-bottom iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 11 | 1 | 1 | -- | +----- Anodontina
--+
| +-- Utterbackia **
+--+
+-- Pyganodon **
Froufe et al., 2015 [Publ 3664], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 10 | 1 | 1 | -- | +----- Anodontina
--+
| +-- Utterbackia **
+--+
+-- Pyganodon **
Wen et al., 2017 [Publ 3795], fig. 3-M iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 4 | 1 | 1 | -- | +----- Anodontina
--+
| +-- Utterbackia **
+--+
+-- Pyganodon **
Fonseca et al., 2016 [Publ 3775], fig. 1-M iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 7 | 1 | 1 | -- | +----- Anodontina
--+
| +-- Utterbackia **
+--+
+-- Pyganodon **
Hoeh et al., 2002 [Publ 2673], fig. 1-F iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 7 | 0 | 0 | -- | Burzynski et al., 2017 [Publ 3774], fig. 1-top iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 11 | 1 | 1 | -- | +----- Anodontina
--+
| +-- Utterbackia **
+--+
+-- Pyganodon **
Whelan et al., 2011 [Publ 3380], fig. 1-ML iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 34 | 0 | 0 | -- | +-- Pyganodon **
--+
+-- Lasmigona **
Whelan et al., 2011 [Publ 3380], fig. 1-MP iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 34 | 0 | 0 | -- | +-- Pyganodon **
--+
+-- Lasmigona **
Hoeh et al., 2001 [Publ 2652], fig. 14f4 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 29 | 1 | 1 | -- | +----- Anodontina
--+
| +-- Strophitus **
+--+
+-- Pyganodon **
Wang et al., 2018 [Publ 3865], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 29 | 5 | 2 | -- | +-------- Cristariina
--+
| +----- Anodontina
+--+
| +-- Pyganodon **
+--+
+-- Platynaias **
Pavan-Kumar et al., 2022 [Publ 4263], fig. S5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 38 | 5 | 2 | -- | +----- Cristariina
--+
| +-- Anodontina
+--+
+-- Pyganodon **
Curole & Kocher, 2002 [Publ 2712], fig. 1b-M iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 3 | 0 | 0 | -- | -- Utterbackiana_implicata **
Lopes-Lima et al., 2019 [Publ 3901], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 27 | 1 | 1 | -- | +-- Anodontina
--+
+-- Pyganodon **
Curole & Kocher, 2005 [Publ 2682], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 10 | 0 | 0 | -- | Inoue et al., 2013 [Publ 3579], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 8 | 0 | 0 | -- | Zanatta & Murphy, 2006 [Publ 2672], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 56 | 0 | 0 | -- | Inoue et al., 2013 [Publ 3528], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 16 | 0 | 0 | -- | Pfeiffer et al., 2019 [Publ 3937], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 56 | 3 | 2 | -- | +----- Cristariina
--+
| +-- Anodontina
+--+
+-- Pyganodon **
Sheng et al., 2014 [Publ 3704], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 7 | 1 | 1 | -- | Plazzi et al., 2011 [Publ 3676], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 17 | 4 | 2 | -- | +-- Pyganodon **
--+
+-- Anodontina
Inoue et al., 2020 [Publ 3941], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 41 | 0 | 0 | -- | -- Arcidens_confragosus **
Graf & Ó Foighil, 2000 [Publ 2649], fig. 1_2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 10 | 0 | 0 | -- | Metzger et al., 2018 [Publ 3952], fig. 1C iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 2 | 0 | 0 | -- | Pfeiffer et al., 2019 [Publ 3937], fig. 4-5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 56 | 3 | 2 | -- | +----- Cristariina
--+
| +-- Anodontina
+--+
+-- Pyganodon **
Pavan-Kumar et al., 2022 [Publ 4263], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 36 | 4 | 2 | -- | +----- Cristariina
--+
| +-- Anodontina
+--+
+-- Pyganodon **
Marshall et al., 2014 [Publ 3593], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 13 | 0 | 0 | -- | Xue et al., 2016 [Publ 3659], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 14 | 4 | 1 | -- | +-- Cristaria
+--+
| +-- Platynaias **
--+
| +-- Anemina
+--+
+-- Sinanodonta
Wei et al., 2016 [Publ 3665], fig. 0 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 5 | 1 | 1 | -- | Hewitt et al., 2019 [Publ 3962], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 9 | 0 | 0 | -- | Pfeiffer et al., 2016 [Publ 3660], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 37 | 0 | 0 | -- | -- Utterbackiana_suborbiculata **
Zanatta & Harris, 2013 [Publ 3530], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 4 | 0 | 0 | -- | Pieri et al., 2018 [Publ 3867], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 19 | 0 | 0 | -- | -- Arcidens_confragosus **
Curole & Kocher, 2002 [Publ 2712], fig. 1a-M iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 3 | 0 | 0 | -- | -- Utterbackiana_implicata **
Soroka, 2008 [Publ 3695], fig. 1-M iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 3 | 2 | 2 | -- | +----- Cristariina
--+
| +-- Anodontina
+--+
+-- Pyganodon **
Lopes-Lima et al., 2019 [Publ 3901], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 25 | 1 | 1 | -- | +-- Anodontina
--+
+-- Pyganodon **
Zanatta & Murphy, 2006 [Publ 2672], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 55 | 0 | 0 | -- | Soroka, 2008 [Publ 3695], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 3 | 2 | 2 | -- | +----- Cristariina
--+
| +-- Anodontina
+--+
+-- Pyganodon **
Lopes-Lima et al., 2019 [Publ 3901], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 28 | 1 | 1 | -- | +-- Anodontina
--+
+-- Pyganodon **
Chase et al., 2018 [Publ 3847], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 9 | 1 | 1 | -- | +-- Anodontina
--+
+-- Utterbackia **
Campbell, 2000 [Publ 2724], fig. 1-5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 2 | 0 | 0 | -- | -- Utterbackia_imbecillis **
Turner et al., 2000 [Publ 2722], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 13 | 0 | 0 | -- | -- Utterbackiana_couperiana **
Graf, 2000 [Publ 2647], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 16 | 0 | 0 | -- | Inoue et al., 2020 [Publ 3941], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 40 | 0 | 0 | -- | -- Arcidens_confragosus **
Chen et al., 2012 [Publ 3624], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 6 | 1 | 1 | -- | +-- Pyganodon **
--+
+-- Cristariina
Mulvey et al., 1997 [Publ 3390], fig. 1_3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 4 | 0 | 0 | -- | -- Utterbackia_imbecillis **
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