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Page last updated
Thu 3 September 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Search the database pages:

Bivalvia | Unionida | Unionoidea | Unionidae | Unioninae | Anodontini

subtribe ANODONTINA Rafinesque, 1820: cladomics
     3 genera

Cladomics Summary

number of species = 11 (24 total)
number of subtaxa (rank: genus) = 3
analyzed in 226 cladograms in 122 publications
maximum number of terminals in an analysis = 10 (mean = 2.02, median = 1)
maximum number of subtaxa (genus) = 3 (mean = 1.43, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 110
maximum number of included terminals = 10, (mean = 3.09 median = 3)
maximum number of subtaxa (genus) = 3 (mean = 1.88, median = 2)
proportion of cladograms that support basic monophyly = 0.6

Rigorous Monophyly

number of cladograms that test rigorous monophyly = 59
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (genus) taxon and at least one rigorous outgroup (family).
maximum number of included terminals = 10 (mean = 3.63, median = 3)
maximum number of subtaxa (genus) = 3 (mean = 2.37, median = 2)
proportion of cladograms that support rigorous monophyly = 0.37

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.


PCoA Ordination


NJ Tree

Whelan et al., 2023 [Publ 4287], fig. 2

** Top Monophyly Cladogram **

iTtTbiriSoTroTroSiD
437383373720
  +----------- Cristariina     
  |                            
--+  +-------- Anodonta_cygnea ** 
  |  |                         
  +--+     +-- Brachyanodon **    
     |  +--+                   
     |  |  +-- Pseudanodonta **   
     +--+                      
        |  +-- Anodonta_anatina **
        +--+                   
           +-- Alasmidontina

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
10109362269247
  +----------------------- Anodonta_anatina **           
  |                                                   
--+  +-------------------- Pseudanodonta **              
  |  |                                                
  +--+  +----------------- Brachyanodon_oregonensis__2 **
     |  |                                             
     |  |           +----- Anodonta_cygnea **            
     +--+  +--------+                                 
        |  |        |  +-- Anodonta_seddonae **          
        |  |        +--+                              
        +--+           +-- Anodonta_exulcerata **        
           |                                          
           |  +----------- Brachyanodon_impura **        
           |  |                                       
           +--+  +-------- Brachyanodon **               
              |  |                                    
              +--+  +----- Brachyanodon_nuttalliana__1 **
                 |  |                                 
                 +--+----- Brachyanodon_nuttalliana__2 **
                    |                                 
                    |  +-- Brachyanodon_californiensis **
                    +--+                              
                       +-- Brachyanodon_oregonensis__1 **

Galal-Khallaf et al., 2024 [Publ 4459], fig. 6

iTtTbiriSoTroTroSiD
7972216239
  +-------------- Cristariina                
  |                                          
  |           +-- Anodonta_exulcerata **        
--+  +--------+                              
  |  |        +-- Anodonta_cygnea **            
  |  |                                       
  +--+        +-- Brachyanodon_californiensis **
     |  +-----+                              
     |  |     +-- Brachyanodon_nuttalliana **   
     |  |                                    
     +--+     +-- Brachyanodon_kennerlyi **     
        |  +--+                              
        |  |  +-- Brachyanodon_oregonensis **   
        +--+                                 
           |  +-- Anodonta_anatina **           
           +--+                              
              +-- Alasmidontina

Chong et al., 2008 [Publ 2668], fig. 3

iTtTbiriSoTroTroSiD
7181031110246
  +----------------- Anodonta_cygnea **            
  |                                             
  +----------------- Pyganodon                  
  |                                             
  +----------------- Alasminota                 
  |                                             
  +----------------- Anodonta_anatina **           
  |                                             
  +----------------- Unionina                   
  |                                             
  +----------------- Cristariina                
  |                                             
  |              +-- Brachyanodon_nuttalliana **   
--+--------------+                              
  |              +-- Brachyanodon_californiensis **
  |                                             
  |              +-- Pseudodontoideus           
  |              |                              
  +--------------+-- Alasmidonta                
  |              |                              
  |              +-- Pterosyna                  
  |                                             
  |  +-------------- Pseudanodonta **              
  |  |                                          
  |  |        +----- Strophitus                 
  |  +--------+                                 
  +--+        |  +-- Prolasmidonta              
     |        +--+                              
     |           +-- Platynaias                 
     |                                          
     |  +----------- Brachyanodon_oregonensis__4 **
     +--+                                       
        |  +-------- Brachyanodon_oregonensis__3 **
        +--+                                    
           |  +----- Brachyanodon_oregonensis__2 **
           +--+                                 
              |  +-- Brachyanodon_oregonensis__1 **
              +--+                              
                 +-- Brachyanodon_kennerlyi **

Wu et al., 2024 [Publ 4379], fig. 5

iTtTbiriSoTroTroSiD
5121034913238
  +----------- Cristariina  
--+                         
  |  +-------- Brachyanodon ** 
  +--+                      
     |  +----- Alasmidontina
     +--+                   
        |  +-- Pseudanodonta **
        +--+                
           +-- Anodonta **

Hou et al., 2025 [Publ 4445], fig. 5

iTtTbiriSoTroTroSiD
5121034813238
  +----------- Cristariina  
--+                         
  |  +-------- Brachyanodon ** 
  +--+                      
     |  +----- Alasmidontina
     +--+                   
        |  +-- Pseudanodonta **
        +--+                
           +-- Anodonta **

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iTtTbiriSoTroTroSiD
512839113236
  +----------- Cristariina        
  |                               
--+  +-------- Brachyanodon **       
  |  |                            
  |  +-------- Pseudanodonta **      
  +--+                            
     |  +----- Anodonta_cygnea **    
     |  |                         
     +--+----- Alasmidontina      
        |                         
        |  +-- Anodonta_exulcerata **
        +--+                      
           +-- Anodonta_anatina **

Hou et al., 2024 [Publ 4400], fig. 4

iTtTbiriSoTroTroSiD
5121037113238
  +-------- Cristariina  
  |                      
--+     +-- Pseudanodonta **
  |  +--+                
  |  |  +-- Anodonta **     
  +--+                   
     |  +-- Brachyanodon ** 
     +--+                
        +-- Alasmidontina

Riccardi et al., 2020 [Publ 4089], fig. 2

iTtTbiriSoTroTroSiD
5973148239
  +-------------- Cristariina        
  |                                  
--+  +----------- Anodonta_anatina **   
  |  |                               
  +--+        +-- Brachyanodon **       
     |  +-----+                      
     |  |     +-- Alasmidontina      
     +--+                            
        |  +----- Pseudanodonta **      
        +--+                         
           |  +-- Anodonta_exulcerata **
           +--+                      
              +-- Anodonta_cygnea **

Lyubas et al., 2022 [Publ 4255], fig. S1

iTtTbiriSoTroTroSiD
5532260041
  +----------- Anodonta_anatina **   
--+                               
  |  +-------- Pseudanodonta **      
  +--+                            
     |  +----- Anodonta_exulcerata **
     +--+                         
        |  +-- Anodonta_seddonae **  
        +--+                      
           +-- Anodonta_cygnea **

