The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)
FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!
Bivalvia | Unionoida | Unionoidea | Unionidae | Unioninae tribe ANODONTINI Rafinesque, 1820: cladomics
3 subtribes, 1 genus Cladomics Summarynumber of species = 95 (116 total)
number of subtaxa (rank: subtribe) = 4
analyzed in 336 cladograms in 187 publications
maximum number of terminals in an analysis =
41 (mean = 6.27, median = 4)
maximum number of subtaxa (subtribe) =
3 (mean = 1.79, median = 2) Basic Monophylynumber of cladograms that test basic monophyly
(i.e., more than 1 included species) = 285
maximum number of included terminals = 41, (mean =
7.21 median = 5)
maximum number of subtaxa (subtribe) =
3 (mean = 1.93, median = 2)
proportion of cladograms that support basic monophyly
= 0.86 Rigorous Monophylynumber of cladograms that test rigorous monophyly = 133
(Rigorous monophyly refers to cladograms with terminal species representing
more than one subtaxon (subtribe) taxon and at least one rigorous outgroup (family).
maximum number of included terminals = 33 (mean = 8.27, median = 7)
maximum number of subtaxa (subtribe) =
3 (mean = 2.59, median = 3)
proportion of cladograms that support rigorous monophyly = 0.77
Pairwise Cladogram ClusteringThe graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.
Dai et al., 2025 [Publ 4440], fig. 1 ** Top Monophyly Cladogram ** iT| tT | bi | riS | oT | roT | roS | iD | | 10 | 39 | 32 | 3 | 35 | 26 | 4 | 0 | +-- Anemina **
+--------------+
| +-- Amuranodonta **
|
| +----- Parreysiini
+-----------+
| | +-- Coelaturini
| +--+
--+ +-- Indochinellini
|
| +----- Pletholophus **
| +--------+
| | | +-- Sinanodonta **
| | +--+
| | +-- Beringiana **
+--+
| +----------- Cristaria **
| |
| | +-- Alasmidontina **
+--+--------+
| +-- Anodontina **
|
| +-------- Lanceolariini
+--+
| +----- Lepidodesmini
+--+
| +-- Acuticostini
+--+
+-- Unionini
Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4 ** Top Sister-Group Cladogram ** iT| tT | bi | riS | oT | roT | roS | iD | | 33 | 33 | 25 | 3 | 88 | 37 | 4 | 57 | +----------- Acuticostini
|
+----------- Lepidodesmini
|
--+----------- Unionini
|
| +-------- Lanceolariini
| |
+--+ +----- Cristariina **
| |
+--+ +-- Pseudanodonta **
| |
+--+-- Anodonta **
|
+-- Alasmidontina **
Whelan et al., 2023 [Publ 4287], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 41 | 41 | 42 | 3 | 0 | 0 | 0 | 74 | +----------- Cristariina **
|
--+ +-------- Anodonta_cygnea **
| |
+--+ +-- Anodonta_nuttalliana **
| +--+
| | +-- Pseudanodonta **
+--+
| +-- Anodonta_anatina **
+--+
+-- Alasmidontina **
Wu et al., 2025 [Publ 4442], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 33 | 33 | 31 | 1 | 2 | 2 | 1 | 64 | +-- Unionini
--+
+-- Cristariina **
Bogan et al., 2023 [Publ 4295], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 28 | 28 | 25 | 2 | 3 | 1 | 1 | 59 | +----- Unionini
--+
| +-- Anodontina **
+--+
+-- Cristariina **
Whelan et al., 2023 [Publ 4287], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 28 | 28 | 27 | 2 | 0 | 0 | 0 | 63 | +-- Anodontina **
--+
+-- Alasmidontina **
Dai et al., 2024 [Publ 4385], fig. 3-ML iT| tT | bi | riS | oT | roT | roS | iD | | 27 | 27 | 25 | 1 | 4 | 0 | 0 | 54 | Dai et al., 2024 [Publ 4385], fig. 3-BI iT| tT | bi | riS | oT | roT | roS | iD | | 27 | 27 | 24 | 1 | 4 | 0 | 0 | 54 | Xiang et al., 2025 [Publ 4433], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 27 | 27 | 25 | 1 | 1 | 0 | 0 | 59 | Whelan et al., 2023 [Publ 4287], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 27 | 27 | 25 | 2 | 0 | 0 | 0 | 63 | +-- Anodontina **
--+
+-- Alasmidontina **
Huang et al., 2019 [Publ 3843], fig. S3 iT| tT | bi | riS | oT | roT | roS | iD | | 20 | 20 | 17 | 3 | 177 | 24 | 4 | 49 | +-------------------------- Acuticostini
|
--+ +----------------------- Unionini
| |
+--+ +-------------------- Lepidodesmini
| |
+--+ +----------------- Lanceolariini
| |
+--+ +-------------- Cristariina **
| |
| | +----------- Anodonta **
+--+ |
| | +-- Alasmidonta **
| +--------+
+--+ +-- Lasmigona **
|
| +-- Pyganodon **
| +-----+
| | +-- Utterbackia **
+--+
| +----- Pseudanodonta **
+--+
| +-- Simpsonaias **
+--+
+-- Strophitus **
Huang et al., 2019 [Publ 3843], fig. S4 iT| tT | bi | riS | oT | roT | roS | iD | | 20 | 45 | 22 | 3 | 179 | 25 | 4 | 54 | +-------- Acuticostini
|
+-------- Alasmidonta **
|
+-------- Anodonta_anatina **
|
+-------- Anemina **
|
+-------- Anodonta_cygnea **
|
+-------- Cristaria **
|
+-------- Anodonta_nuttalliana **
|
+-------- Lasmigona **
|
+-------- Lepidodesmini
|
+-------- Pseudanodonta **
|
+-------- Pyganodon **
|
--+-------- Simpsonaias **
|
+-------- Strophitus **
|
+-------- Schistodesmus
|
+-------- Utterbackia **
|
+-------- Lanceolariini
|
+-------- Unio
|
+-------- Aculamprotula
|
| +-- Pletholophus **
+-----+
| +-- Sinanodonta **
|
| +----- Cuneopsis_celtiformis
| |
| +----- Tchangsinaia
| |
+--+----- Cuneopsis_rufescens
|
+----- Nodularia
|
| +-- Cuneopsis_szechenyii
+--+
+-- Cuneopsis_heudei
Huang et al., 2019 [Publ 3843], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 20 | 20 | 13 | 3 | 179 | 25 | 4 | 50 | +----------------- Acuticostini
|
--+ +-------------- Lepidodesmini
| |
| +-------------- Unionini
+--+
| +----------- Lanceolariini
| |
| | +-------- Cristariina **
+--+ |
| | +----- Alasmidonta **
| | |
+--+ +----- Lasmigona **
| |
| +----- Pseudanodonta **
| |
