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Page last updated
Thu 3 September 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Search the database pages:

Bivalvia | Unionoida | Unionoidea | Unionidae | Unioninae | Anodontini

subtribe ANODONTINA Rafinesque, 1820: cladomics
     3 genera

Cladomics Summary

number of species = 11 (24 total)
number of subtaxa (rank: genus) = 3
analyzed in 216 cladograms in 117 publications
maximum number of terminals in an analysis = 7 (mean = 1.96, median = 1)
maximum number of subtaxa (genus) = 2 (mean = 1.28, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 103
maximum number of included terminals = 7, (mean = 3.02 median = 3)
maximum number of subtaxa (genus) = 2 (mean = 1.59, median = 2)
proportion of cladograms that support basic monophyly = 0.6

Rigorous Monophyly

number of cladograms that test rigorous monophyly = 45
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (genus) taxon and at least one rigorous outgroup (family).
maximum number of included terminals = 7 (mean = 3.69, median = 4)
maximum number of subtaxa (genus) = 2 (mean = 2, median = 2)
proportion of cladograms that support rigorous monophyly = 0.4

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.


PCoA Ordination


NJ Tree

Whelan et al., 2023 [Publ 4287], fig. 2

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
437382373720
  +----------- Cristariina         
  |                                
--+  +-------- Anodonta_cygnea **     
  |  |                             
  +--+     +-- Anodonta_nuttalliana **
     |  +--+                       
     |  |  +-- Pseudanodonta **       
     +--+                          
        |  +-- Anodonta_anatina **    
        +--+                       
           +-- Alasmidontina

Galal-Khallaf et al., 2024 [Publ 4459], fig. 6

iTtTbiriSoTroTroSiD
7971216239
  +-------------- Cristariina            
  |                                      
  |           +-- Anodonta_exulcerata **    
--+  +--------+                          
  |  |        +-- Anodonta_cygnea **        
  |  |                                   
  +--+        +-- Anodonta_californiensis **
     |  +-----+                          
     |  |     +-- Anodonta_nuttalliana **   
     |  |                                
     +--+     +-- Anodonta_kennerlyi **     
        |  +--+                          
        |  |  +-- Anodonta_oregonensis **   
        +--+                             
           |  +-- Anodonta_anatina **       
           +--+                          
              +-- Alasmidontina

Chong et al., 2008 [Publ 2668], fig. 3

iTtTbiriSoTroTroSiD
7181021110246
  +----------------- Anodonta_cygnea **        
  |                                         
  +----------------- Pyganodon              
  |                                         
  +----------------- Alasminota             
  |                                         
  +----------------- Anodonta_anatina **       
  |                                         
  +----------------- Unio                   
  |                                         
  +----------------- Cristariina            
  |                                         
  |              +-- Anodonta_nuttalliana **   
--+--------------+                          
  |              +-- Anodonta_californiensis **
  |                                         
  |              +-- Pseudodontoideus       
  |              |                          
  +--------------+-- Alasmidonta            
  |              |                          
  |              +-- Pterosyna              
  |                                         
  |  +-------------- Pseudanodonta **          
  |  |                                      
  |  |        +----- Strophitus             
  |  +--------+                             
  +--+        |  +-- Prolasmidonta          
     |        +--+                          
     |           +-- Platynaias             
     |                                      
     |  +----------- Anodonta_oregonensis__4 **
     +--+                                   
        |  +-------- Anodonta_oregonensis__3 **
        +--+                                
           |  +----- Anodonta_oregonensis__2 **
           +--+                             
              |  +-- Anodonta_oregonensis__1 **
              +--+                          
                 +-- Anodonta_kennerlyi **

Wu et al., 2024 [Publ 4379], fig. 5

iTtTbiriSoTroTroSiD
5121024813238
  +----------------- Cristariina         
--+                                      
  |  +-------------- Anodonta_nuttalliana **
  +--+                                   
     |  +----------- Alasmidontina       
     +--+                                
        |  +-------- Pseudanodonta **       
        +--+                             
           |  +----- Anodonta_anatina **    
           +--+                          
              |  +-- Anodonta_cygnea **     
              +--+                       
                 +-- Anodonta_exulcerata **

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iTtTbiriSoTroTroSiD
5121028713238
  +----------------- Cristariina         
--+                                      
  |  +-------------- Anodonta_nuttalliana **
  +--+                                   
     |  +----------- Alasmidontina       
     +--+                                
        |  +-------- Pseudanodonta **       
        +--+                             
           |  +----- Anodonta_anatina **    
           +--+                          
              |  +-- Anodonta_cygnea **     
              +--+                       
                 +-- Anodonta_exulcerata **

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iTtTbiriSoTroTroSiD
55329113239
  +-------------- Cristariina         
  |                                   
--+  +----------- Alasmidontina       
  |  |                                
  +--+        +-- Anodonta_nuttalliana **
     |  +-----+                       
     |  |     +-- Pseudanodonta **       
     +--+                             
        |  +----- Anodonta_anatina **    
        +--+                          
           |  +-- Anodonta_cygnea **     
           +--+                       
              +-- Anodonta_exulcerata **

Riccardi et al., 2020 [Publ 4089], fig. 2

iTtTbiriSoTroTroSiD
5972148239
  +-------------- Cristariina         
  |                                   
--+  +----------- Anodonta_anatina **    
  |  |                                
  +--+        +-- Anodonta_nuttalliana **
     |  +-----+                       
     |  |     +-- Alasmidontina       
     +--+                             
        |  +----- Pseudanodonta **       
        +--+                          
           |  +-- Anodonta_exulcerata ** 
           +--+                       
              +-- Anodonta_cygnea **

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iTtTbiriSoTroTroSiD
512829113236
  +----------- Cristariina         
  |                                
--+  +-------- Anodonta_nuttalliana **
  |  |                             
  |  +-------- Pseudanodonta **       
  +--+                             
     |  +----- Anodonta_cygnea **     
     |  |                          
     +--+----- Alasmidontina       
        |                          
        |  +-- Anodonta_exulcerata ** 
        +--+                       
           +-- Anodonta_anatina **

Lyubas et al., 2022 [Publ 4255], fig. 7

iTtTbiriSoTroTroSiD
5532260041
  +-- Pseudanodonta **
--+                
  +-- Anodonta **

Lyubas et al., 2022 [Publ 4255], fig. S1

iTtTbiriSoTroTroSiD
5532260041
  +----------- Anodonta_anatina **   
--+                               
  |  +-------- Pseudanodonta **      
  +--+                            
     |  +----- Anodonta_exulcerata **
     +--+                         
        |  +-- Anodonta_seddonae **  
        +--+                      
           +-- Anodonta_cygnea **