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iTtTbiriSoTroTroSiD
55339113239
  +----------- Cristariina  
--+                         
  |  +-------- Alasmidontina
  +--+                      
     |  +----- Anodonta **     
     +--+                   
        |  +-- Brachyanodon ** 
        +--+                
           +-- Pseudanodonta **

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iTtTbiriSoTroTroSiD
5121038713238
  +-------- Cristariina  
  |                      
--+     +-- Pseudanodonta **
  |  +--+                
  |  |  +-- Anodonta **     
  +--+                   
     |  +-- Brachyanodon ** 
     +--+                
        +-- Alasmidontina

Lyubas et al., 2022 [Publ 4255], fig. 7

iTtTbiriSoTroTroSiD
5532260041
  +-- Pseudanodonta **
--+                
  +-- Anodonta **

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iTtTbiriSoTroTroSiD
5121038713238
  +----------- Cristariina  
--+                         
  |  +-------- Brachyanodon ** 
  +--+                      
     |  +----- Alasmidontina
     +--+                   
        |  +-- Pseudanodonta **
        +--+                
           +-- Anodonta **

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

iTtTbiriSoTroTroSiD
51311313612235
  +-------------- Cristariina        
  |                                  
--+           +-- Anodonta_cygnea **    
  |  +--------+                      
  |  |        +-- Anodonta_exulcerata **
  +--+                               
     |  +-------- Anodonta_anatina **   
     +--+                            
        |  +----- Alasmidontina      
        +--+                         
           |  +-- Brachyanodon **       
           +--+                      
              +-- Pseudanodonta **

Tomilova et al., 2020 [Publ 4097], fig. 3

iTtTbiriSoTroTroSiD
442242140
  +-------- Anodonta_anatina **   
--+                            
  |  +----- Pseudanodonta **      
  +--+                         
     |  +-- Anodonta_cygnea **    
     +--+                      
        +-- Anodonta_exulcerata **

Bolotov et al., 2017 [Publ 3761], fig. S1

iTtTbiriSoTroTroSiD
497313411243
  +-------------- Cristariina     
--+                               
  |  +----------- Anodonta_anatina **
  +--+                            
     |  +-------- Alasmidontina   
     +--+                         
        |  +----- Pseudanodonta **   
        +--+                      
           |  +-- Anodonta_cygnea ** 
           +--+                   
              +-- Brachyanodon **

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
4118319416242
  +-------------- Cristariina     
  |                               
  |           +-- Alasmidonta     
  |  +--------+                   
--+  |        +-- Lasmigona       
  |  |                            
  |  |     +----- Anodonta_anatina **
  |  +-----+                      
  |  |     |  +-- Anodonta_cygnea ** 
  +--+     +--+                   
     |        +-- Brachyanodon **    
     |                            
     |        +-- Pyganodon       
     |  +-----+                   
     |  |     +-- Utterbackia     
     +--+                         
        |  +----- Pseudanodonta **   
        +--+                      
           |  +-- Simpsonaias     
           +--+                   
              +-- Strophitus

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iTtTbiriSoTroTroSiD
494311829241
  +----------- Cristariina     
  |                            
--+  +-------- Pseudanodonta **   
  |  |                         
  +--+-------- Alasmidontina   
     |                         
     |  +----- Anodonta_cygnea ** 
     +--+                      
        |  +-- Anodonta_anatina **
        +--+                   
           +-- Brachyanodon **

Bolotov et al., 2017 [Publ 3761], fig. S2

iTtTbiriSoTroTroSiD
493313411240
  +----- Anodonta_anatina **
  |                      
  +----- Pseudanodonta **   
--+                      
  +----- Alasmidontina   
  |                      
  |  +-- Anodonta_cygnea ** 
  +--+                   
     +-- Brachyanodon **

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
44724319616273
  +-------- Acuticosta           
  |                              
  +-------- Alasmidonta          
  |                              
  +-------- Anodonta_anatina **     
  |                              
  +-------- Anemina              
  |                              
  +-------- Anodonta_cygnea **      
  |                              
  +-------- Cristaria            
  |                              
  +-------- Brachyanodon **         
  |                              
  +-------- Lasmigona            
  |                              
  +-------- Lepidodesma          
  |                              
  +-------- Pseudanodonta **        
  |                              
  +-------- Pyganodon            
  |                              
--+-------- Simpsonaias          
  |                              
  +-------- Strophitus           
  |                              
  +-------- Utterbackia          
  |                              
  +-------- Lanceolaria          
  |                              
  +-------- Schistodesmus        
  |                              
  +-------- Unionina             
  |                              
  +-------- Aculamprotulina      
  |                              
  |     +-- Pletholophus         
  +-----+                        
  |     +-- Sinanodonta          
  |                              
  |  +----- Cuneopsis_celtiformis
  |  |                           
  |  +----- Tchangsinaia         
  |  |                           
  +--+----- Cuneopsis_rufescens  
     |                           
     +----- Nodularia            
     |                           
     |  +-- Cuneopsis_szechenyii 
     +--+                        
        +-- Cuneopsis_heudei

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iTtTbiriSoTroTroSiD
44137610239
  +----------- Cristariina     
  |                            
--+  +-------- Alasmidontina   
  |  |                         
  +--+  +----- Pseudanodonta **   
     |  |                      
     +--+  +-- Anodonta_anatina **
        |  |                   
        +--+-- Anodonta_cygnea ** 
           |                   
           +-- Brachyanodon **

Bolotov et al., 2017 [Publ 3764], fig. S1

iTtTbiriSoTroTroSiD
4170312011246
  +-- Alasmidonta          
  |                        
  +-- Anodonta_anatina **     
  |                        
  +-- Anodonta_cygnea **      
  |                        
  +-- Brachyanodon **         
  |                        
  +-- Lasmigona            
  |                        
  +-- Pseudanodonta **        
  |                        
  +-- Pyganodon            
  |                        
  +-- Simpsonaias          
  |                        
  +-- Strophitus           
--+                        
  +-- Sinanodonta_lauta    
  |                        
  +-- Cristaria_plicata    
  |                        
  +-- Cristaria_radiata    
  |                        
  +-- Pletholophus         
  |                        
  +-- Sinanodonta_lucida   
  |                        
  +-- Sinanodonta_jourdyi  
  |                        
  +-- Lanceolaria_gladiola 
  |                        
  +-- Lanceolaria_grayii__1
  |                        
  +-- Lanceolaria_grayii__2

Wu et al., 2026 [Publ 4811], fig. 4

iTtTbiriSoTroTroSiD
411929222237
  +-------- Cristariina  
--+                      
  |  +----- Anodonta **     
  +--+                   
     |  +-- Pseudanodonta **
     +--+                
        +-- Alasmidontina

Nagel et al., 1996 [Publ 3400], fig. 351

iTtTbiriSoTroTroSiD
454211141
  +-------------- Pseudanodonta **   
--+                               
  |  +----------- Anodonta_cygnea ** 
  +--+                            
     |  +-------- Cristariina     
     +--+                         
        |  +----- Anodonta__2 **     
        +--+                      
           |  +-- Anodonta_anatina **
           +--+                   
              +-- Anodonta__1 **