+--+----- Simpsonaias **
|
+----- Strophitus **
|
+----- Anodonta **
|
| +-- Pyganodon **
+--+
+-- Utterbackia **
Inoue et al., 2018 [Publ 3858], fig. 1_3a-c iT| tT | bi | riS | oT | roT | roS | iD | | 20 | 20 | 0 | 3 | 84 | 0 | 0 | 54 | +-- Alasmidonta_marginata **
|
+-- Prolasmidonta **
|
+-- Alasmidonta_undulata **
|
+-- Cristariina **
|
+-- Anodonta_kennerlyi **
|
+-- Anodonta_nuttalliana **
|
+-- Anodonta_oregonensis **
|
+-- Arcidens_confragosus **
|
+-- Arcidens_wheeleri **
|
+-- Lasmigona_complanata **
|
--+-- Platynaias_compressa **
|
+-- Lasmigona_costata **
|
+-- Platynaias_decorata **
|
+-- Platynaias_subviridis **
|
+-- Pyganodon_grandis **
|
+-- Pyganodon_lacustris **
|
+-- Pseudodontoideus **
|
+-- Strophitus **
|
+-- Utterbackia_imbecillis **
|
+-- Utterbackia_peninsularis__1 **
|
+-- Utterbackia_peninsularis__2 **
Sano et al., 2020 [Publ 4138], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 19 | 35 | 37 | 3 | 49 | 16 | 3 | 54 | +-- Anemina **
+--------------+
| +-- Buldowskia **
|
--+ +----- Sinanodonta **
| +--------+
| | | +-- Pletholophus **
| | +--+
+--+ +-- Beringiana **
|
| +----------- Cristaria **
| |
+--+ +-- Anodontina **
| +-----+
| | +-- Alasmidontina **
+--+
| +----- Lepidodesmini
+--+
| +-- Lanceolariini
+--+
+-- Unionini
Sano et al., 2020 [Publ 4138], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 19 | 35 | 37 | 3 | 49 | 16 | 3 | 52 | +-- Anemina **
+--------------+
| +-- Buldowskia **
|
--+ +----- Sinanodonta **
| +--------+
| | | +-- Pletholophus **
| | +--+
+--+ +-- Beringiana **
|
| +----------- Cristaria **
| |
+--+ +-- Anodontina **
| +-----+
| | +-- Alasmidontina **
+--+
| +----- Lanceolariini
+--+
| +-- Lepidodesmini
+--+
+-- Unionini
Sano et al., 2020 [Publ 4138], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 19 | 35 | 36 | 3 | 49 | 16 | 3 | 50 | +-- Anemina **
+--------------+
| +-- Buldowskia **
|
--+ +----- Pletholophus **
| +--------+
| | | +-- Beringiana **
| | +--+
+--+ +-- Sinanodonta **
|
| +----------- Cristaria **
| |
+--+ +-- Anodontina **
| +-----+
| | +-- Alasmidontina **
+--+
| +----- Lanceolariini
+--+
| +-- Lepidodesmini
+--+
+-- Unionini
Sano et al., 2022 [Publ 4311], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 18 | 18 | 16 | 1 | 0 | 0 | 0 | 51 | Wu et al., 2024 [Publ 4379], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 18 | 18 | 16 | 3 | 35 | 33 | 3 | 48 | +-------------------------- Lepidodesmini
--+
| +----------------------- Unionini
+--+
| +-------------------- Lanceolariini
+--+
| +----------------- Cristariina **
+--+
| +-------------- Anodonta_nuttalliana **
+--+
| +----------- Alasmidontina **
+--+
| +-------- Pseudanodonta **
+--+
| +----- Anodonta_anatina **
+--+
| +-- Anodonta_cygnea **
+--+
+-- Anodonta_exulcerata **
Hou et al., 2024 [Publ 4400], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 18 | 18 | 16 | 3 | 58 | 54 | 3 | 48 | +----------------------- Lepidodesmini
|
--+ +-------------------- Unionini
| |
+--+ +----------------- Lanceolariini
| |
+--+ +-------------- Cristariina **
| |
+--+ +-- Anodonta_nuttalliana **
| +--------+
| | +-- Alasmidontina **
+--+
| +-------- Pseudanodonta **
+--+
| +----- Anodonta_anatina **
+--+
| +-- Anodonta_cygnea **
+--+
+-- Anodonta_exulcerata **
Smith et al., 2018 [Publ 3790], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 18 | 18 | 17 | 2 | 0 | 0 | 0 | 54 | +-- Cristariina **
--+
+-- Alasmidontina **
Wu et al., 2024 [Publ 4350], fig. 8a_S1 iT| tT | bi | riS | oT | roT | roS | iD | | 18 | 18 | 16 | 3 | 73 | 28 | 4 | 48 | +----------------------- Lepidodesmini
|
--+ +-------------------- Unionini
| |
+--+ +-- Acuticostini
| +--------------+
| | +-- Lanceolariini
+--+
| +-------------- Cristariina **
| |
+--+ +-- Anodonta_nuttalliana **
| +--------+
| | +-- Alasmidontina **
+--+
| +-------- Pseudanodonta **
+--+
| +----- Anodonta_anatina **
+--+
| +-- Anodonta_cygnea **
+--+
+-- Anodonta_exulcerata **
Wu et al., 2024 [Publ 4350], fig. 8b_S2 iT| tT | bi | riS | oT | roT | roS | iD | | 18 | 18 | 16 | 3 | 73 | 28 | 4 | 48 | +-------------------------- Lepidodesmini
|
--+ +----------------------- Unionini
| |
+--+ +-- Acuticostini
| +-----------------+
| | +-- Lanceolariini
+--+
| +----------------- Cristariina **
+--+
| +-------------- Anodonta_nuttalliana **
+--+
| +----------- Alasmidontina **
+--+
| +-------- Pseudanodonta **
+--+
| +----- Anodonta_anatina **
+--+
| +-- Anodonta_cygnea **
+--+
+-- Anodonta_exulcerata **
Wu et al., 2023 [Publ 4339], fig. 4_S5-nt iT| tT | bi | riS | oT | roT | roS | iD | | 18 | 18 | 14 | 3 | 77 | 28 | 4 | 50 | +-------------------- Lepidodesmini
|
--+ +----------------- Unionini
| |
+--+ +-- Acuticostini
| +-----------+
| | +-- Lanceolariini
+--+
| +----------- Cristariina **
| |
+--+ +-------- Anodonta_nuttalliana **
| |
| +-------- Pseudanodonta **
+--+
| +----- Anodonta_cygnea **
| |
+--+----- Alasmidontina **
|
| +-- Anodonta_exulcerata **
+--+
+-- Anodonta_anatina **
Wu et al., 2023 [Publ 4339], fig. 4_S5-aa iT| tT | bi | riS | oT | roT | roS | iD | | 18 | 18 | 16 | 3 | 77 | 28 | 4 | 48 | +-------------- Lepidodesmini
|
--+ +----------- Unionini
| |
+--+ +-- Acuticostini
| +-----+
| | +-- Lanceolariini
+--+
| +----- Cristariina **
+--+
| +-- Anodontina **
+--+
+-- Alasmidontina **
Chong et al., 2008 [Publ 2668], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 17 | 18 | 10 | 3 | 1 | 1 | 1 | 48 | +----------------- Anodonta_cygnea **
|