Hou et al., 2025 [Publ 4445], fig. 5

iTtTbiriSoTroTroSiD
5121024813238
  +----------------- Cristariina         
--+                                      
  |  +-------------- Anodonta_nuttalliana **
  +--+                                   
     |  +----------- Alasmidontina       
     +--+                                
        |  +-------- Pseudanodonta **       
        +--+                             
           |  +----- Anodonta_anatina **    
           +--+                          
              |  +-- Anodonta_cygnea **     
              +--+                       
                 +-- Anodonta_exulcerata **

Hou et al., 2024 [Publ 4400], fig. 4

iTtTbiriSoTroTroSiD
5121027113238
  +-------------- Cristariina         
  |                                   
--+           +-- Anodonta_nuttalliana **
  |  +--------+                       
  |  |        +-- Alasmidontina       
  +--+                                
     |  +-------- Pseudanodonta **       
     +--+                             
        |  +----- Anodonta_anatina **    
        +--+                          
           |  +-- Anodonta_cygnea **     
           +--+                       
              +-- Anodonta_exulcerata **

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iTtTbiriSoTroTroSiD
5121028713238
  +-------------- Cristariina         
  |                                   
--+           +-- Anodonta_nuttalliana **
  |  +--------+                       
  |  |        +-- Alasmidontina       
  +--+                                
     |  +-------- Pseudanodonta **       
     +--+                             
        |  +----- Anodonta_anatina **    
        +--+                          
           |  +-- Anodonta_cygnea **     
           +--+                       
              +-- Anodonta_exulcerata **

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

iTtTbiriSoTroTroSiD
44227210240
  +-------- Cristariina  
--+                      
  |  +----- Alasmidontina
  +--+                   
     |  +-- Pseudanodonta **
     +--+                
        +-- Anodonta **

Lopes-Lima et al., 2021 [Publ 4217], fig. 11

iTtTbiriSoTroTroSiD
442100040
1

Bolotov et al., 2017 [Publ 3761], fig. S1

iTtTbiriSoTroTroSiD
497213511243
  +-------------- Cristariina         
--+                                   
  |  +----------- Anodonta_anatina **    
  +--+                                
     |  +-------- Alasmidontina       
     +--+                             
        |  +----- Pseudanodonta **       
        +--+                          
           |  +-- Anodonta_cygnea **     
           +--+                       
              +-- Anodonta_nuttalliana **

Nagel et al., 1996 [Publ 3400], fig. 351

iTtTbiriSoTroTroSiD
454211141
  +-------------- Pseudanodonta **   
--+                               
  |  +----------- Anodonta_cygnea ** 
  +--+                            
     |  +-------- Cristariina     
     +--+                         
        |  +----- Anodonta__2 **     
        +--+                      
           |  +-- Anodonta_anatina **
           +--+                   
              +-- Anodonta__1 **

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
4114219716239
  +-------- Cristariina  
  |                      
  |  +----- Alasmidonta  
  |  |                   
--+  +----- Lasmigona    
  |  |                   
  |  +----- Pseudanodonta **
  |  |                   
  +--+----- Simpsonaias  
     |                   
     +----- Strophitus   
     |                   
     +----- Anodonta **     
     |                   
     |  +-- Pyganodon    
     +--+                
        +-- Utterbackia

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
44724219716273
  +-------- Acuticosta           
  |                              
  +-------- Alasmidonta          
  |                              
  +-------- Anodonta_anatina **     
  |                              
  +-------- Anemina              
  |                              
  +-------- Anodonta_cygnea **      
  |                              
  +-------- Cristaria            
  |                              
  +-------- Anodonta_nuttalliana ** 
  |                              
  +-------- Lasmigona            
  |                              
  +-------- Lepidodesma          
  |                              
  +-------- Pseudanodonta **        
  |                              
  +-------- Pyganodon            
  |                              
--+-------- Simpsonaias          
  |                              
  +-------- Strophitus           
  |                              
  +-------- Utterbackia          
  |                              
  +-------- Lanceolaria          
  |                              
  +-------- Schistodesmus        
  |                              
  +-------- Unio                 
  |                              
  +-------- Aculamprotula        
  |                              
  |     +-- Pletholophus         
  +-----+                        
  |     +-- Sinanodonta          
  |                              
  |  +----- Cuneopsis_celtiformis
  |  |                           
  |  +----- Tchangsinaia         
  |  |                           
  +--+----- Cuneopsis_rufescens  
     |                           
     +----- Nodularia            
     |                           
     |  +-- Cuneopsis_szechenyii 
     +--+                        
        +-- Cuneopsis_heudei

Riccardi et al., 2020 [Publ 4089], fig. 1

iTtTbiriSoTroTroSiD
442222138
  +-------- Cristariina        
  |                            
--+     +-- Anodonta_anatina **   
  |  +--+                      
  |  |  +-- Pseudanodonta **      
  +--+                         
     |  +-- Anodonta_exulcerata **
     +--+                      
        +-- Anodonta_cygnea **

Bolotov et al., 2017 [Publ 3761], fig. S2

iTtTbiriSoTroTroSiD
493213511240
  +----- Anodonta_anatina **    
  |                          
  +----- Pseudanodonta **       
--+                          
  +----- Alasmidontina       
  |                          
  |  +-- Anodonta_cygnea **     
  +--+                       
     +-- Anodonta_nuttalliana **

Tomilova et al., 2020 [Publ 4097], fig. 3

iTtTbiriSoTroTroSiD
442242140
  +-------- Anodonta_anatina **   
--+                            
  |  +----- Pseudanodonta **      
  +--+                         
     |  +-- Anodonta_cygnea **    
     +--+                      
        +-- Anodonta_exulcerata **

Nagel et al., 1998 [Publ 1946], fig. 7

iTtTbiriSoTroTroSiD
454211141
  +-------------- Pseudanodonta **   
--+                               
  |  +----------- Anodonta_cygnea ** 
  +--+                            
     |  +-------- Cristariina     
     +--+                         
        |  +----- Anodonta__2 **     
        +--+                      
           |  +-- Anodonta_anatina **
           +--+                   
              +-- Anodonta__1 **

Bolotov et al., 2017 [Publ 3764], fig. S1

iTtTbiriSoTroTroSiD
4170212111246
  +-- Alasmidonta          
  |                        
  +-- Anodonta_anatina **     
  |                        
  +-- Anodonta_cygnea **      
  |                        
  +-- Anodonta_nuttalliana ** 
  |                        
  +-- Lasmigona            
  |                        
  +-- Pseudanodonta **        
  |                        
  +-- Pyganodon            
  |                        
  +-- Simpsonaias          
  |                        
  +-- Strophitus           
--+                        
  +-- Sinanodonta_lauta    
  |                        
  +-- Cristaria_plicata    
  |                        
  +-- Cristaria_radiata    
  |                        
  +-- Pletholophus         
  |                        
  +-- Sinanodonta_lucida   
  |                        
  +-- Sinanodonta_jourdyi  
  |                        
  +-- Lanceolaria_gladiola 
  |                        
  +-- Lanceolaria_grayii__1
  |                        
  +-- Lanceolaria_grayii__2