Nagel et al., 1998 [Publ 1946], fig. 7

iTtTbiriSoTroTroSiD
454211141
  +-------------- Pseudanodonta **   
--+                               
  |  +----------- Anodonta_cygnea ** 
  +--+                            
     |  +-------- Cristariina     
     +--+                         
        |  +----- Anodonta__2 **     
        +--+                      
           |  +-- Anodonta_anatina **
           +--+                   
              +-- Anodonta__1 **

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

iTtTbiriSoTroTroSiD
44237210240
  +----------- Cristariina  
--+                         
  |  +-------- Alasmidontina
  +--+                      
     |  +----- Pseudanodonta **
     +--+                   
        |  +-- Brachyanodon ** 
        +--+                
           +-- Anodonta **

Riccardi et al., 2020 [Publ 4089], fig. 1

iTtTbiriSoTroTroSiD
442222138
  +-------- Cristariina        
  |                            
--+     +-- Anodonta_anatina **   
  |  +--+                      
  |  |  +-- Pseudanodonta **      
  +--+                         
     |  +-- Anodonta_exulcerata **
     +--+                      
        +-- Anodonta_cygnea **

Lopes-Lima et al., 2021 [Publ 4217], fig. 11

iTtTbiriSoTroTroSiD
442100040
1

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
4114319616239
  +----------- Cristariina     
  |                            
  |  +-------- Alasmidonta     
  |  |                         
--+  +-------- Lasmigona       
  |  |                         
  |  +-------- Pseudanodonta **   
  |  |                         
  |  +-------- Simpsonaias     
  +--+                         
     +-------- Strophitus      
     |                         
     |     +-- Pyganodon       
     +-----+                   
     |     +-- Utterbackia     
     |                         
     |  +----- Anodonta_anatina **
     +--+                      
        |  +-- Anodonta_cygnea ** 
        +--+                   
           +-- Brachyanodon **

Wang et al., 2025 [Publ 4488], fig. 3B

iTtTbiriSoTroTroSiD
411926122237
  +-------- Cristariina  
--+                      
  |  +----- Anodonta **     
  +--+                   
     |  +-- Pseudanodonta **
     +--+                
        +-- Alasmidontina

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

iTtTbiriSoTroTroSiD
44237210240
  +-------------- Cristariina     
--+                               
  |  +----------- Alasmidontina   
  +--+                            
     |  +-------- Pseudanodonta **   
     +--+                         
        |  +----- Anodonta_anatina **
        +--+                      
           |  +-- Brachyanodon **    
           +--+                   
              +-- Anodonta_cygnea **

Konopleva et al., 2023 [Publ 4269], fig. S1_4

iTtTbiriSoTroTroSiD
33125313139
  +-------- Cristariina     
--+                         
  |  +----- Anodonta_anatina **
  +--+                      
     |  +-- Pseudanodonta **   
     +--+                   
        +-- Anodonta_cygnea **

Mezhzherin et al., 2014 [Publ 3705], fig. 2

iTtTbiriSoTroTroSiD
331211139
  +----- Cristariina  
--+                   
  |  +-- Pseudanodonta **
  +--+                
     +-- Anodonta **

Soroka, 2010 [Publ 3404], fig. 4

iTtTbiriSoTroTroSiD
375241143
        +-- Anodonta_anatina **
  +-----+                   
  |     +-- Unionina        
--+                         
  |  +----- Cristariina     
  +--+                      
     |  +-- Anodonta_cygnea ** 
     +--+                   
        +-- Pseudanodonta **

Inoue et al., 2018 [Publ 3858], fig. 1_3a-c

iTtTbiriSoTroTroSiD
3200110217242
  +-- Alasmidonta_marginata      
  |                              
  +-- Prolasmidonta              
  |                              
  +-- Alasmidonta_undulata       
  |                              
  +-- Cristariina                
  |                              
  +-- Brachyanodon_kennerlyi **     
  |                              
  +-- Brachyanodon_nuttalliana **   
  |                              
  +-- Brachyanodon_oregonensis **   
  |                              
  +-- Arcidens_confragosus       
  |                              
  +-- Arcidens_wheeleri          
  |                              
  +-- Pterosyna                  
  |                              
--+-- Platynaias_compressa       
  |                              
  +-- Lasmigona                  
  |                              
  +-- Platynaias_decorata        
  |                              
  +-- Platynaias_subviridis      
  |                              
  +-- Pyganodon_grandis          
  |                              
  +-- Pyganodon_lacustris        
  |                              
  +-- Pseudodontoideus           
  |                              
  +-- Strophitus                 
  |                              
  +-- Utterbackia_imbecillis     
  |                              
  +-- Utterbackia_peninsularis__1
  |                              
  +-- Utterbackia_peninsularis__2

Lopes-Lima et al., 2021 [Publ 4217], fig. 1

iTtTbiriSoTroTroSiD
3311220039
1

Hinzmann et al., 2013 [Publ 3541], fig. 5

iTtTbiriSoTroTroSiD
331200039
  +-- Pseudanodonta **
--+                
  +-- Anodonta **

Froufe et al., 2014 [Publ 3592], fig. 2

iTtTbiriSoTroTroSiD
331200039
  +----- Anodonta_anatina **
--+                      
  |  +-- Pseudanodonta **   
  +--+                   
     +-- Anodonta_cygnea **

Manendo et al., 2008 [Publ 2669], fig. 1

iTtTbiriSoTroTroSiD
3532102139
  +----- Anodonta **     
--+                   
  |  +-- Pseudanodonta **
  +--+                
     +-- Alasmidontina

Soroka, 2010 [Publ 3404], fig. 5b

iTtTbiriSoTroTroSiD
385252144
  +-------- Anodonta_anatina **
  |                         
--+  +----- Cristariina     
  |  |                      
  +--+----- Unionina        
     |                      
     |  +-- Anodonta_cygnea ** 
     +--+                   
        +-- Pseudanodonta **

Zieritz et al., 2012 [Publ 3673], fig. 4

iTtTbiriSoTroTroSiD
330251138
  +-- Pseudanodonta **   
  |                   
--+-- Anodonta_anatina **
  |                   
  +-- Anodonta_cygnea **

Soroka, 2010 [Publ 3404], fig. 5a

iTtTbiriSoTroTroSiD
353252141
  +-------- Anodonta_anatina **
--+                         
  |  +----- Cristariina     
  +--+                      
     |  +-- Anodonta_cygnea ** 
     +--+                   
        +-- Pseudanodonta **

Mezhzherin et al., 2014 [Publ 3705], fig. 4

iTtTbiriSoTroTroSiD
331211139
  +-------- Cristariina     
--+                         
  |  +----- Anodonta_anatina **
  +--+                      
     |  +-- Anodonta_cygnea ** 
     +--+                   
        +-- Pseudanodonta **

Källersjö et al., 2005 [Publ 2684], fig. 1A

iTtTbiriSoTroTroSiD
330240038
  +-- Anodonta_anatina **
  |                   
--+-- Pseudanodonta **   
  |                   
  +-- Anodonta_cygnea **

Nagel et al., 1998 [Publ 1946], fig. 1

iTtTbiriSoTroTroSiD
331260039
  +-- Pseudanodonta **
--+                
  +-- Anodonta **

Källersjö et al., 2005 [Publ 2684], fig. 1B-F

iTtTbiriSoTroTroSiD
331240039
  +----- Anodonta_anatina **
--+                      
  |  +-- Pseudanodonta **   
  +--+                   
     +-- Anodonta_cygnea **