+----------------- Pyganodon **
|
+----------------- Alasminota **
|
+----------------- Anodonta_anatina **
|
+----------------- Unionini
|
+----------------- Cristariina **
|
| +-- Anodonta_nuttalliana **
--+--------------+
| +-- Anodonta_californiensis **
|
| +-- Pseudodontoideus **
| |
+--------------+-- Alasmidonta **
| |
| +-- Lasmigona **
|
| +-------------- Pseudanodonta **
| |
| | +----- Strophitus **
| +--------+
+--+ | +-- Prolasmidonta **
| +--+
| +-- Platynaias **
|
| +----------- Anodonta_oregonensis__4 **
+--+
| +-------- Anodonta_oregonensis__3 **
+--+
| +----- Anodonta_oregonensis__2 **
+--+
| +-- Anodonta_oregonensis__1 **
+--+
+-- Anodonta_kennerlyi **
Zanatta et al., 2007 [Publ 2698], fig. 1A iT| tT | bi | riS | oT | roT | roS | iD | | 16 | 16 | 10 | 2 | 0 | 0 | 0 | 52 | +-- Platynaias **
|
+--------+-- Lasmigona_costata **
| |
| +-- Lasmigona_complanata **
--+
| +-------- Anodonta_cygnea **
| |
+--+ +-- Anodonta_kennerlyi **
| +--+
| | +-- Utterbackiana **
+--+
| +-- Utterbackia **
+--+
+-- Pyganodon **
Konopleva et al., 2023 [Publ 4269], fig. S1_4 iT| tT | bi | riS | oT | roT | roS | iD | | 16 | 16 | 14 | 2 | 40 | 17 | 4 | 45 | +----------- Unionini
|
--+ +-- Lepidodesmini
| +-----+
| | +-- Lanceolariini
+--+
| +----- Acuticostini
+--+
| +-- Anodontina **
+--+
+-- Cristariina **
Zanatta et al., 2007 [Publ 2698], fig. 1B iT| tT | bi | riS | oT | roT | roS | iD | | 16 | 16 | 13 | 2 | 0 | 0 | 0 | 52 | +-- Lasmigona_costata **
|
+--------+-- Platynaias **
| |
| +-- Lasmigona_complanata **
--+
| +-- Utterbackia **
| +-----+
| | +-- Pyganodon **
+--+
| +----- Anodonta_cygnea **
+--+
| +-- Anodonta_kennerlyi **
+--+
+-- Utterbackiana **
Bolotov et al., 2017 [Publ 3761], fig. S1 iT| tT | bi | riS | oT | roT | roS | iD | | 15 | 15 | 13 | 3 | 124 | 12 | 2 | 46 | +-------------------- Unionini
--+
| +----------------- Lanceolariini
+--+
| +-------------- Cristariina **
+--+
| +----------- Anodonta_anatina **
+--+
| +-------- Alasmidontina **
+--+
| +----- Pseudanodonta **
+--+
| +-- Anodonta_cygnea **
+--+
+-- Anodonta_nuttalliana **
Hoeh, 1990 [Publ 2717], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 15 | 15 | 13 | 2 | 0 | 0 | 0 | 51 | +-- Lasmigona_complanata **
|
+--------+-- Platynaias **
| |
| +-- Lasmigona_costata **
--+
| +-- Utterbackia **
| +-----+
| | +-- Pyganodon **
+--+
| +----- Anodonta_cygnea **
+--+
| +-- Anodonta_kennerlyi **
+--+
+-- Utterbackiana **
Bolotov et al., 2017 [Publ 3764], fig. S1 iT| tT | bi | riS | oT | roT | roS | iD | | 15 | 17 | 0 | 3 | 110 | 12 | 2 | 47 | +----- Unionini
|
| +-- Alasmidonta **
| |
| +-- Anodonta_anatina **
| |
| +-- Anodonta_cygnea **
| |
| +-- Anodonta_nuttalliana **
--+ |
| +-- Lasmigona **
| |
| +-- Pseudanodonta **
| |
| +-- Pyganodon **
| |
| +-- Simpsonaias **
| |
| +-- Strophitus **
+--+
+-- Sinanodonta_lauta **
|
+-- Cristaria_plicata **
|
+-- Cristaria_radiata **
|
+-- Pletholophus **
|
+-- Sinanodonta_lucida **
|
+-- Sinanodonta_jourdyi **
|
+-- Lanceolaria_gladiola
|
+-- Lanceolaria_grayii__1
|
+-- Lanceolaria_grayii__2
Bolotov et al., 2017 [Publ 3761], fig. S2 iT| tT | bi | riS | oT | roT | roS | iD | | 15 | 17 | 10 | 3 | 124 | 12 | 2 | 45 | +----------- Unionini
|
--+ +-------- Lanceolariini
| |
| +-------- Cristariina **
+--+
| +----- Anodonta_anatina **
| |
| +----- Pseudanodonta **
+--+
+----- Alasmidontina **
|
| +-- Anodonta_cygnea **
+--+
+-- Anodonta_nuttalliana **
Hoeh, 1990 [Publ 2717], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 15 | 15 | 2 | 2 | 0 | 0 | 0 | 51 | +----- Pyganodon_fragilis **
|
+----- Utterbackia_imbecillis **
|
+----- Utterbackia_peggyae **
|
+----- Anodonta_cygnea **
|
+----- Anodonta_kennerlyi **
|
+----- Utterbackiana_couperiana__1 **
|
--+----- Utterbackiana_couperiana__2 **
|
+----- Utterbackiana_suborbiculata **
|
| +-- Lasmigona_costata **
+--+
| +-- Utterbackiana_implicata **
|
| +-- Lasmigona_complanata **
| |
| +-- Platynaias **
| |
| +-- Pyganodon_cataracta **
+--+
+-- Pyganodon_gibbosa **
|
+-- Pyganodon_grandis **
|
+-- Pyganodon_lacustris **
Hoeh, 1990 [Publ 2717], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 15 | 15 | 13 | 2 | 0 | 0 | 0 | 51 | +-- Lasmigona_complanata **
|
+--------+-- Platynaias **
| |
| +-- Lasmigona_costata **
--+
| +-- Utterbackia **
| +-----+
| | +-- Pyganodon **
+--+
| +----- Anodonta_cygnea **
+--+
| +-- Anodonta_kennerlyi **
+--+
+-- Utterbackiana **
Wu et al., 2025 [Publ 4442], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 15 | 15 | 13 | 1 | 2 | 2 | 1 | 48 | +-- Unionini
--+
+-- Cristariina **
Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2 iT| tT | bi | riS | oT | roT | roS | iD | | 14 | 14 | 12 | 3 | 62 | 12 | 2 | 43 | +----------- Unionini
--+
| +-------- Lanceolariini
+--+
| +----- Cristariina **
+--+
| +-- Anodontina **
+--+
+-- Alasmidontina **
Chase et al., 2018 [Publ 3847], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 14 | 14 | 13 | 3 | 24 | 2 | 1 | 45 | +-------- Unionini
--+
| +----- Cristariina **
+--+
| +-- Anodontina **
+--+
+-- Alasmidontina **
Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1 iT| tT | bi | riS | oT | roT | roS | iD | | 14 | 14 | 12 | 3 | 62 | 12 | 2 | 43 | +----------- Unionini
--+
| +-------- Lanceolariini
+--+
| +----- Cristariina **
+--+
| +-- Anodontina **
+--+
+-- Alasmidontina **