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iTtTbiriSoTroTroSiD
44127610239
  +-------- Cristariina  
--+                      
  |  +----- Alasmidontina
  +--+                   
     |  +-- Pseudanodonta **
     +--+                
        +-- Anodonta **

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iTtTbiriSoTroTroSiD
494211829241
  +----- Cristariina  
  |                   
--+  +-- Pseudanodonta **
  |  |                
  +--+-- Anodonta **     
     |                
     +-- Alasmidontina

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
4118219516242
  +-------------- Cristariina  
  |                            
  |  +----------- Anodonta **     
--+  |                         
  |  |        +-- Alasmidonta  
  |  +--------+                
  +--+        +-- Lasmigona    
     |                         
     |        +-- Pyganodon    
     |  +-----+                
     |  |     +-- Utterbackia  
     +--+                      
        |  +----- Pseudanodonta **
        +--+                   
           |  +-- Simpsonaias  
           +--+                
              +-- Strophitus

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

iTtTbiriSoTroTroSiD
44227210240
  +-------- Cristariina  
--+                      
  |  +----- Alasmidontina
  +--+                   
     |  +-- Pseudanodonta **
     +--+                
        +-- Anodonta **

Mezhzherin et al., 2014 [Publ 3705], fig. 4

iTtTbiriSoTroTroSiD
331211139
  +-------- Cristariina     
--+                         
  |  +----- Anodonta_anatina **
  +--+                      
     |  +-- Anodonta_cygnea ** 
     +--+                   
        +-- Pseudanodonta **

Källersjö et al., 2005 [Publ 2684], fig. 1B-F

iTtTbiriSoTroTroSiD
331240039
  +----- Anodonta_anatina **
--+                      
  |  +-- Pseudanodonta **   
  +--+                   
     +-- Anodonta_cygnea **

Lopes-Lima et al., 2021 [Publ 4217], fig. 1

iTtTbiriSoTroTroSiD
3311220039
1

Froufe et al., 2017 [Publ 3802], fig. 7

iTtTbiriSoTroTroSiD
331171139
  +-- Cristariina
--+              
  +-- Anodonta **

Prié & Puillandre, 2014 [Publ 3529], fig. 2

iTtTbiriSoTroTroSiD
3312140039
  +----- Anodonta_anatina **
--+                      
  |  +-- Pseudanodonta **   
  +--+                   
     +-- Anodonta_cygnea **

Manendo et al., 2008 [Publ 2669], fig. 1

iTtTbiriSoTroTroSiD
3532102139
  +----- Anodonta **     
--+                   
  |  +-- Pseudanodonta **
  +--+                
     +-- Alasmidontina

Reis et al., 2013 [Publ 3580], fig. 4

iTtTbiriSoTroTroSiD
3312110037
  +----- Anodonta_cygnea ** 
--+                      
  |  +-- Anodonta_anatina **
  +--+                   
     +-- Pseudanodonta **

Lopes-Lima et al., 2017 [Publ 3734], fig. 1

iTtTbiriSoTroTroSiD
3312140039
  +----- Anodonta_anatina **
--+                      
  |  +-- Pseudanodonta **   
  +--+                   
     +-- Anodonta_cygnea **

Hinzmann et al., 2013 [Publ 3541], fig. 5

iTtTbiriSoTroTroSiD
331200039
  +-- Pseudanodonta **
--+                
  +-- Anodonta **

Reis et al., 2013 [Publ 3580], fig. 5

iTtTbiriSoTroTroSiD
3302120038
  +-- Anodonta_anatina **
  |                   
--+-- Anodonta_cygnea ** 
  |                   
  +-- Pseudanodonta **

Nagel et al., 1998 [Publ 1946], fig. 1

iTtTbiriSoTroTroSiD
331260039
  +-- Pseudanodonta **
--+                
  +-- Anodonta **

Wu et al., 2023 [Publ 4339], fig. 5

iTtTbiriSoTroTroSiD
3312166239
  +----------- Cristariina     
--+                            
  |  +-------- Alasmidontina   
  +--+                         
     |  +----- Anodonta_anatina **
     +--+                      
        |  +-- Pseudanodonta **   
        +--+                   
           +-- Anodonta_cygnea **

Wu et al., 2021 [Publ 4366], fig. 2_I

iTtTbiriSoTroTroSiD
3532887239
  +-------------- Cristariina  
--+                            
  |  +----------- Lasmigona    
  +--+                         
     |  +-------- Anodonta **     
     +--+                      
        |  +----- Pseudanodonta **
        +--+                   
           |  +-- Pyganodon    
           +--+                
              +-- Strophitus

Perkins et al., 2025 [Publ 4447], fig. 2_mtDNA-ITS

iTtTbiriSoTroTroSiD
3363723333132
  +----- Anodonta **     
--+                   
  |  +-- Pseudanodonta **
  +--+                
     +-- Alasmidontina

Plazzi et al., 2011 [Publ 3676], fig. 5

iTtTbiriSoTroTroSiD
317102152252
  +----------------- Cristariina  
  |                               
  |  +-------------- Lanceolaria  
  |  |                            
--+  +-------------- Anodonta **     
  |  |                            
  |  +-------------- Unio         
  |  |                            
  |  |           +-- Alasmidontina
  +--+-----------+                
     |           +-- Pseudanodonta **
     |                            
     |           +-- Alathyria    
     |           |                
     |  +--------+-- Pronodularia 
     |  |        |                
     |  |        +-- Sinohyriopsis
     +--+                         
        |  +-------- Quadrulina   
        |  |                      
        +--+-------- Parvaspina   
           |                      
           |  +----- Lampsilis    
           +--+                   
              |  +-- Epioblasma   
              +--+                
                 +-- Venustaconcha

Soroka, 2010 [Publ 3404], fig. 2

iTtTbiriSoTroTroSiD
331241137
  +----- Anodonta_cygnea ** 
--+                      
  |  +-- Anodonta_anatina **
  +--+                   
     +-- Pseudanodonta **

Froufe et al., 2017 [Publ 3802], fig. 2

iTtTbiriSoTroTroSiD
331111139
  +-- Cristariina
--+              
  +-- Anodonta **

Inoue et al., 2018 [Publ 3858], fig. 1_3a-c

iTtTbiriSoTroTroSiD
3200110117242
  +-- Alasmidonta_marginata      
  |                              
  +-- Prolasmidonta              
  |                              
  +-- Alasmidonta_undulata       
  |                              
  +-- Cristariina                
  |                              
  +-- Anodonta_kennerlyi **         
  |                              
  +-- Anodonta_nuttalliana **       
  |                              
  +-- Anodonta_oregonensis **       
  |                              
  +-- Arcidens_confragosus       
  |                              
  +-- Arcidens_wheeleri          
  |                              
  +-- Pterosyna                  
  |                              
--+-- Platynaias_compressa       
  |                              
  +-- Lasmigona                  
  |                              
  +-- Platynaias_decorata        
  |                              
  +-- Platynaias_subviridis      
  |                              
  +-- Pyganodon_grandis          
  |                              
  +-- Pyganodon_lacustris        
  |                              
  +-- Pseudodontoideus           
  |                              
  +-- Strophitus                 
  |                              
  +-- Utterbackia_imbecillis     
  |                              
  +-- Utterbackia_peninsularis__1
  |                              
  +-- Utterbackia_peninsularis__2