Soroka, 2010 [Publ 3404], fig. 3a

iTtTbiriSoTroTroSiD
331262139
  +----- Anodonta_anatina **
--+                      
  |  +-- Anodonta_cygnea ** 
  +--+                   
     +-- Pseudanodonta **

Bolotov et al., 2020 [Publ 4084], fig. 1

iTtTbiriSoTroTroSiD
3312339139
  +-------- Cristariina     
--+                         
  |  +----- Anodonta_anatina **
  +--+                      
     |  +-- Pseudanodonta **   
     +--+                   
        +-- Anodonta_cygnea **

Reis et al., 2013 [Publ 3580], fig. 5

iTtTbiriSoTroTroSiD
3302120038
  +-- Anodonta_anatina **
  |                   
--+-- Anodonta_cygnea ** 
  |                   
  +-- Pseudanodonta **

Wu et al., 2021 [Publ 4366], fig. 6

iTtTbiriSoTroTroSiD
3642887240
  +-------------- Cristariina     
  |                               
--+  +----------- Anodonta_cygnea ** 
  |  |                            
  +--+        +-- Lasmigona       
     |  +-----+                   
     |  |     +-- Anodonta_anatina **
     +--+                         
        |  +----- Pseudanodonta **   
        +--+                      
           |  +-- Pyganodon       
           +--+                   
              +-- Strophitus

Reis et al., 2013 [Publ 3580], fig. 4

iTtTbiriSoTroTroSiD
3312110037
  +----- Anodonta_cygnea ** 
--+                      
  |  +-- Anodonta_anatina **
  +--+                   
     +-- Pseudanodonta **

Froufe et al., 2017 [Publ 3802], fig. 2

iTtTbiriSoTroTroSiD
331111139
  +-- Cristariina
--+              
  +-- Anodonta **

Pfarrer et al., 2022 [Publ 4305], fig. 2-5

iTtTbiriSoTroTroSiD
331192139
  +-- Cristariina
--+              
  +-- Anodonta **

Wu et al., 2023 [Publ 4339], fig. 5

iTtTbiriSoTroTroSiD
3312166239
  +----------- Cristariina     
--+                            
  |  +-------- Alasmidontina   
  +--+                         
     |  +----- Anodonta_anatina **
     +--+                      
        |  +-- Pseudanodonta **   
        +--+                   
           +-- Anodonta_cygnea **

Gomes-dos-Santos et al., 2019 [Publ 4042], fig. 8

iTtTbiriSoTroTroSiD
331133139
  +-- Anodonta **   
--+              
  +-- Cristariina

Plazzi et al., 2011 [Publ 3676], fig. 5

iTtTbiriSoTroTroSiD
317102152252
  +----------------- Cristariina  
  |                               
  |  +-------------- Lanceolaria  
  |  |                            
--+  +-------------- Anodonta **     
  |  |                            
  |  +-------------- Unionina     
  |  |                            
  |  |           +-- Alasmidontina
  +--+-----------+                
     |           +-- Pseudanodonta **
     |                            
     |           +-- Alathyria    
     |           |                
     |  +--------+-- Pronodularia 
     |  |        |                
     |  |        +-- Sinohyriopsis
     +--+                         
        |  +-------- Quadrulina   
        |  |                      
        +--+-------- Parvaspina   
           |                      
           |  +----- Lampsilis    
           +--+                   
              |  +-- Epioblasma   
              +--+                
                 +-- Venustaconcha

Lopes-Lima et al., 2016 [Publ 3757], fig. 1

iTtTbiriSoTroTroSiD
331200039
  +-- Pseudanodonta **
--+                
  +-- Anodonta **

Prié & Puillandre, 2014 [Publ 3529], fig. 2

iTtTbiriSoTroTroSiD
3312140039
  +----- Anodonta_anatina **
--+                      
  |  +-- Pseudanodonta **   
  +--+                   
     +-- Anodonta_cygnea **

Wu et al., 2021 [Publ 4366], fig. 2_I

iTtTbiriSoTroTroSiD
3532887239
  +-------------- Cristariina  
--+                            
  |  +----------- Lasmigona    
  +--+                         
     |  +-------- Anodonta **     
     +--+                      
        |  +----- Pseudanodonta **
        +--+                   
           |  +-- Pyganodon    
           +--+                
              +-- Strophitus

Perkins et al., 2025 [Publ 4447], fig. 2_mtDNA-ITS

iTtTbiriSoTroTroSiD
3363723333132
  +----- Anodonta **     
--+                   
  |  +-- Pseudanodonta **
  +--+                
     +-- Alasmidontina

Soroka, 2010 [Publ 3404], fig. 2

iTtTbiriSoTroTroSiD
331241137
  +----- Anodonta_cygnea ** 
--+                      
  |  +-- Anodonta_anatina **
  +--+                   
     +-- Pseudanodonta **

Soroka, 2010 [Publ 3404], fig. 3b

iTtTbiriSoTroTroSiD
395262144
  +----- Anodonta_anatina **
  |                      
  +----- Anodonta_cygnea ** 
--+                      
  +----- Pseudanodonta **   
  |                      
  |  +-- Cristariina     
  +--+                   
     +-- Unionina

Froufe et al., 2017 [Publ 3802], fig. 7

iTtTbiriSoTroTroSiD
331171139
  +-- Cristariina
--+              
  +-- Anodonta **

Lopes-Lima et al., 2017 [Publ 3734], fig. 1

iTtTbiriSoTroTroSiD
3312140039
  +----- Anodonta_anatina **
--+                      
  |  +-- Pseudanodonta **   
  +--+                   
     +-- Anodonta_cygnea **

Hoeh, 1990 [Publ 2717], fig. 3

iTtTbiriSoTroTroSiD
215021313146
  +----- Pyganodon_fragilis         
  |                                 
  +----- Utterbackia_imbecillis     
  |                                 
  +----- Utterbackia_peggyae        
  |                                 
  +----- Anodonta **                   
  |                                 
  +----- Brachyanodon **               
  |                                 
  +----- Utterbackiana_couperiana__1
  |                                 
--+----- Utterbackiana_couperiana__2
  |                                 
  +----- Utterbackiana_suborbiculata
  |                                 
  |  +-- Lasmigona                  
  +--+                              
  |  +-- Utterbackiana_implicata    
  |                                 
  |  +-- Pterosyna                  
  |  |                              
  |  +-- Platynaias                 
  |  |                              
  |  +-- Pyganodon_cataracta        
  +--+                              
     +-- Pyganodon_gibbosa          
     |                              
     +-- Pyganodon_grandis          
     |                              
     +-- Pyganodon_lacustris

Lopes-Lima et al., 2024 [Publ 4376], fig. 4-5_MtG

iTtTbiriSoTroTroSiD
22012000--
1

Whelan et al., 2023 [Publ 4287], fig. 4

iTtTbiriSoTroTroSiD
220125251--
  +-- Anodonta **     
--+                
  +-- Alasmidontina

Wu et al., 2021 [Publ 4366], fig. 4

iTtTbiriSoTroTroSiD
28016710234
  +-------- Cristariina     
--+                         
  |  +----- Anodonta_cygnea ** 
  +--+                      
     |  +-- Anodonta_anatina **
     +--+                   
        +-- Alasmidontina