Inoue et al., 2014 [Publ 3635], fig. 3A iT| tT | bi | riS | oT | roT | roS | iD | | 14 | 14 | 13 | 1 | 6 | 0 | 0 | 49 | Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2 iT| tT | bi | riS | oT | roT | roS | iD | | 14 | 14 | 11 | 3 | 66 | 14 | 2 | 43 | +----------- Unionini
--+
| +-------- Lanceolariini
+--+
| +----- Cristariina **
+--+
| +-- Anodontina **
+--+
+-- Alasmidontina **
Wu et al., 2018 [Publ 3836], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 13 | 13 | 9 | 3 | 40 | 20 | 4 | 45 | +-------------- Lepidodesmini
|
+-------------- Unionini
--+
| +-- Acuticostini
| +--------+
| | +-- Lanceolariini
+--+
| +-------- Cristariina **
+--+
| +----- Anodonta_cygnea **
+--+
| +-- Anodonta_anatina **
+--+
+-- Alasmidontina **
Wu et al., 2021 [Publ 4366], fig. 3-ML iT| tT | bi | riS | oT | roT | roS | iD | | 13 | 13 | 11 | 3 | 60 | 20 | 4 | 47 | +----------------- Lepidodesmini
|
--+ +-------------- Unionini
| |
+--+ +-- Acuticostini
| +--------+
| | +-- Lanceolariini
+--+
| +-------- Cristariina **
+--+
| +----- Anodonta_cygnea **
+--+
| +-- Anodonta_anatina **
+--+
+-- Alasmidontina **
Wu et al., 2021 [Publ 4366], fig. 3-BI iT| tT | bi | riS | oT | roT | roS | iD | | 13 | 13 | 11 | 3 | 60 | 20 | 4 | 47 | +----------------- Lepidodesmini
|
--+ +-------------- Unionini
| |
+--+ +-- Acuticostini
| +--------+
| | +-- Lanceolariini
+--+
| +-------- Cristariina **
+--+
| +----- Anodonta_cygnea **
+--+
| +-- Anodonta_anatina **
+--+
+-- Alasmidontina **
Douda et al., 2024 [Publ 4364], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 13 | 13 | 12 | 1 | 1 | 0 | 0 | 48 | Riccardi et al., 2020 [Publ 4089], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 13 | 13 | 11 | 3 | 6 | 4 | 2 | 47 | +-- Lanceolariini
+--------------+
| +-- Unionini
--+
| +-------------- Cristariina **
| |
+--+ +----------- Anodonta_anatina **
| |
+--+ +-- Anodonta_nuttalliana **
| +-----+
| | +-- Alasmidontina **
+--+
| +----- Pseudanodonta **
+--+
| +-- Anodonta_exulcerata **
+--+
+-- Anodonta_cygnea **
Sano et al., 2022 [Publ 4311], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 12 | 12 | 12 | 1 | 0 | 0 | 0 | 46 | Breton et al., 2011 [Publ 3376], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 12 | 12 | 11 | 2 | 25 | 0 | 0 | 45 | +-- Cristariina **
--+
+-- Alasmidontina **
Froufe et al., 2017 [Publ 3802], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 12 | 12 | 9 | 2 | 0 | 0 | 0 | 45 | +-- Anodontina **
--+
+-- Cristariina **
Bogan et al., 2023 [Publ 4295], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 12 | 12 | 9 | 1 | 1 | 0 | 0 | 49 | Wu et al., 2021 [Publ 4366], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 12 | 12 | 10 | 3 | 56 | 20 | 4 | 46 | +----------------- Lepidodesmini
|
--+ +-------------- Unionini
| |
+--+ +-- Acuticostini
| +--------+
| | +-- Lanceolariini
+--+
| +-------- Cristariina **
+--+
| +----- Anodonta_cygnea **
+--+
| +-- Anodonta_anatina **
+--+
+-- Alasmidontina **
Lopes-Lima et al., 2020 [Publ 4080], fig. 10 iT| tT | bi | riS | oT | roT | roS | iD | | 12 | 12 | 8 | 1 | 2 | 0 | 0 | 47 | Bolotov et al., 2020 [Publ 4084], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 12 | 12 | 10 | 2 | 24 | 7 | 2 | 41 | +-------- Unionini
--+
| +----- Lanceolariini
+--+
| +-- Anodontina **
+--+
+-- Cristariina **
Aunins et al., 2018 [Publ 3863], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 11 | 11 | 9 | 3 | 29 | 15 | 4 | 45 | +-------------- Lepidodesmini
|
--+ +----------- Unionini
| |
+--+ +-- Acuticostini
| +-----+
| | +-- Lanceolariini
+--+
| +----- Cristariina **
+--+
| +-- Anodontina **
+--+
+-- Alasmidontina **
Chen et al., 2020 [Publ 4170], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 11 | 34 | 32 | 3 | 23 | 4 | 2 | 66 | +----- Prolasmidonta **
+--+
| | +-- Alasmidonta **
| +--+
+-----------------+ +-- Anemina **
| |
| | +----- Pseudodontini
| +--+
| | +-- Lamprotula_caveata
| +--+
| +-- Chamberlainia
|
| +----- Schistodesmus
| +-----+
| | | +-- Lepidodesmini
--+ | +--+
| | +-- Aculamprotula
| +-----------+
| | | +----- Platynaias **
| | | +--+
| | | | | +-- Utterbackia_imbecillis **
| | | | +--+
| | +--+ +-- Pyganodon **
| | |
| | | +----- Sinanodonta **
| | +--+
| | | +-- Cristaria **
+--+ +--+
| +-- Anodontina **
|
| +-- Utterbackia_peninsularis **
| +-----+
| | +-- Potamilus_alatus
| +-----------+
| | | +----- Lamprotula_leaii
| | +--+
| | | +-- Sinohyriopsis
+--+ +--+
| +-- Gonideini
|
| +----------------- Gibbosula
+--+
| +-------------- Quadrulini
+--+
| +----------- Toxolasma
+--+
| +-------- Lampsilis_ornata
+--+
| +----- Potamilus_leptodon
+--+
| +-- Lampsilis_siliquoidea
+--+
+-- Lampsilis_powellii
Wu et al., 2021 [Publ 4366], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 10 | 10 | 8 | 3 | 81 | 16 | 4 | 43 | +-------------------------- Acuticostini
|
--+ +----------------------- Unionini
| |
+--+ +-------------------- Lanceolariini
| |
+--+ +----------------- Lepidodesmini
| |
+--+ +-------------- Cristariina **
| |
+--+ +----------- Anodonta_cygnea **
| |
+--+ +-- Lasmigona **
| +-----+
| | +-- Anodonta_anatina **
+--+
| +----- Pseudanodonta **
+--+
| +-- Pyganodon **
+--+
+-- Strophitus **
Wu et al., 2022 [Publ 4238], fig. 2d iT| tT | bi | riS | oT | roT | roS | iD | | 10 | 10 | 8 | 3 | 28 | 28 | 4 | 44 | +----------------- Lepidodesmini
--+
| +-------------- Unionini
+--+
| +----------- Lanceolariini
+--+
| +-------- Acuticostini
+--+
| +----- Cristariina **
+--+
| +-- Anodontina **
+--+
+-- Alasmidontina **
Krebs et al., 2003 [Publ 2691], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 10 | 10 | 5 | 2 | 23 | 0 | 0 | 45.5 | +----------- Utterbackiana **