Källersjö et al., 2005 [Publ 2684], fig. 1A

iTtTbiriSoTroTroSiD
330240038
  +-- Anodonta_anatina **
  |                   
--+-- Pseudanodonta **   
  |                   
  +-- Anodonta_cygnea **

Bolotov et al., 2020 [Publ 4084], fig. 1

iTtTbiriSoTroTroSiD
3312339139
  +-------- Cristariina     
--+                         
  |  +----- Anodonta_anatina **
  +--+                      
     |  +-- Pseudanodonta **   
     +--+                   
        +-- Anodonta_cygnea **

Soroka, 2010 [Publ 3404], fig. 5a

iTtTbiriSoTroTroSiD
353252141
  +-------- Anodonta_anatina **
--+                         
  |  +----- Cristariina     
  +--+                      
     |  +-- Anodonta_cygnea ** 
     +--+                   
        +-- Pseudanodonta **

Lopes-Lima et al., 2016 [Publ 3757], fig. 1

iTtTbiriSoTroTroSiD
331200039
  +-- Pseudanodonta **
--+                
  +-- Anodonta **

Soroka, 2010 [Publ 3404], fig. 5b

iTtTbiriSoTroTroSiD
385252144
  +-------- Anodonta_anatina **
  |                         
--+  +----- Cristariina     
  |  |                      
  +--+----- Unio            
     |                      
     |  +-- Anodonta_cygnea ** 
     +--+                   
        +-- Pseudanodonta **

Mezhzherin et al., 2014 [Publ 3705], fig. 2

iTtTbiriSoTroTroSiD
331211139
  +----- Cristariina  
--+                   
  |  +-- Pseudanodonta **
  +--+                
     +-- Anodonta **

Gomes-dos-Santos et al., 2019 [Publ 4042], fig. 8

iTtTbiriSoTroTroSiD
331133139
  +-- Anodonta **   
--+              
  +-- Cristariina

Soroka, 2010 [Publ 3404], fig. 4

iTtTbiriSoTroTroSiD
375241143
        +-- Anodonta_anatina **
  +-----+                   
  |     +-- Unio            
--+                         
  |  +----- Cristariina     
  +--+                      
     |  +-- Anodonta_cygnea ** 
     +--+                   
        +-- Pseudanodonta **

Wu et al., 2021 [Publ 4366], fig. 6

iTtTbiriSoTroTroSiD
3642887240
  +-------------- Cristariina     
  |                               
--+  +----------- Anodonta_cygnea ** 
  |  |                            
  +--+        +-- Lasmigona       
     |  +-----+                   
     |  |     +-- Anodonta_anatina **
     +--+                         
        |  +----- Pseudanodonta **   
        +--+                      
           |  +-- Pyganodon       
           +--+                   
              +-- Strophitus

Froufe et al., 2014 [Publ 3592], fig. 2

iTtTbiriSoTroTroSiD
331200039
  +----- Anodonta_anatina **
--+                      
  |  +-- Pseudanodonta **   
  +--+                   
     +-- Anodonta_cygnea **

Soroka, 2010 [Publ 3404], fig. 3b

iTtTbiriSoTroTroSiD
395262144
  +----- Anodonta_anatina **
  |                      
  +----- Anodonta_cygnea ** 
--+                      
  +----- Pseudanodonta **   
  |                      
  |  +-- Cristariina     
  +--+                   
     +-- Unio

Konopleva et al., 2023 [Publ 4269], fig. S1_4

iTtTbiriSoTroTroSiD
33125313139
  +-------- Cristariina     
--+                         
  |  +----- Anodonta_anatina **
  +--+                      
     |  +-- Pseudanodonta **   
     +--+                   
        +-- Anodonta_cygnea **

Zieritz et al., 2012 [Publ 3673], fig. 4

iTtTbiriSoTroTroSiD
330251138
  +-- Pseudanodonta **   
  |                   
--+-- Anodonta_anatina **
  |                   
  +-- Anodonta_cygnea **

Pfarrer et al., 2022 [Publ 4305], fig. 2-5

iTtTbiriSoTroTroSiD
331192139
  +-- Cristariina
--+              
  +-- Anodonta **

Soroka, 2010 [Publ 3404], fig. 3a

iTtTbiriSoTroTroSiD
331262139
  +----- Anodonta_anatina **
--+                      
  |  +-- Anodonta_cygnea ** 
  +--+                   
     +-- Pseudanodonta **

Klishko et al., 2019 [Publ 3886], fig. 9

iTtTbiriSoTroTroSiD
2201900--
1

Lopes-Lima et al., 2024 [Publ 4376], fig. 4-5_AHE

iTtTbiriSoTroTroSiD
22012000--
1

Soroka, 2010 [Publ 3404], fig. 8b

iTtTbiriSoTroTroSiD
2201411--
1

Zanatta et al., 2007 [Publ 2698], fig. 1A

iTtTbiriSoTroTroSiD
213011414139
           +-- Platynaias        
           |                     
  +--------+-- Lasmigona         
  |        |                     
  |        +-- Pterosyna         
--+                              
  |  +-------- Anodonta_cygnea **   
  |  |                           
  +--+     +-- Anodonta_kennerlyi **
     |  +--+                     
     |  |  +-- Utterbackiana     
     +--+                        
        |  +-- Utterbackia       
        +--+                     
           +-- Pyganodon

Wu et al., 2022 [Publ 4238], fig. 3

iTtTbiriSoTroTroSiD
22017282--
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Nagel et al., 1998 [Publ 1946], fig. 9

iTtTbiriSoTroTroSiD
2202400--
  +-- Anodonta **     
--+                
  +-- Pseudanodonta **

Dai et al., 2025 [Publ 4440], fig. 2

iTtTbiriSoTroTroSiD
22024382--
  +----- Alasmidontina
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Pseudanodonta **

Huang et al., 2019 [Publ 3843], fig. 3

iTtTbiriSoTroTroSiD
2601548234
  +-------- Cristariina     
--+                         
  |  +----- Anodonta_cygnea ** 
  +--+                      
     |  +-- Anodonta_anatina **
     +--+                   
        +-- Alasmidontina

Soroka & Burzynski, 2017 [Publ 3837], fig. 1

iTtTbiriSoTroTroSiD
22012182--
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Wu et al., 2018 [Publ 3836], fig. 2

iTtTbiriSoTroTroSiD
27015111238
  +-------- Cristariina     
--+                         
  |  +----- Anodonta_cygnea ** 
  +--+                      
     |  +-- Anodonta_anatina **
     +--+                   
        +-- Alasmidontina