Zanatta et al., 2007 [Publ 2698], fig. 1A

iTtTbiriSoTroTroSiD
213021414140
           +-- Platynaias   
           |                
  +--------+-- Lasmigona    
  |        |                
  |        +-- Pterosyna    
--+                         
  |  +-------- Anodonta **     
  |  |                      
  +--+     +-- Brachyanodon ** 
     |  +--+                
     |  |  +-- Utterbackiana
     +--+                   
        |  +-- Utterbackia  
        +--+                
           +-- Pyganodon

Campbell et al., 2005 [Publ 2660], fig. 2

iTtTbiriSoTroTroSiD
220210041--
  +-------- Pyganodon       
  |                         
  |     +-- Lasmigona       
--+-----+                   
  |     +-- Pseudodontoideus
  |                         
  |  +----- Alasminota      
  +--+                      
     |  +-- Anodonta **        
     +--+                   
        +-- Brachyanodon **

Klishko et al., 2019 [Publ 3886], fig. 9

iTtTbiriSoTroTroSiD
2201900--
1

Sano et al., 2020 [Publ 4138], fig. 3

iTtTbiriSoTroTroSiD
220166172--
  +-- Anodonta **     
--+                
  +-- Alasmidontina

Jadhav & Jamkhedkar, 2009 [Publ 3697], fig. 11

iTtTbiriSoTroTroSiD
2801234141
  +----------- Anodonta_cygnea ** 
  |                            
--+  +-------- Unio_pictorum__2
  |  |                         
  +--+     +-- Unio_pictorum__1
     |  +--+                   
     |  |  +-- Unio_caffer     
     +--+                      
        |  +-- Anodonta **        
        +--+                   
           +-- Alasmidontina

Wu et al., 2021 [Publ 4366], fig. 3-ML

iTtTbiriSoTroTroSiD
29017211233
  +-------- Cristariina     
--+                         
  |  +----- Anodonta_cygnea ** 
  +--+                      
     |  +-- Anodonta_anatina **
     +--+                   
        +-- Alasmidontina

Chen et al., 2025 [Publ 4487], fig. 3

iTtTbiriSoTroTroSiD
22023982--
  +----- Alasmidontina
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Pseudanodonta **

Nagel et al., 1998 [Publ 1946], fig. 9

iTtTbiriSoTroTroSiD
2202400--
  +-- Anodonta **     
--+                
  +-- Pseudanodonta **

Wu et al., 2022 [Publ 4238], fig. 3

iTtTbiriSoTroTroSiD
22017282--
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Soroka & Burzynski, 2017 [Publ 3837], fig. 1

iTtTbiriSoTroTroSiD
22012182--
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Froufe et al., 2017 [Publ 3802], fig. 6

iTtTbiriSoTroTroSiD
220110101--
  +-- Anodonta **   
--+              
  +-- Cristariina

Huang et al., 2019 [Publ 3843], fig. 3

iTtTbiriSoTroTroSiD
2601548234
  +-------- Cristariina     
--+                         
  |  +----- Anodonta_cygnea ** 
  +--+                      
     |  +-- Anodonta_anatina **
     +--+                   
        +-- Alasmidontina

Dai et al., 2025 [Publ 4440], fig. 2

iTtTbiriSoTroTroSiD
22024382--
  +----- Alasmidontina
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Pseudanodonta **

Zanatta et al., 2007 [Publ 2698], fig. 1B

iTtTbiriSoTroTroSiD
26021414139
           +-- Lasmigona    
           |                
  +--------+-- Platynaias   
  |        |                
  |        +-- Pterosyna    
--+                         
  |        +-- Utterbackia  
  |  +-----+                
  |  |     +-- Pyganodon    
  +--+                      
     |  +----- Anodonta **     
     +--+                   
        |  +-- Brachyanodon ** 
        +--+                
           +-- Utterbackiana

Soroka, 2010 [Publ 3404], fig. 8b

iTtTbiriSoTroTroSiD
2201411--
1

Fukata & Iigo, 2020 [Publ 4106], fig. 1

iTtTbiriSoTroTroSiD
22015282--
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Dyduch-Falniowska & Koziol, 1989 [Publ 3814], fig. 5_7

iTtTbiriSoTroTroSiD
2202300--
  +-- Pseudanodonta **
--+                
  +-- Anodonta **

Xiang et al., 2025 [Publ 4433], fig. 7

iTtTbiriSoTroTroSiD
22011171--
  +-- Anodonta **   
--+              
  +-- Cristariina

Dai et al., 2025 [Publ 4440], fig. 1

iTtTbiriSoTroTroSiD
22024382--
  +----- Alasmidontina
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Pseudanodonta **

Soroka, 2010 [Publ 3404], fig. 8a

iTtTbiriSoTroTroSiD
2201411--
  +-- Cristariina
--+              
  +-- Anodonta **

Hoeh, 1990 [Publ 2717], fig. 2

iTtTbiriSoTroTroSiD
25021313139
           +-- Pterosyna    
           |                
  +--------+-- Platynaias   
  |        |                
  |        +-- Lasmigona    
--+                         
  |        +-- Utterbackia  
  |  +-----+                
  |  |     +-- Pyganodon    
  +--+                      
     |  +----- Anodonta **     
     +--+                   
        |  +-- Brachyanodon ** 
        +--+                
           +-- Utterbackiana

Aunins et al., 2018 [Publ 3863], fig. 1

iTtTbiriSoTroTroSiD
22013892--
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Bogatov et al., 2018 [Publ 3859], fig. 2

iTtTbiriSoTroTroSiD
2202400--
  +-- Anodonta **     
--+                
  +-- Pseudanodonta **

Wu et al., 2021 [Publ 4366], fig. 3-BI

iTtTbiriSoTroTroSiD
29017211233
  +-------- Cristariina     
--+                         
  |  +----- Anodonta_cygnea ** 
  +--+                      
     |  +-- Anodonta_anatina **
     +--+                   
        +-- Alasmidontina

Wu et al., 2026 [Publ 4811], fig. 3

iTtTbiriSoTroTroSiD
22011500--
1

Sano et al., 2020 [Publ 4138], fig. 2

iTtTbiriSoTroTroSiD
220166172--
  +-- Anodonta **     
--+                
  +-- Alasmidontina

Wu et al., 2022 [Publ 4238], fig. 2d

iTtTbiriSoTroTroSiD
22013782--
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Chen et al., 2023 [Publ 4302], fig. 1

iTtTbiriSoTroTroSiD
22023982--
  +----- Alasmidontina
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Pseudanodonta **

Chen et al., 2020 [Publ 4170], fig. 1

iTtTbiriSoTroTroSiD
22013292--
  +-------- Alasmidontina
--+                      
  |  +----- Sinanodonta  
  +--+                   
     |  +-- Cristaria    
     +--+                
        +-- Anodonta **

Wu et al., 2018 [Publ 3836], fig. 2

iTtTbiriSoTroTroSiD
27015111238
  +-------- Cristariina     
--+                         
  |  +----- Anodonta_cygnea ** 
  +--+                      
     |  +-- Anodonta_anatina **
     +--+                   
        +-- Alasmidontina