|
+----------- Platynaias **
|
--+----------- Alasmidonta **
|
+----------- Lasmigona **
|
| +-------- Strophitus **
+--+
| +----- Utterbackia **
+--+
| +-- Pyganodon **
+--+
+-- Anodontina **
Bolotov et al., 2018 [Publ 3826], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 10 | 10 | 8 | 3 | 73 | 7 | 2 | 41 | +----------- Unionini
--+
| +-------- Lanceolariini
+--+
| +----- Cristariina **
+--+
| +-- Anodontina **
+--+
+-- Alasmidontina **
Dai et al., 2025 [Publ 4440], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 10 | 10 | 8 | 3 | 78 | 34 | 4 | 44 | +-------------- Lepidodesmini
|
--+ +----------- Unionini
| |
+--+ +-- Acuticostini
| +-----+
| | +-- Lanceolariini
+--+
| +----- Cristariina **
+--+
| +-- Anodontina **
+--+
+-- Alasmidontina **
Soroka & Burzynski, 2017 [Publ 3837], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 10 | 10 | 8 | 3 | 13 | 13 | 3 | 44 | +----- Cristariina **
+--+
| | +-- Anodontina **
| +--+
--+ +-- Alasmidontina **
|
| +----- Lanceolariini
+--+
| +-- Lepidodesmini
+--+
+-- Unionini
Dai et al., 2025 [Publ 4440], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 10 | 36 | 30 | 3 | 35 | 26 | 4 | 13 | +-- Anemina **
+-----------+
| +-- Amuranodonta **
|
| +----- Pletholophus **
--+ +-----+
| | | +-- Sinanodonta **
| | +--+
| | +-- Beringiana **
+--+
| +-- Alasmidontina **
| +-----+
| | +-- Anodontina **
| |
+--+ +----- Lanceolaria_gladiola
| |
| +----- Lanceolaria_oxyrhyncha
| |
+--+----- Cristaria **
|
| +-- Acuticostini
| |
+--+-- Lepidodesmini
|
+-- Unionini
Wu et al., 2022 [Publ 4238], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 10 | 10 | 8 | 3 | 63 | 25 | 4 | 44 | +-------------- Lepidodesmini
|
--+ +----------- Unionini
| |
+--+ +-- Acuticostini
| +-----+
| | +-- Lanceolariini
+--+
| +----- Cristariina **
+--+
| +-- Anodontina **
+--+
+-- Alasmidontina **
Huang et al., 2019 [Publ 3843], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 10 | 10 | 8 | 3 | 46 | 16 | 4 | 43 | +-------------------- Lepidodesmini
--+
| +----------------- Unionini
+--+
| +-------------- Lanceolariini
+--+
| +----------- Acuticostini
+--+
| +-------- Cristariina **
+--+
| +----- Anodonta_cygnea **
+--+
| +-- Anodonta_anatina **
+--+
+-- Alasmidontina **
Wu et al., 2021 [Publ 4366], fig. 2_I iT| tT | bi | riS | oT | roT | roS | iD | | 10 | 10 | 8 | 3 | 81 | 16 | 4 | 41 | +----------------------- Acuticostini
|
--+ +-- Lepidodesmini
| +-----------------+
| | +-- Unionini
+--+
| +----------------- Lanceolariini
+--+
| +-------------- Cristariina **
+--+
| +----------- Lasmigona **
+--+
| +-------- Anodonta **
+--+
| +----- Pseudanodonta **
+--+
| +-- Pyganodon **
+--+
+-- Strophitus **
Chen et al., 2023 [Publ 4302], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 10 | 10 | 8 | 3 | 30 | 21 | 4 | 35 | +-------------- Acuticostini
|
--+ +----------- Lepidodesmini
| |
+--+----------- Unionini
|
| +-------- Lanceolariini
+--+
| +----- Cristariina **
+--+
| +-- Alasmidontina **
+--+
+-- Anodontina **
Chen et al., 2023 [Publ 4302], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 10 | 31 | 23 | 3 | 30 | 21 | 4 | 24 | +-- Anemina **
+--------+
| +-- Amuranodonta **
|
| +----- Pletholophus **
--+ +--+
| | | +-- Sinanodonta **
| | +--+
| | +-- Beringiana **
| |
+--+ +----- Lanceolaria_gladiola
| |
| +----- Acuticostini
| |
| +----- Lepidodesmini
| |
+--+----- Lanceolaria_oxyrhyncha
|
+----- Cristaria **
|
+----- Unionini
|
| +-- Alasmidontina **
+--+
+-- Anodontina **
Sano et al., 2020 [Publ 4138], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 10 | 22 | 5 | 1 | 24 | 12 | 4 | 51 | +----- Lanceolaria_grayii__1
|
+----- Lanceolaria_grayii__2
|
+----- Cristaria_plicata__1 **
|
+----- Cristaria_plicata__2 **
|
+----- Sinanodonta_lauta **
|
+----- Beringiana_japonica **
|
+----- Beringiana_fukuharai__1 **
|
+----- Beringiana_beringiana **
|
+----- Beringiana_fukuharai__2 **
|
+----- Beringiana_fukuharai__3 **
|
+----- Buldowskia_iwakawai__1 **
|
+----- Buldowskia_iwakawai__2 **
|
+----- Sinanodonta_tumens__1 **
--+
+----- Sinanodonta_tumens__2 **
|
+----- Pletholophus **
|
+----- Anemina **
|
+----- Lanceolaria_lanceolata
|
+----- Cristaria_plicata__3 **
|
+----- Lanceolaria_gladiola
|
+----- Sinanodonta_angula **
|
+----- Lepidodesmini
|
| +-- Aculamprotula
+--+
| +-- Acuticostini
|
| +-- Cuneopsis
+--+
| +-- Nodularia
|
| +-- Schistodesmus
+--+
+-- Inversiunio
Fukata & Iigo, 2020 [Publ 4106], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 10 | 10 | 8 | 3 | 43 | 17 | 4 | 44 | +----------------- Lepidodesmini
--+
| +-------------- Lanceolariini
+--+
| +----------- Unionini
+--+
| +-------- Acuticostini
+--+
| +----- Cristariina **
+--+
| +-- Anodontina **
+--+
+-- Alasmidontina **
Xiang et al., 2025 [Publ 4433], fig. 7 iT| tT | bi | riS | oT | roT | roS | iD | | 9 | 9 | 7 | 2 | 4 | 3 | 1 | 44 | +----- Lanceolariini
--+
| +-- Anodontina **
+--+
+-- Cristariina **
Wu et al., 2019 [Publ 3864], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 9 | 9 | 7 | 3 | 37 | 17 | 4 | 43 | +-------------- Lepidodesmini
|
--+ +----------- Unionini
| |
+--+ +-- Acuticostini
| +-----+
| | +-- Lanceolariini
+--+
| +----- Cristariina **
+--+
| +-- Anodontina **
+--+
+-- Alasmidontina **
Wu et al., 2023 [Publ 4339], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 9 | 9 | 7 | 3 | 10 | 2 | 1 | 41 | +-------- Lepidodesmini
--+
| +----- Cristariina **