Dyduch-Falniowska & Koziol, 1989 [Publ 3814], fig. 5_7

iTtTbiriSoTroTroSiD
2202300--
  +-- Pseudanodonta **
--+                
  +-- Anodonta **

Chen et al., 2023 [Publ 4302], fig. 2

iTtTbiriSoTroTroSiD
22023982--
  +-------- Cristariina  
--+                      
  |  +----- Alasmidontina
  +--+                   
     |  +-- Anodonta **     
     +--+                
        +-- Pseudanodonta **

Lopes-Lima et al., 2024 [Publ 4376], fig. 4-5_MtG

iTtTbiriSoTroTroSiD
22012000--
1

Gustafson & Iwamoto, 2005 [Publ 2741], fig. 2

iTtTbiriSoTroTroSiD
2201311--
  +-- Cristariina
--+              
  +-- Anodonta **

Hoeh, 1990 [Publ 2717], fig. 2

iTtTbiriSoTroTroSiD
25011313140
           +-- Pterosyna         
           |                     
  +--------+-- Platynaias        
  |        |                     
  |        +-- Lasmigona         
--+                              
  |        +-- Utterbackia       
  |  +-----+                     
  |  |     +-- Pyganodon         
  +--+                           
     |  +----- Anodonta_cygnea **   
     +--+                        
        |  +-- Anodonta_kennerlyi **
        +--+                     
           +-- Utterbackiana

Bogatov et al., 2018 [Publ 3859], fig. 2

iTtTbiriSoTroTroSiD
2202400--
  +-- Anodonta **     
--+                
  +-- Pseudanodonta **

Wu et al., 2021 [Publ 4366], fig. 3-BI

iTtTbiriSoTroTroSiD
29017211233
  +-------- Cristariina     
--+                         
  |  +----- Anodonta_cygnea ** 
  +--+                      
     |  +-- Anodonta_anatina **
     +--+                   
        +-- Alasmidontina

Fukata & Iigo, 2020 [Publ 4106], fig. 1

iTtTbiriSoTroTroSiD
22015282--
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Sano et al., 2020 [Publ 4138], fig. 2

iTtTbiriSoTroTroSiD
220166172--
  +-- Anodonta **     
--+                
  +-- Alasmidontina

Dai et al., 2025 [Publ 4440], fig. 1

iTtTbiriSoTroTroSiD
22024382--
  +----- Alasmidontina
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Pseudanodonta **

Aunins et al., 2018 [Publ 3863], fig. 1

iTtTbiriSoTroTroSiD
22013892--
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Froufe et al., 2017 [Publ 3802], fig. 6

iTtTbiriSoTroTroSiD
220110101--
  +-- Anodonta **   
--+              
  +-- Cristariina

Wu et al., 2021 [Publ 4366], fig. 3-ML

iTtTbiriSoTroTroSiD
29017211233
  +-------- Cristariina     
--+                         
  |  +----- Anodonta_cygnea ** 
  +--+                      
     |  +-- Anodonta_anatina **
     +--+                   
        +-- Alasmidontina

Campbell et al., 2005 [Publ 2660], fig. 2

iTtTbiriSoTroTroSiD
220110041--
  +----- Pyganodon       
  |                      
  |  +-- Lasmigona       
--+--+                   
  |  +-- Pseudodontoideus
  |                      
  |  +-- Alasminota      
  +--+                   
     +-- Anodonta **

Zanatta et al., 2007 [Publ 2698], fig. 1B

iTtTbiriSoTroTroSiD
26011414140
           +-- Lasmigona         
           |                     
  +--------+-- Platynaias        
  |        |                     
  |        +-- Pterosyna         
--+                              
  |        +-- Utterbackia       
  |  +-----+                     
  |  |     +-- Pyganodon         
  +--+                           
     |  +----- Anodonta_cygnea **   
     +--+                        
        |  +-- Anodonta_kennerlyi **
        +--+                     
           +-- Utterbackiana

Chen et al., 2023 [Publ 4302], fig. 1

iTtTbiriSoTroTroSiD
22023982--
  +----- Alasmidontina
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Pseudanodonta **

Jadhav & Jamkhedkar, 2009 [Publ 3697], fig. 11

iTtTbiriSoTroTroSiD
2801234141
  +----------- Anodonta_cygnea ** 
  |                            
--+  +-------- Unio_pictorum__2
  |  |                         
  +--+     +-- Unio_pictorum__1
     |  +--+                   
     |  |  +-- Unio_caffer     
     +--+                      
        |  +-- Anodonta **        
        +--+                   
           +-- Alasmidontina

Dai et al., 2025 [Publ 4440], fig. 3

iTtTbiriSoTroTroSiD
22018682--
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Whelan et al., 2023 [Publ 4287], fig. 4

iTtTbiriSoTroTroSiD
220125251--
  +-- Anodonta **     
--+                
  +-- Alasmidontina

Chen et al., 2020 [Publ 4170], fig. 1

iTtTbiriSoTroTroSiD
22013292--
  +-------- Alasmidontina
--+                      
  |  +----- Sinanodonta  
  +--+                   
     |  +-- Cristaria    
     +--+                
        +-- Anodonta **

Xiang et al., 2025 [Publ 4433], fig. 7

iTtTbiriSoTroTroSiD
22011171--
  +-- Anodonta **   
--+              
  +-- Cristariina

Soroka, 2010 [Publ 3404], fig. 8a

iTtTbiriSoTroTroSiD
2201411--
  +-- Cristariina
--+              
  +-- Anodonta **

Wu et al., 2022 [Publ 4238], fig. 2d

iTtTbiriSoTroTroSiD
22013782--
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Bogatov et al., 2018 [Publ 3859], fig. 1

iTtTbiriSoTroTroSiD
2202400--
  +-- Anodonta **     
--+                
  +-- Pseudanodonta **

Hoeh, 1990 [Publ 2717], fig. 4

iTtTbiriSoTroTroSiD
25011313140
           +-- Pterosyna         
           |                     
  +--------+-- Platynaias        
  |        |                     
  |        +-- Lasmigona         
--+                              
  |        +-- Utterbackia       
  |  +-----+                     
  |  |     +-- Pyganodon         
  +--+                           
     |  +----- Anodonta_cygnea **   
     +--+                        
        |  +-- Anodonta_kennerlyi **
        +--+                     
           +-- Utterbackiana

Chen et al., 2025 [Publ 4448], fig. 3

iTtTbiriSoTroTroSiD
22018492--
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Sano et al., 2020 [Publ 4138], fig. 4

iTtTbiriSoTroTroSiD
220166172--
  +-- Anodonta **     
--+                
  +-- Alasmidontina