Chen et al., 2025 [Publ 4487], fig. 2

iTtTbiriSoTroTroSiD
22023982--
  +-------- Cristariina  
--+                      
  |  +----- Alasmidontina
  +--+                   
     |  +-- Anodonta **     
     +--+                
        +-- Pseudanodonta **

Chen et al., 2023 [Publ 4302], fig. 2

iTtTbiriSoTroTroSiD
22023982--
  +-------- Cristariina  
--+                      
  |  +----- Alasmidontina
  +--+                   
     |  +-- Anodonta **     
     +--+                
        +-- Pseudanodonta **

Dai et al., 2025 [Publ 4440], fig. 3

iTtTbiriSoTroTroSiD
22018682--
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Bogatov et al., 2018 [Publ 3859], fig. 1

iTtTbiriSoTroTroSiD
2202400--
  +-- Anodonta **     
--+                
  +-- Pseudanodonta **

Gustafson & Iwamoto, 2005 [Publ 2741], fig. 2

iTtTbiriSoTroTroSiD
2201311--
  +-- Cristariina 
--+               
  +-- Brachyanodon **

Lopes-Lima et al., 2024 [Publ 4376], fig. 4-5_AHE

iTtTbiriSoTroTroSiD
22012000--
1

Hoeh, 1990 [Publ 2717], fig. 4

iTtTbiriSoTroTroSiD
25021313139
           +-- Pterosyna    
           |                
  +--------+-- Platynaias   
  |        |                
  |        +-- Lasmigona    
--+                         
  |        +-- Utterbackia  
  |  +-----+                
  |  |     +-- Pyganodon    
  +--+                      
     |  +----- Anodonta **     
     +--+                   
        |  +-- Brachyanodon ** 
        +--+                
           +-- Utterbackiana

Chen et al., 2025 [Publ 4448], fig. 3

iTtTbiriSoTroTroSiD
22018492--
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Sano et al., 2020 [Publ 4138], fig. 4

iTtTbiriSoTroTroSiD
220166172--
  +-- Anodonta **     
--+                
  +-- Alasmidontina

Chase et al., 2018 [Publ 3847], fig. 4

iTtTbiriSoTroTroSiD
220137122--
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Campbell et al., 2005 [Publ 2660], fig. 1

iTtTbiriSoTroTroSiD
26021004141
  +----------- Brachyanodon **    
  |                            
--+        +-- Anodonta **        
  |  +-----+                   
  |  |     +-- Alasminota      
  +--+                         
     |  +----- Pyganodon       
     +--+                      
        |  +-- Lasmigona       
        +--+                   
           +-- Pseudodontoideus

Huang et al., 2019 [Publ 3843], fig. 3-M

iTtTbiriSoTroTroSiD
11012232--
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Gomes-dos-Santos et al., 2019 [Publ 3943], fig. 1

iTtTbiriSoTroTroSiD
1101900--
-- Anodonta_anatina **

Bogan et al., 2023 [Publ 4295], fig. 3

iTtTbiriSoTroTroSiD
110130271--
  +-- Anodonta **   
--+              
  +-- Cristariina

Rosenberg et al., 1997 [Publ 2720], fig. 2A

iTtTbiriSoTroTroSiD
11012131--
-- Anodonta_anatina **

Soroka, 2010 [Publ 3404], fig. 3a-M

iTtTbiriSoTroTroSiD
1101521--
-- Anodonta_anatina **

Bolotov et al., 2016 [Publ 3743], fig. 2

iTtTbiriSoTroTroSiD
11012900--
-- Anodonta_anatina **

Sonowal et al., 2021 [Publ 4221], fig. 4

iTtTbiriSoTroTroSiD
11011500--
-- Anodonta_cygnea **

Froufe et al., 2019 [Publ 3936], fig. 3

iTtTbiriSoTroTroSiD
11014382--
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Taylor et al., 2007 [Publ 2696], fig. 1-3

iTtTbiriSoTroTroSiD
1101300--
-- Anodonta_cygnea **

Walker et al., 2006 [Publ 2666], fig. 3

iTtTbiriSoTroTroSiD
11011421--
  +-- Anodonta **     
--+                
  +-- Alasmidontina

Wen et al., 2017 [Publ 3795], fig. 3

iTtTbiriSoTroTroSiD
11011521--
  +-- Anodonta **   
--+              
  +-- Cristariina

Feng et al., 2016 [Publ 3800], fig. 2

iTtTbiriSoTroTroSiD
11011352--
  +-- Anodonta **     
--+                
  +-- Alasmidontina

Chase et al., 2018 [Publ 3861], fig. 1-M

iTtTbiriSoTroTroSiD
11011021--
  +-- Anodonta **     
--+                
  +-- Alasmidontina

Fonseca et al., 2016 [Publ 3775], fig. 1

iTtTbiriSoTroTroSiD
11013082--
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Cyr et al., 2007 [Publ 2745], fig. 2B

iTtTbiriSoTroTroSiD
1101441--
  +-- Brachyanodon ** 
--+                
  +-- Alasmidontina

Zieritz et al., 2024 [Publ 4381], fig. 2

iTtTbiriSoTroTroSiD
11014400--
-- Anodonta_cygnea **

Mock et al., 2004 [Publ 2686], fig. 4

iTtTbiriSoTroTroSiD
1101111--
  +-- Anodonta **   
--+              
  +-- Cristariina

Froufe et al., 2019 [Publ 3936], fig. 6

iTtTbiriSoTroTroSiD
11014382--
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Wu et al., 2018 [Publ 4143], fig. 2

iTtTbiriSoTroTroSiD
11012162--
  +----------- Sinanodonta  
--+                         
  |  +-------- Anemina      
  +--+                      
     |  +----- Cristaria    
     +--+                   
        |  +-- Anodonta **     
        +--+                
           +-- Alasmidontina

Hoeh et al., 2001 [Publ 2652], fig. 14f2

iTtTbiriSoTroTroSiD
11013021--
  +----- Pyganodon 
--+                
  |  +-- Anodonta **  
  +--+             
     +-- Strophitus

Araujo et al., 2018 [Publ 3779], fig. 1

iTtTbiriSoTroTroSiD
11011721--
  +-- Anodonta **     
--+                
  +-- Alasmidontina

Bogan & Hoeh, 2000 [Publ 2650], fig. 1

iTtTbiriSoTroTroSiD
11013321--
  +-- Anodonta **     
--+                
  +-- Alasmidontina

Burzynski et al., 2017 [Publ 3774], fig. 1-top

iTtTbiriSoTroTroSiD
11011221--
  +-- Anodonta **     
--+                
  +-- Alasmidontina

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iTtTbiriSoTroTroSiD
11015632--
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Burzynski et al., 2017 [Publ 3774], fig. 1-bottom

iTtTbiriSoTroTroSiD
11011221--
  +-- Anodonta **     
--+                
  +-- Alasmidontina

Whelan et al., 2023 [Publ 4287], fig. 3

iTtTbiriSoTroTroSiD
110127271--
  +-- Anodonta **     
--+                
  +-- Alasmidontina

Wang et al., 2018 [Publ 3865], fig. 2

iTtTbiriSoTroTroSiD
11013062--
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Hoeh et al., 2001 [Publ 2652], fig. 14f4

iTtTbiriSoTroTroSiD
11013021--
  +-- Anodonta **     
--+                
  +-- Alasmidontina