+--+
| +-- Alasmidontina **
+--+
+-- Anodontina **
Boyer et al., 2011 [Publ 2718], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 9 | 9 | 6 | 1 | 29 | 0 | 0 | 44 | Lopes-Lima et al., 2020 [Publ 4080], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 9 | 9 | 7 | 1 | 2 | 0 | 0 | 42 | Sano et al., 2017 [Publ 3799], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 9 | 9 | 7 | 1 | 24 | 12 | 4 | 42 | +-------------- Cristariina **
|
| +-- Tchangsinaia
--+ +--------+
| | +-- Nodularia
| |
+--+ +-- Schistodesmus
| +-----+
| | +-- Acuticostini
+--+
| +----- Inversiunio
+--+
| +-- Lepidodesmini
+--+
+-- Lanceolariini
Froufe et al., 2019 [Publ 3936], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 9 | 9 | 7 | 3 | 35 | 13 | 3 | 43 | +-------------- Lepidodesmini
--+
| +----------- Unionini
+--+
| +-------- Lanceolariini
+--+
| +----- Cristariina **
+--+
| +-- Anodontina **
+--+
+-- Alasmidontina **
Wu et al., 2020 [Publ 4121], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 9 | 9 | 7 | 3 | 40 | 18 | 4 | 43 | +-------------- Lepidodesmini
|
--+ +----------- Unionini
| |
+--+ +-- Acuticostini
| +-----+
| | +-- Lanceolariini
+--+
| +----- Cristariina **
+--+
| +-- Anodontina **
+--+
+-- Alasmidontina **
Cha et al., 2018 [Publ 3916], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 9 | 9 | 7 | 3 | 29 | 12 | 3 | 43 | +-------------- Lepidodesmini
--+
| +----------- Unionini
+--+
| +-------- Lanceolariini
+--+
| +----- Cristariina **
+--+
| +-- Anodontina **
+--+
+-- Alasmidontina **
Zieritz et al., 2021 [Publ 4231], fig. 3-F_5 iT| tT | bi | riS | oT | roT | roS | iD | | 9 | 9 | 7 | 3 | 39 | 13 | 3 | 43 | +-------------- Lepidodesmini
--+
| +----------- Unionini
+--+
| +-------- Lanceolariini
+--+
| +----- Cristariina **
+--+
| +-- Anodontina **
+--+
+-- Alasmidontina **
Froufe et al., 2019 [Publ 3936], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 9 | 9 | 7 | 3 | 35 | 13 | 3 | 43 | +-------------- Lepidodesmini
--+
| +----------- Unionini
+--+
| +-------- Lanceolariini
+--+
| +----- Cristariina **
+--+
| +-- Anodontina **
+--+
+-- Alasmidontina **
Huang et al., 2018 [Publ 3828], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 9 | 9 | 7 | 3 | 33 | 9 | 1 | 43 | +-------- Unionini
--+
| +----- Cristariina **
+--+
| +-- Anodontina **
+--+
+-- Alasmidontina **
King et al., 1999 [Publ 2699], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 9 | 9 | 7 | 1 | 0 | 0 | 0 | 46 | Lopes-Lima et al., 2017 [Publ 3792], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 9 | 9 | 7 | 3 | 23 | 9 | 2 | 43 | +-- Lanceolariini
+-----+
| +-- Unionini
--+
| +----- Cristariina **
+--+
| +-- Anodontina **
+--+
+-- Alasmidontina **
Fonseca et al., 2016 [Publ 3775], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 9 | 9 | 7 | 3 | 22 | 9 | 2 | 43 | +-- Lanceolariini
+-----+
| +-- Unionini
--+
| +----- Cristariina **
+--+
| +-- Anodontina **
+--+
+-- Alasmidontina **
Guerra et al., 2017 [Publ 3772], fig. 2b iT| tT | bi | riS | oT | roT | roS | iD | | 8 | 8 | 6 | 3 | 17 | 2 | 1 | 42 | +----------- Unio
--+
| +-------- Aculamprotula
+--+
| +----- Cristariina **
+--+
| +-- Anodontina **
+--+
+-- Alasmidontina **
Huang et al., 2019 [Publ 3843], fig. S1 iT| tT | bi | riS | oT | roT | roS | iD | | 8 | 31 | 36 | 1 | 37 | 23 | 4 | 56 | +-------------- Anemina **
|
--+ +----------- Sinanodonta **
| |
+--+ +-------- Acuticostini
| |
+--+ +-- Lepidodesmini
| +--+
| | +-- Cristaria **
+--+
| +-- Lanceolariini
+--+
+-- Unionini
Kondakov et al., 2018 [Publ 3914], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 8 | 8 | 6 | 1 | 0 | 0 | 0 | 43 | Sano et al., 2017 [Publ 3799], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 8 | 8 | 6 | 1 | 13 | 5 | 2 | 40 | +----- Cristariina **
--+
| +-- Lanceolariini
+--+
+-- Unionini
Guerra et al., 2017 [Publ 3772], fig. 2a iT| tT | bi | riS | oT | roT | roS | iD | | 8 | 8 | 6 | 3 | 17 | 2 | 1 | 40 | +----------- Unionini
--+
| +-------- Sinanodonta **
+--+
| +----- Cristaria **
+--+
| +-- Anodontina **
+--+
+-- Alasmidontina **
Huang et al., 2019 [Publ 3843], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 8 | 29 | 27 | 1 | 35 | 21 | 4 | 50 | +----------------------------------- Anemina **
|
--+ +-------------------------------- Acuticostini
| |
+--+ +----------------------------- Sinanodonta **
| |
+--+ +-------------------------- Aculamprotula
| |
+--+ +-- Cristaria **
| +--------------------+
| | +-- Lanceolariini
+--+
| +-------------------- Lepidodesmini
+--+
| +----------------- Schistodesmus
+--+
| +-------------- Nodularia
+--+
| +----------- Cuneopsis_rufescens
+--+
| +-------- Tchangsinaia
+--+
| +----- Cuneopsis_celtiformis
+--+
| +-- Cuneopsis_szechenyii
+--+
+-- Cuneopsis_heudei
Vikhrev et al., 2017 [Publ 3810], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 8 | 8 | 6 | 1 | 2 | 0 | 0 | 43 | Bespalaya et al., 2018 [Publ 3811], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 8 | 8 | 7 | 1 | 2 | 0 | 0 | 44 | Bauer et al., 2021 [Publ 4365], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 8 | 8 | 5 | 1 | 0 | 0 | 0 | 43 | Wu et al., 2024 [Publ 4388], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 7 | 5 | 3 | 47 | 11 | 3 | 42 | +-------------- Lepidodesmini
--+
| +----------- Unionini
+--+
| +-------- Lanceolariini
+--+
| +----- Cristariina **
+--+
| +-- Anodontina **
+--+
+-- Alasmidontina **
Wu et al., 2020 [Publ 4147], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 7 | 5 | 3 | 22 | 10 | 1 | 42 | +-------- Unionini