Hoeh, 1990 [Publ 2717], fig. 3

iTtTbiriSoTroTroSiD
215011313146
  +----- Pyganodon_fragilis         
  |                                 
  +----- Utterbackia_imbecillis     
  |                                 
  +----- Utterbackia_peggyae        
  |                                 
  +----- Anodonta_cygnea **            
  |                                 
  +----- Anodonta_kennerlyi **         
  |                                 
  +----- Utterbackiana_couperiana__1
  |                                 
--+----- Utterbackiana_couperiana__2
  |                                 
  +----- Utterbackiana_suborbiculata
  |                                 
  |  +-- Lasmigona                  
  +--+                              
  |  +-- Utterbackiana_implicata    
  |                                 
  |  +-- Pterosyna                  
  |  |                              
  |  +-- Platynaias                 
  |  |                              
  |  +-- Pyganodon_cataracta        
  +--+                              
     +-- Pyganodon_gibbosa          
     |                              
     +-- Pyganodon_grandis          
     |                              
     +-- Pyganodon_lacustris

Chase et al., 2018 [Publ 3847], fig. 4

iTtTbiriSoTroTroSiD
220136122--
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Wu et al., 2021 [Publ 4366], fig. 4

iTtTbiriSoTroTroSiD
28016710234
  +-------- Cristariina     
--+                         
  |  +----- Anodonta_cygnea ** 
  +--+                      
     |  +-- Anodonta_anatina **
     +--+                   
        +-- Alasmidontina

Campbell et al., 2005 [Publ 2660], fig. 1

iTtTbiriSoTroTroSiD
26011004140
  +----------- Anodonta_oregonensis **
  |                                
--+        +-- Anodonta_cygnea **     
  |  +-----+                       
  |  |     +-- Alasminota          
  +--+                             
     |  +----- Pyganodon           
     +--+                          
        |  +-- Lasmigona           
        +--+                       
           +-- Pseudodontoideus

Sano et al., 2020 [Publ 4138], fig. 3

iTtTbiriSoTroTroSiD
220166172--
  +-- Anodonta **     
--+                
  +-- Alasmidontina

Dai et al., 2024 [Publ 4386], fig. 5-BI

iTtTbiriSoTroTroSiD
11012011--
-- Anodonta_anatina **

Mock et al., 2004 [Publ 2686], fig. 4

iTtTbiriSoTroTroSiD
1101111--
  +-- Anodonta **   
--+              
  +-- Cristariina

Wu et al., 2024 [Publ 4388], fig. 6

iTtTbiriSoTroTroSiD
11015362--
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A

iTtTbiriSoTroTroSiD
11012951--
  +-- Anodonta **     
--+                
  +-- Alasmidontina

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

iTtTbiriSoTroTroSiD
11013852--
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Wu et al., 2022 [Publ 4238], fig. 2e

iTtTbiriSoTroTroSiD
11011732--
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Wu et al., 2019 [Publ 3864], fig. 3

iTtTbiriSoTroTroSiD
11014582--
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Wu et al., 2016 [Publ 3711], fig. 1

iTtTbiriSoTroTroSiD
11012062--
  +----------- Sinanodonta  
--+                         
  |  +-------- Anemina      
  +--+                      
     |  +----- Cristaria    
     +--+                   
        |  +-- Anodonta **     
        +--+                
           +-- Alasmidontina

Araujo et al., 2018 [Publ 3779], fig. 1

iTtTbiriSoTroTroSiD
11011721--
  +-- Anodonta **     
--+                
  +-- Alasmidontina

Bogan & Hoeh, 2000 [Publ 2650], fig. 1

iTtTbiriSoTroTroSiD
11013321--
  +-- Anodonta **     
--+                
  +-- Alasmidontina

Burzynski & Soroka, 2018 [Publ 3862], fig. 3

iTtTbiriSoTroTroSiD
1101332--
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Soroka, 2010 [Publ 3404], fig. 4-M

iTtTbiriSoTroTroSiD
1101111--
  +-- Anodonta **   
--+              
  +-- Cristariina

Rosenberg et al., 1997 [Publ 2720], fig. 2B

iTtTbiriSoTroTroSiD
11012131--
-- Anodonta **

Dai et al., 2024 [Publ 4358], fig. 4

iTtTbiriSoTroTroSiD
11015262--
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Guerra et al., 2017 [Publ 3772], fig. 2a

iTtTbiriSoTroTroSiD
11012472--
  +-------- Sinanodonta  
--+                      
  |  +----- Cristaria    
  +--+                   
     |  +-- Anodonta **     
     +--+                
        +-- Alasmidontina

Hoeh et al., 2002 [Publ 2653], fig. 2

iTtTbiriSoTroTroSiD
11013021--
  +-- Anodonta **     
--+                
  +-- Alasmidontina

Mock et al., 2004 [Publ 2686], fig. 3

iTtTbiriSoTroTroSiD
1101111--
  +-- Anodonta **   
--+              
  +-- Cristariina

Lydeard et al., 2000 [Publ 2654], fig. 2d

iTtTbiriSoTroTroSiD
11011300--
-- Anodonta_cygnea **

Wang et al., 2018 [Publ 3846], fig. 2

iTtTbiriSoTroTroSiD
11013062--
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Lopes-Lima et al., 2019 [Publ 3901], fig. 2

iTtTbiriSoTroTroSiD
11012711--
  +-- Anodonta **     
--+                
  +-- Alasmidontina

Soroka & Burzynski, 2018 [Publ 3762], fig. 3

iTtTbiriSoTroTroSiD
1101621--
  +-- Anodonta **     
--+                
  +-- Alasmidontina

Walker et al., 2006 [Publ 2666], fig. 2

iTtTbiriSoTroTroSiD
11011421--
  +-- Anodonta **     
--+                
  +-- Alasmidontina

Froufe et al., 2019 [Publ 3936], fig. 3-M

iTtTbiriSoTroTroSiD
11011832--
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Soroka, 2010 [Publ 3404], fig. 5a-M

iTtTbiriSoTroTroSiD
1101111--
  +-- Anodonta **   
--+              
  +-- Cristariina

Lydeard et al., 1996 [Publ 2656], fig. 4

iTtTbiriSoTroTroSiD
11012751--
     +----- Utterbackia  
  +--+                   
  |  |  +-- Utterbackiana
  |  +--+                
--+     +-- Anodonta **     
  |                      
  |  +----- Strophitus   
  +--+                   
     |  +-- Lasmigona    
     +--+                
        +-- Alasmidonta

Soroka, 2010 [Publ 3404], fig. 5b-M

iTtTbiriSoTroTroSiD
1101111--
  +-- Anodonta **   
--+              
  +-- Cristariina

Roe & Hoeh, 2003 [Publ 2655], fig. 4f3

iTtTbiriSoTroTroSiD
11013021--
  +-- Anodonta **     
--+                
  +-- Alasmidontina