Froufe et al., 2015 [Publ 3664], fig. 1-M

iTtTbiriSoTroTroSiD
1101721--
  +-- Anodonta **     
--+                
  +-- Alasmidontina

Wu et al., 2019 [Publ 3864], fig. 3

iTtTbiriSoTroTroSiD
11014582--
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Araujo et al., 2017 [Publ 3801], fig. 2

iTtTbiriSoTroTroSiD
11011300--
-- Anodonta_anatina **

Dai et al., 2023 [Publ 4285], fig. 5

iTtTbiriSoTroTroSiD
11015262--
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Lopes-Lima et al., 2017 [Publ 3792], fig. 2

iTtTbiriSoTroTroSiD
11013182--
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Gomes-dos-Santos et al., 2019 [Publ 3943], fig. 1-M

iTtTbiriSoTroTroSiD
1101500--
-- Anodonta_anatina **

Mock et al., 2004 [Publ 2686], fig. 3

iTtTbiriSoTroTroSiD
1101111--
  +-- Anodonta **   
--+              
  +-- Cristariina

Wu et al., 2023 [Publ 4296], fig. 4

iTtTbiriSoTroTroSiD
11012231--
  +-- Pseudanodonta **
--+                
  +-- Alasmidontina

Huff et al., 2004 [Publ 2697], fig. 5-6

iTtTbiriSoTroTroSiD
11011100--
-- Anodonta **

Bolotov et al., 2016 [Publ 3743], fig. 4

iTtTbiriSoTroTroSiD
11012900--
-- Anodonta_anatina **

Wu et al., 2020 [Publ 4121], fig. 3

iTtTbiriSoTroTroSiD
11014882--
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Hoeh et al., 2002 [Publ 2653], fig. 2

iTtTbiriSoTroTroSiD
11013021--
  +-- Anodonta **     
--+                
  +-- Alasmidontina

Araujo et al., 2009 [Publ 2589], fig. 7B

iTtTbiriSoTroTroSiD
11011000--
-- Anodonta_cygnea **

Burzynski & Soroka, 2018 [Publ 3862], fig. 5a

iTtTbiriSoTroTroSiD
11011932--
  +-- Cristariina  
  |                
--+-- Anodonta **     
  |                
  +-- Alasmidontina

Guerra et al., 2019 [Publ 4168], fig. 2-M

iTtTbiriSoTroTroSiD
11012132--
  +----- Alasmidontina
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Cristariina

Hoeh et al., 1998 [Publ 2695], fig. 4

iTtTbiriSoTroTroSiD
1101400--
-- Anodonta_cygnea **

Sonowal et al., 2021 [Publ 4221], fig. 3a

iTtTbiriSoTroTroSiD
11011400--
-- Anodonta_cygnea **

Lopes-Lima et al., 2019 [Publ 3901], fig. 2

iTtTbiriSoTroTroSiD
11012711--
  +-- Brachyanodon ** 
--+                
  +-- Alasmidontina

Soroka, 2010 [Publ 3404], fig. 2-M

iTtTbiriSoTroTroSiD
1101411--
-- Anodonta_anatina **

Klymus et al., 2025 [Publ 4814], fig. 3B

iTtTbiriSoTroTroSiD
11013532--
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Huff et al., 2004 [Publ 2697], fig. 2

iTtTbiriSoTroTroSiD
11011100--
-- Anodonta **

Lydeard et al., 2000 [Publ 2654], fig. 2b

iTtTbiriSoTroTroSiD
11011300--
-- Anodonta_cygnea **

Soroka, 2008 [Publ 3695], fig. 1

iTtTbiriSoTroTroSiD
1101322--
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Rosenberg et al., 1997 [Publ 2720], fig. 2B

iTtTbiriSoTroTroSiD
11012131--
-- Anodonta **

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

iTtTbiriSoTroTroSiD
11013852--
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Roe & Hoeh, 2003 [Publ 2655], fig. 4f3

iTtTbiriSoTroTroSiD
11013021--
  +-- Anodonta **     
--+                
  +-- Alasmidontina

Zieritz et al., 2021 [Publ 4231], fig. 3-M_4

iTtTbiriSoTroTroSiD
11012232--
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Lydeard et al., 2000 [Publ 2654], fig. 2a

iTtTbiriSoTroTroSiD
11011300--
-- Anodonta_cygnea **

Wu et al., 2022 [Publ 4238], fig. 2e

iTtTbiriSoTroTroSiD
11011732--
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Dai et al., 2024 [Publ 4358], fig. 4

iTtTbiriSoTroTroSiD
11015262--
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Wen et al., 2017 [Publ 3795], fig. 3-M

iTtTbiriSoTroTroSiD
1101521--
  +-- Anodonta **     
--+                
  +-- Alasmidontina

Burzynski & Soroka, 2018 [Publ 3862], fig. 3

iTtTbiriSoTroTroSiD
1101332--
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A

iTtTbiriSoTroTroSiD
11012951--
  +-- Anodonta **     
--+                
  +-- Alasmidontina

Guerra et al., 2019 [Publ 4168], fig. 1-M

iTtTbiriSoTroTroSiD
11012132--
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Hoeh et al., 1998 [Publ 2695], fig. 3

iTtTbiriSoTroTroSiD
1101400--
-- Anodonta_cygnea **

Soroka, 2010 [Publ 3404], fig. 5a-M

iTtTbiriSoTroTroSiD
1101111--
  +-- Anodonta **   
--+              
  +-- Cristariina

Burzynski & Soroka, 2018 [Publ 3862], fig. 5b

iTtTbiriSoTroTroSiD
1101932--
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Lopes-Lima et al., 2017 [Publ 3792], fig. 2-M

iTtTbiriSoTroTroSiD
1101921--
  +-- Anodonta **     
--+                
  +-- Alasmidontina

Lydeard et al., 1996 [Publ 2656], fig. 4

iTtTbiriSoTroTroSiD
11012751--
     +----- Utterbackia  
  +--+                   
  |  |  +-- Utterbackiana
  |  +--+                
--+     +-- Anodonta **     
  |                      
  |  +----- Strophitus   
  +--+                   
     |  +-- Lasmigona    
     +--+                
        +-- Alasmidonta

Lydeard et al., 2000 [Publ 2654], fig. 2c

iTtTbiriSoTroTroSiD
11011300--
-- Anodonta_cygnea **

Roe & Hoeh, 2003 [Publ 2655], fig. 4f2

iTtTbiriSoTroTroSiD
11013021--
  +----- Pyganodon 
--+                
  |  +-- Anodonta **  
  +--+             
     +-- Strophitus

Chase et al., 2018 [Publ 3861], fig. 1

iTtTbiriSoTroTroSiD
11011021--
  +-- Anodonta **     
--+                
  +-- Alasmidontina

Soroka & Burzynski, 2018 [Publ 3762], fig. 3

iTtTbiriSoTroTroSiD
1101621--
  +-- Anodonta **     
--+                
  +-- Alasmidontina