--+
| +----- Cristariina **
+--+
| +-- Anodontina **
+--+
+-- Alasmidontina **
Inoue et al., 2014 [Publ 3635], fig. 3B iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 7 | 2 | 1 | 1 | 0 | 0 | 45 | Wu et al., 2016 [Publ 3711], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 7 | 5 | 3 | 14 | 5 | 1 | 40 | +-------------- Unionini
--+
| +----------- Sinanodonta **
+--+
| +-------- Anemina **
+--+
| +----- Cristaria **
+--+
| +-- Anodontina **
+--+
+-- Alasmidontina **
Dai et al., 2023 [Publ 4285], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 7 | 5 | 3 | 46 | 10 | 4 | 42 | +-------------- Lepidodesmini
|
--+ +----------- Unionini
| |
+--+ +-- Acuticostini
| +-----+
| | +-- Lanceolariini
+--+
| +----- Cristariina **
+--+
| +-- Anodontina **
+--+
+-- Alasmidontina **
Wu et al., 2018 [Publ 4143], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 7 | 5 | 3 | 15 | 6 | 1 | 40 | +-------------- Unionini
--+
| +----------- Sinanodonta **
+--+
| +-------- Anemina **
+--+
| +----- Cristaria **
+--+
| +-- Anodontina **
+--+
+-- Alasmidontina **
Huang et al., 2002 [Publ 2657], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 14 | 5 | 2 | 27 | 7 | 4 | 44 | +-------- Strophitus **
|
+-------- Schistodesmus
|
+-------- Lepidodesmini
|
+-------- Acuticostini
|
+-------- Cristariina **
--+
+-------- Lanceolariini
|
| +-- Nodularia
+-----+
| +-- Tchangsinaia
|
| +----- Utterbackia **
| |
+--+----- Utterbackiana **
|
| +-- Alasmidonta **
+--+
+-- Lasmigona **
Huang et al., 2002 [Publ 2657], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 7 | 5 | 2 | 27 | 7 | 4 | 41 | +-- Cristariina **
+--------+
| +-- Alasmidontina **
|
--+ +----- Acuticostini
| +--+
| | | +-- Nodularia
| | +--+
+--+ +-- Tchangsinaia
|
| +----- Lanceolariini
+--+
| +-- Schistodesmus
+--+
+-- Lepidodesmini
Wang et al., 2018 [Publ 3865], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 7 | 5 | 3 | 24 | 10 | 3 | 41 | +----------- Unionini
|
--+ +-- Acuticostini
| +-----+
| | +-- Lanceolariini
+--+
| +----- Cristariina **
+--+
| +-- Anodontina **
+--+
+-- Alasmidontina **
Hoeh et al., 1995 [Publ 3391], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 7 | 4 | 1 | 0 | 0 | 0 | 45 | Smith et al., 2023 [Publ 4291], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 7 | 5 | 1 | 0 | 0 | 0 | 45 | Wang et al., 2018 [Publ 3846], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 7 | 5 | 3 | 24 | 10 | 2 | 41 | +-- Lanceolariini
+-----+
| +-- Unionini
--+
| +----- Cristariina **
+--+
| +-- Anodontina **
+--+
+-- Alasmidontina **
Guerra et al., 2019 [Publ 4168], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 7 | 5 | 3 | 34 | 6 | 2 | 41 | +-------------- Aculamprotula
|
--+ +-- Schistodesmus
| +--------+
| | +-- Unio
+--+
| +-------- Lanceolariini
+--+
| +----- Cristariina **
+--+
| +-- Anodontina **
+--+
+-- Alasmidontina **
Guerra et al., 2019 [Publ 4168], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 7 | 5 | 3 | 34 | 6 | 2 | 41 | +-- Lanceolariini
+-----+
| +-- Unionini
--+
| +----- Cristariina **
+--+
| +-- Anodontina **
+--+
+-- Alasmidontina **
Dai et al., 2024 [Publ 4358], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 7 | 7 | 5 | 3 | 45 | 20 | 4 | 41 | +----------------- Lepidodesmini
--+
| +-------------- Unionini
+--+
| +----------- Lanceolariini
+--+
| +-------- Acuticostini
+--+
| +----- Cristariina **
+--+
| +-- Anodontina **
+--+
+-- Alasmidontina **
Zhou et al., 2016 [Publ 3710], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 4 | 2 | 14 | 3 | 2 | 40 | +-- Lepidodesmini
+-----+
| +-- Unionini
--+
| +----- Sinanodonta **
+--+
| +-- Cristaria **
+--+
+-- Alasmidontina **
Boyer et al., 2011 [Publ 2718], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 2 | 1 | 29 | 0 | 0 | 41 | Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 4 | 2 | 24 | 2 | 1 | 40 | +-- Anodontina **
--+
+-- Alasmidontina **
Huang et al., 2013 [Publ 3629], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 4 | 2 | 10 | 1 | 1 | 42 | +----- Unionini
--+
| +-- Cristariina **
+--+
+-- Alasmidontina **
Pavan-Kumar et al., 2022 [Publ 4263], fig. S5 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 4 | 3 | 32 | 10 | 4 | 40 | +----- Cristariina **
+-----------+
| | +-- Anodontina **
| +--+
| +-- Alasmidontina **
--+
| +-- Acuticostini
| +-----------+
| | +-- Lanceolariini
| |
+--+ +-- Lepidodesmini
| +--------+
| | +-- Aculamprotula
+--+
| +-------- Unio
+--+
| +----- Schistodesmus
+--+
| +-- Nodularia
+--+
+-- Tchangsinaia
Hewitt et al., 2018 [Publ 3758], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 3 | 1 | 3 | 0 | 0 | 43 | Lydeard et al., 1996 [Publ 2656], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 4 | 1 | 21 | 0 | 0 | 42 | Gomes-dos-Santos et al., 2019 [Publ 4042], fig. 8 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 4 | 2 | 0 | 0 | 0 | 42 | +-- Anodontina **
--+
+-- Cristariina **
King et al., 1999 [Publ 2699], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 3 | 1 | 0 | 0 | 0 | 43 | Lopes-Lima et al., 2020 [Publ 4080], fig. 7 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 4 | 1 | 2 | 0 | 0 | 41 | Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 4 | 2 | 24 | 2 | 1 | 40 | +-- Anodontina **
--+
+-- Alasmidontina **
Tomilova et al., 2020 [Publ 4097], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 8 | 5 | 2 | 2 | 2 | 1 | 40 | +-- Unionini
|