Wu et al., 2020 [Publ 4147], fig. 3

iTtTbiriSoTroTroSiD
11012962--
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Dai et al., 2024 [Publ 4386], fig. 5-ML

iTtTbiriSoTroTroSiD
11012011--
-- Anodonta_anatina **

Taylor et al., 2007 [Publ 2696], fig. 1-3

iTtTbiriSoTroTroSiD
1101300--
-- Anodonta_cygnea **

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iTtTbiriSoTroTroSiD
11015632--
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Hoeh et al., 1998 [Publ 2695], fig. 4

iTtTbiriSoTroTroSiD
1101400--
-- Anodonta_cygnea **

Dai et al., 2024 [Publ 4386], fig. 3

iTtTbiriSoTroTroSiD
11011200--
-- Anodonta_anatina **

Huff et al., 2004 [Publ 2697], fig. 4

iTtTbiriSoTroTroSiD
11011000--
-- Anodonta **

Fonseca et al., 2016 [Publ 3775], fig. 1

iTtTbiriSoTroTroSiD
11013082--
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Araujo et al., 2009 [Publ 2589], fig. 7C

iTtTbiriSoTroTroSiD
1101900--
-- Anodonta_cygnea **

Soroka, 2010 [Publ 3404], fig. 3a-M

iTtTbiriSoTroTroSiD
1101521--
-- Anodonta_anatina **

Hoeh et al., 1998 [Publ 2695], fig. 3

iTtTbiriSoTroTroSiD
1101400--
-- Anodonta_cygnea **

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iTtTbiriSoTroTroSiD
11015632--
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

iTtTbiriSoTroTroSiD
11013642--
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Chase et al., 2018 [Publ 3861], fig. 1-M

iTtTbiriSoTroTroSiD
11011021--
  +-- Anodonta **     
--+                
  +-- Alasmidontina

Huff et al., 2004 [Publ 2697], fig. 3

iTtTbiriSoTroTroSiD
1101700--
-- Anodonta **

Soroka & Burzynski, 2018 [Publ 3762], fig. 3-M

iTtTbiriSoTroTroSiD
1101621--
  +-- Anodonta **     
--+                
  +-- Alasmidontina

Huang et al., 2019 [Publ 3843], fig. 3-M

iTtTbiriSoTroTroSiD
11012232--
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Huff et al., 2004 [Publ 2697], fig. 5-6

iTtTbiriSoTroTroSiD
11011100--
-- Anodonta **

Huang et al., 2018 [Publ 3828], fig. 6-M

iTtTbiriSoTroTroSiD
11011632--
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Guerra et al., 2017 [Publ 3772], fig. 2b

iTtTbiriSoTroTroSiD
11012472--
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Chase et al., 2018 [Publ 3861], fig. 1

iTtTbiriSoTroTroSiD
11011021--
  +-- Anodonta **     
--+                
  +-- Alasmidontina

Lydeard et al., 2000 [Publ 2654], fig. 2c

iTtTbiriSoTroTroSiD
11011300--
-- Anodonta_cygnea **

Cha et al., 2018 [Publ 3916], fig. 1

iTtTbiriSoTroTroSiD
11013782--
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

iTtTbiriSoTroTroSiD
11014782--
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Guerra et al., 2019 [Publ 4168], fig. 2-M

iTtTbiriSoTroTroSiD
11012132--
  +----- Alasmidontina
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Cristariina

Soroka, 2008 [Publ 3695], fig. 1-M

iTtTbiriSoTroTroSiD
1101322--
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Froufe et al., 2019 [Publ 3936], fig. 4

iTtTbiriSoTroTroSiD
11011832--
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Lopes-Lima et al., 2019 [Publ 3901], fig. 4

iTtTbiriSoTroTroSiD
11012511--
  +-- Anodonta **     
--+                
  +-- Alasmidontina

Soroka, 2010 [Publ 3404], fig. 2-M

iTtTbiriSoTroTroSiD
1101411--
-- Anodonta_anatina **

Walker et al., 2006 [Publ 2666], fig. 4

iTtTbiriSoTroTroSiD
11011421--
  +-- Anodonta **     
--+                
  +-- Alasmidontina

Bolotov et al., 2016 [Publ 3743], fig. 2

iTtTbiriSoTroTroSiD
11012900--
-- Anodonta_anatina **

Burzynski & Soroka, 2018 [Publ 3862], fig. 5b

iTtTbiriSoTroTroSiD
1101932--
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Krebs et al., 2003 [Publ 2691], fig. 3

iTtTbiriSoTroTroSiD
11013291--
  +----------- Utterbackiana
  |                         
  +----------- Platynaias   
  |                         
  +----------- Alasmidonta  
--+                         
  |        +-- Pterosyna    
  +--------+                
  |        +-- Lasmigona    
  |                         
  |  +-------- Strophitus   
  +--+                      
     |  +----- Utterbackia  
     +--+                   
        |  +-- Pyganodon    
        +--+                
           +-- Anodonta **

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B

iTtTbiriSoTroTroSiD
11012951--
  +-- Anodonta **     
--+                
  +-- Alasmidontina

Guerra et al., 2019 [Publ 4168], fig. 2

iTtTbiriSoTroTroSiD
11014062--
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Soroka, 2010 [Publ 3404], fig. 3b-M

iTtTbiriSoTroTroSiD
1101521--
-- Anodonta_anatina **

Liu et al., 2023 [Publ 4310], fig. 4

iTtTbiriSoTroTroSiD
11012331--
  +-- Pseudanodonta **
--+                
  +-- Alasmidontina

Feng et al., 2016 [Publ 3800], fig. 2

iTtTbiriSoTroTroSiD
11011352--
  +-- Anodonta **     
--+                
  +-- Alasmidontina

Bogan et al., 2023 [Publ 4295], fig. 3

iTtTbiriSoTroTroSiD
110130271--
  +-- Anodonta **   
--+              
  +-- Cristariina

Froufe et al., 2019 [Publ 3936], fig. 6

iTtTbiriSoTroTroSiD
11014382--
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Liu et al., 2023 [Publ 4310], fig. 5

iTtTbiriSoTroTroSiD
11012331--
  +-------- Lasmigona    
--+                      
  |  +----- Pseudanodonta **
  +--+                   
     |  +-- Pyganodon    
     +--+                
        +-- Strophitus

Bolotov et al., 2018 [Publ 3826], fig. 2

iTtTbiriSoTroTroSiD
11018292--
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Soroka, 2008 [Publ 3695], fig. 1

iTtTbiriSoTroTroSiD
1101322--
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Zieritz et al., 2024 [Publ 4381], fig. 2

iTtTbiriSoTroTroSiD
11014500--
-- Anodonta_cygnea **

Roe & Hoeh, 2003 [Publ 2655], fig. 4f2

iTtTbiriSoTroTroSiD
11013021--
  +----- Pyganodon 
--+                
  |  +-- Anodonta **  
  +--+             
     +-- Strophitus

Araujo et al., 2009 [Publ 2589], fig. 7A

iTtTbiriSoTroTroSiD
11011000--
-- Anodonta_cygnea **

Lopes-Lima et al., 2019 [Publ 3901], fig. 3

iTtTbiriSoTroTroSiD
11012811--
  +-- Anodonta **     
--+                
  +-- Alasmidontina