Cha et al., 2018 [Publ 3916], fig. 1

iTtTbiriSoTroTroSiD
11013782--
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

iTtTbiriSoTroTroSiD
11014782--
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Wu et al., 2020 [Publ 4147], fig. 3

iTtTbiriSoTroTroSiD
11012962--
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Wang et al., 2018 [Publ 3846], fig. 2

iTtTbiriSoTroTroSiD
11013062--
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Fonseca et al., 2016 [Publ 3775], fig. 1-M

iTtTbiriSoTroTroSiD
1101821--
  +-- Anodonta **     
--+                
  +-- Alasmidontina

Dai et al., 2024 [Publ 4386], fig. 5-ML

iTtTbiriSoTroTroSiD
11012111--
-- Anodonta_anatina **

Soroka, 2010 [Publ 3404], fig. 3b-M

iTtTbiriSoTroTroSiD
1101521--
-- Anodonta_anatina **

Araujo et al., 2009 [Publ 2589], fig. 7C

iTtTbiriSoTroTroSiD
1101900--
-- Anodonta_cygnea **

Huang et al., 2018 [Publ 3828], fig. 6

iTtTbiriSoTroTroSiD
11014182--
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Lopes-Lima et al., 2019 [Publ 3901], fig. 4

iTtTbiriSoTroTroSiD
11012511--
  +-- Brachyanodon ** 
--+                
  +-- Alasmidontina

Walker et al., 2006 [Publ 2666], fig. 2

iTtTbiriSoTroTroSiD
11011421--
  +-- Anodonta **     
--+                
  +-- Alasmidontina

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B

iTtTbiriSoTroTroSiD
11012951--
  +-- Anodonta **     
--+                
  +-- Alasmidontina

Huff et al., 2004 [Publ 2697], fig. 4

iTtTbiriSoTroTroSiD
11011000--
-- Anodonta **

Guerra et al., 2017 [Publ 3772], fig. 2b

iTtTbiriSoTroTroSiD
11012472--
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Dai et al., 2024 [Publ 4386], fig. 3

iTtTbiriSoTroTroSiD
11011300--
-- Anodonta_anatina **

Krebs et al., 2003 [Publ 2691], fig. 3

iTtTbiriSoTroTroSiD
11013291--
  +----------- Utterbackiana
  |                         
  +----------- Platynaias   
  |                         
  +----------- Alasmidonta  
--+                         
  |        +-- Pterosyna    
  +--------+                
  |        +-- Lasmigona    
  |                         
  |  +-------- Strophitus   
  +--+                      
     |  +----- Utterbackia  
     +--+                   
        |  +-- Pyganodon    
        +--+                
           +-- Anodonta **

Cyr et al., 2007 [Publ 2745], fig. 2C

iTtTbiriSoTroTroSiD
1101441--
  +-- Pseudanodonta **
--+                
  +-- Alasmidontina

Froufe et al., 2019 [Publ 3936], fig. 3-M

iTtTbiriSoTroTroSiD
11011832--
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Guerra et al., 2017 [Publ 3772], fig. 2a

iTtTbiriSoTroTroSiD
11012472--
  +-------- Sinanodonta  
--+                      
  |  +----- Cristaria    
  +--+                   
     |  +-- Anodonta **     
     +--+                
        +-- Alasmidontina

Dai et al., 2024 [Publ 4386], fig. 5-BI

iTtTbiriSoTroTroSiD
11012111--
-- Anodonta_anatina **

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iTtTbiriSoTroTroSiD
11015632--
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

iTtTbiriSoTroTroSiD
11013642--
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Araujo et al., 2009 [Publ 2589], fig. 7A

iTtTbiriSoTroTroSiD
11011000--
-- Anodonta_cygnea **

Guerra et al., 2019 [Publ 4168], fig. 2

iTtTbiriSoTroTroSiD
11014062--
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Wu et al., 2024 [Publ 4388], fig. 6

iTtTbiriSoTroTroSiD
11015262--
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Lopes-Lima et al., 2019 [Publ 3901], fig. 3

iTtTbiriSoTroTroSiD
11012811--
  +-- Brachyanodon ** 
--+                
  +-- Alasmidontina

Huff et al., 2004 [Publ 2697], fig. 3

iTtTbiriSoTroTroSiD
1101700--
-- Anodonta **

Bolotov et al., 2018 [Publ 3826], fig. 2

iTtTbiriSoTroTroSiD
11018192--
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Lydeard et al., 2000 [Publ 2654], fig. 2d

iTtTbiriSoTroTroSiD
11011300--
-- Anodonta_cygnea **

Soroka, 2010 [Publ 3404], fig. 4-M

iTtTbiriSoTroTroSiD
1101111--
  +-- Anodonta **   
--+              
  +-- Cristariina

Wu et al., 2016 [Publ 3711], fig. 1

iTtTbiriSoTroTroSiD
11012062--
  +----------- Sinanodonta  
--+                         
  |  +-------- Anemina      
  +--+                      
     |  +----- Cristaria    
     +--+                   
        |  +-- Anodonta **     
        +--+                
           +-- Alasmidontina

Sonowal et al., 2021 [Publ 4221], fig. 3b

iTtTbiriSoTroTroSiD
11011400--
-- Anodonta_cygnea **

Campbell & Lydeard, 2026 [Publ 4810], fig. 4

iTtTbiriSoTroTroSiD
11014632--
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Liu et al., 2023 [Publ 4310], fig. 5

iTtTbiriSoTroTroSiD
11012331--
  +-------- Lasmigona    
--+                      
  |  +----- Pseudanodonta **
  +--+                   
     |  +-- Pyganodon    
     +--+                
        +-- Strophitus

Froufe et al., 2019 [Publ 3936], fig. 4

iTtTbiriSoTroTroSiD
11011832--
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Walker et al., 2006 [Publ 2666], fig. 4

iTtTbiriSoTroTroSiD
11011421--
  +-- Anodonta **     
--+                
  +-- Alasmidontina

Chase et al., 2018 [Publ 3847], fig. 5

iTtTbiriSoTroTroSiD
1101911--
  +-- Anodonta **     
--+                
  +-- Alasmidontina

Huang et al., 2018 [Publ 3828], fig. 6-M

iTtTbiriSoTroTroSiD
11011632--
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Soroka, 2008 [Publ 3695], fig. 1-M

iTtTbiriSoTroTroSiD
1101322--
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Klymus et al., 2025 [Publ 4814], fig. 3A

iTtTbiriSoTroTroSiD
11013532--
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Froufe et al., 2015 [Publ 3664], fig. 1

iTtTbiriSoTroTroSiD
11011121--
  +-- Anodonta **     
--+                
  +-- Alasmidontina

Soroka, 2010 [Publ 3404], fig. 5b-M

iTtTbiriSoTroTroSiD
1101111--
  +-- Anodonta **   
--+              
  +-- Cristariina

Liu et al., 2023 [Publ 4310], fig. 4

iTtTbiriSoTroTroSiD
11012331--
  +-- Pseudanodonta **
--+                
  +-- Alasmidontina

Guerra et al., 2019 [Publ 4168], fig. 1

iTtTbiriSoTroTroSiD
11014062--
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Soroka & Burzynski, 2018 [Publ 3762], fig. 3-M

iTtTbiriSoTroTroSiD
1101621--
  +-- Anodonta **     
--+                
  +-- Alasmidontina
 
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