Chase et al., 2018 [Publ 3847], fig. 5

iTtTbiriSoTroTroSiD
1101911--
  +-- Anodonta **     
--+                
  +-- Alasmidontina

Sonowal et al., 2021 [Publ 4221], fig. 4

iTtTbiriSoTroTroSiD
11011500--
-- Anodonta_cygnea **

Walker et al., 2006 [Publ 2666], fig. 3

iTtTbiriSoTroTroSiD
11011421--
  +-- Anodonta **     
--+                
  +-- Alasmidontina

Wen et al., 2017 [Publ 3795], fig. 3

iTtTbiriSoTroTroSiD
11011521--
  +-- Anodonta **   
--+              
  +-- Cristariina

Zieritz et al., 2021 [Publ 4231], fig. 3-M_4

iTtTbiriSoTroTroSiD
11012232--
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Lopes-Lima et al., 2017 [Publ 3792], fig. 2-M

iTtTbiriSoTroTroSiD
1101921--
  +-- Anodonta **     
--+                
  +-- Alasmidontina

Araujo et al., 2017 [Publ 3801], fig. 2

iTtTbiriSoTroTroSiD
11011300--
-- Anodonta_anatina **

Araujo et al., 2009 [Publ 2589], fig. 7B

iTtTbiriSoTroTroSiD
11011000--
-- Anodonta_cygnea **

Sonowal et al., 2021 [Publ 4221], fig. 3b

iTtTbiriSoTroTroSiD
11011400--
-- Anodonta_cygnea **

Cyr et al., 2007 [Publ 2745], fig. 2B

iTtTbiriSoTroTroSiD
1101441--
  +-- Anodonta **     
--+                
  +-- Alasmidontina

Lopes-Lima et al., 2017 [Publ 3792], fig. 2

iTtTbiriSoTroTroSiD
11013182--
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Whelan et al., 2023 [Publ 4287], fig. 3

iTtTbiriSoTroTroSiD
110127271--
  +-- Anodonta **     
--+                
  +-- Alasmidontina

Burzynski & Soroka, 2018 [Publ 3862], fig. 5a

iTtTbiriSoTroTroSiD
11011932--
  +-- Cristariina  
  |                
--+-- Anodonta **     
  |                
  +-- Alasmidontina

Froufe et al., 2015 [Publ 3664], fig. 1-M

iTtTbiriSoTroTroSiD
1101721--
  +-- Anodonta **     
--+                
  +-- Alasmidontina

Guerra et al., 2019 [Publ 4168], fig. 1-M

iTtTbiriSoTroTroSiD
11012132--
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Hoeh et al., 2001 [Publ 2652], fig. 14f2

iTtTbiriSoTroTroSiD
11013021--
  +----- Pyganodon 
--+                
  |  +-- Anodonta **  
  +--+             
     +-- Strophitus

Rosenberg et al., 1997 [Publ 2720], fig. 2A

iTtTbiriSoTroTroSiD
11012131--
-- Anodonta_anatina **

Lydeard et al., 2000 [Publ 2654], fig. 2a

iTtTbiriSoTroTroSiD
11011300--
-- Anodonta_cygnea **

Gomes-dos-Santos et al., 2019 [Publ 3943], fig. 1-M

iTtTbiriSoTroTroSiD
1101500--
-- Anodonta_anatina **

Huang et al., 2018 [Publ 3828], fig. 6

iTtTbiriSoTroTroSiD
11014182--
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Burzynski et al., 2017 [Publ 3774], fig. 1-bottom

iTtTbiriSoTroTroSiD
11011221--
  +-- Anodonta **     
--+                
  +-- Alasmidontina

Froufe et al., 2015 [Publ 3664], fig. 1

iTtTbiriSoTroTroSiD
11011121--
  +-- Anodonta **     
--+                
  +-- Alasmidontina

Wu et al., 2020 [Publ 4121], fig. 3

iTtTbiriSoTroTroSiD
11014882--
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Wu et al., 2018 [Publ 4143], fig. 2

iTtTbiriSoTroTroSiD
11012162--
  +----------- Sinanodonta  
--+                         
  |  +-------- Anemina      
  +--+                      
     |  +----- Cristaria    
     +--+                   
        |  +-- Anodonta **     
        +--+                
           +-- Alasmidontina

Wu et al., 2023 [Publ 4296], fig. 4

iTtTbiriSoTroTroSiD
11012231--
  +-- Pseudanodonta **
--+                
  +-- Alasmidontina

Huff et al., 2004 [Publ 2697], fig. 2

iTtTbiriSoTroTroSiD
11011100--
-- Anodonta **

Froufe et al., 2019 [Publ 3936], fig. 3

iTtTbiriSoTroTroSiD
11014382--
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Guerra et al., 2019 [Publ 4168], fig. 1

iTtTbiriSoTroTroSiD
11014062--
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Wen et al., 2017 [Publ 3795], fig. 3-M

iTtTbiriSoTroTroSiD
1101521--
  +-- Anodonta **     
--+                
  +-- Alasmidontina

Sonowal et al., 2021 [Publ 4221], fig. 3a

iTtTbiriSoTroTroSiD
11011400--
-- Anodonta_cygnea **

Lydeard et al., 2000 [Publ 2654], fig. 2b

iTtTbiriSoTroTroSiD
11011300--
-- Anodonta_cygnea **

Bolotov et al., 2016 [Publ 3743], fig. 4

iTtTbiriSoTroTroSiD
11012900--
-- Anodonta_anatina **

Dai et al., 2023 [Publ 4285], fig. 5

iTtTbiriSoTroTroSiD
11015262--
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Fonseca et al., 2016 [Publ 3775], fig. 1-M

iTtTbiriSoTroTroSiD
1101821--
  +-- Anodonta **     
--+                
  +-- Alasmidontina

Gomes-dos-Santos et al., 2019 [Publ 3943], fig. 1

iTtTbiriSoTroTroSiD
1101900--
-- Anodonta_anatina **

Burzynski et al., 2017 [Publ 3774], fig. 1-top

iTtTbiriSoTroTroSiD
11011221--
  +-- Anodonta **     
--+                
  +-- Alasmidontina

Hoeh et al., 2001 [Publ 2652], fig. 14f4

iTtTbiriSoTroTroSiD
11013021--
  +-- Anodonta **     
--+                
  +-- Alasmidontina

Wang et al., 2018 [Publ 3865], fig. 2

iTtTbiriSoTroTroSiD
11013062--
  +----- Cristariina  
--+                   
  |  +-- Anodonta **     
  +--+                
     +-- Alasmidontina

Cyr et al., 2007 [Publ 2745], fig. 2C

iTtTbiriSoTroTroSiD
1101441--
  +-- Pseudanodonta **
--+                
  +-- Alasmidontina
 
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