--+-- Cristariina **
|
+-- Anodontina **
Jadhav & Jamkhedkar, 2009 [Publ 3697], fig. 11 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 8 | 6 | 2 | 19 | 3 | 1 | 40.5 | +-------------- Nodularia
|
--+ +----------- Anodonta_cygnea **
| |
+--+ +-------- Unio_pictorum__2
| |
+--+ +-- Unio_pictorum__1
| +--+
| | +-- Unio_caffer
+--+
| +-- Anodonta **
+--+
+-- Alasmidontina **
Riccardi et al., 2020 [Publ 4089], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 4 | 2 | 0 | 0 | 0 | 39 | +-- Cristariina **
--+
+-- Anodontina **
Bogan et al., 2023 [Publ 4295], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 4 | 1 | 1 | 0 | 0 | 41 | Bolotov et al., 2016 [Publ 3732], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 5 | 1 | 2 | 0 | 0 | 44 | Feng et al., 2016 [Publ 3800], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 12 | 10 | 3 | 8 | 1 | 1 | 48 | +-- Anodontina **
+--------+
| +-- Utterbackia **
|
--+ +----- Unionini
| +--+
| | | +-- Pyganodon **
| | +--+
+--+ +-- Cristariina **
|
| +----- Gonideini
+--+
| +-- Quadrulini
+--+
+-- Lampsilini
Campbell et al., 2005 [Publ 2660], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 3 | 2 | 96 | 0 | 0 | 42 | +----- Pyganodon **
|
| +-- Lasmigona **
--+--+
| +-- Pseudodontoideus **
|
| +-- Alasminota **
+--+
+-- Anodontina **
Campbell et al., 2005 [Publ 2660], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 4 | 2 | 96 | 0 | 0 | 42 | +----------- Anodonta_oregonensis **
|
--+ +-- Anodonta_cygnea **
| +-----+
| | +-- Alasminota **
+--+
| +----- Pyganodon **
+--+
| +-- Lasmigona **
+--+
+-- Pseudodontoideus **
Lydeard et al., 1996 [Publ 2656], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 4 | 2 | 22 | 0 | 0 | 40.5 | +----- Utterbackia **
+--+
| | +-- Utterbackiana **
| +--+
--+ +-- Anodontina **
|
| +----- Strophitus **
+--+
| +-- Lasmigona **
+--+
+-- Alasmidonta **
Pavan-Kumar et al., 2022 [Publ 4263], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 3 | 31 | 10 | 4 | 40 | +----------------- Lepidodesmini
--+
| +-------------- Unionini
+--+
| +----------- Lanceolariini
+--+
| +-------- Acuticostini
+--+
| +----- Cristariina **
+--+
| +-- Anodontina **
+--+
+-- Alasmidontina **
Lopes-Lima et al., 2020 [Publ 4080], fig. 8 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 1 | 2 | 0 | 0 | 40 | Pfarrer et al., 2022 [Publ 4305], fig. 2-5 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 2 | 7 | 5 | 1 | 40 | +----- Unionini
--+
| +-- Cristariina **
+--+
+-- Anodontina **
Lyubas et al., 2022 [Publ 4255], fig. S1 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 1 | 26 | 23 | 1 | 40 | +-- Anodontina **
--+
+-- Unionini
Plazzi et al., 2011 [Publ 3676], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 18 | 11 | 3 | 13 | 4 | 2 | 50 | +----------- Cristariina **
|
| +-------- Lanceolariini
| |
--+ +-------- Anodonta **
| |
| +-------- Unionini
| |
| | +-- Alasmidontina **
+--+-----+
| +-- Pseudanodonta **
|
| +-- Alathyria
| |
| +--+-- Lamprotulini
| | |
| | +-- Chamberlainiini
+--+
| +-- Quadrulini
| |
+--+-- Pleurobemini
|
+-- Lampsilini
Bogan et al., 2023 [Publ 4295], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 1 | 1 | 0 | 0 | 40 | Manendo et al., 2008 [Publ 2669], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 2 | 8 | 3 | 1 | 41 | +-------- Unionini
--+
| +----- Anodonta **
+--+
| +-- Pseudanodonta **
+--+
+-- Alasmidontina **
Soroka, 2010 [Publ 3404], fig. 9a iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 1 | 0 | 0 | 0 | 40 | Pfeiffer & Graf, 2013 [Publ 3531], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 2 | 26 | 6 | 2 | 41 | +-- Cristariina **
+--+
| +-- Alasmidontina **
--+
| +-- Acuticostini
+--+
+-- Unionini
Mabuchi & Nishida, 2020 [Publ 4120], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 5 | 1 | 7 | 3 | 2 | 42 | +-------- Cristariina **
--+
| +----- Nodularia
+--+
| +-- Lanceolariini
+--+
+-- Inversiunio
Xue et al., 2016 [Publ 3659], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 2 | 10 | 6 | 2 | 39 | +-- Lanceolariini
+-----+
| +-- Unionini
|
--+ +-- Cristaria **
| +--+
| | +-- Alasmidontina **
+--+
| +-- Anemina **
+--+
+-- Sinanodonta **
Nagel et al., 1998 [Publ 1946], fig. 7 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 4 | 2 | 0 | 0 | 0 | 40 | +-------------- Pseudanodonta **
--+
| +----------- Anodonta_cygnea **
+--+
| +-------- Cristariina **
+--+
| +----- Anodonta__2 **
+--+
| +-- Anodonta_anatina **
+--+
+-- Anodonta__1 **
Nagel et al., 1996 [Publ 3400], fig. 351 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 4 | 2 | 0 | 0 | 0 | 40 | +-------------- Pseudanodonta **
--+
| +----------- Anodonta_cygnea **
+--+
| +-------- Cristariina **
+--+
| +----- Anodonta__2 **
+--+
| +-- Anodonta_anatina **
+--+
+-- Anodonta__1 **
Soroka, 2010 [Publ 3404], fig. 5a iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 2 | 3 | 3 | 1 | 40 | +----------- Unionini
--+
| +-------- Anodonta_anatina **
+--+
| +----- Cristariina **
+--+
| +-- Anodonta_cygnea **
+--+
+-- Pseudanodonta **
Amaro et al., 2017 [Publ 3791], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 1 | 11 | 0 | 0 | 42 | Soroka, 2010 [Publ 3404], fig. 3a iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 9 | 7 | 2 | 4 | 4 | 1 | 40 | +----- Cristariina **
--+
| +-- Anodontina **
+--+
+-- Unionini
Lyubas et al., 2022 [Publ 4255], fig. 7 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 3 | 1 | 26 | 23 | 1 | 40 | +-- Anodontina **
--+
+-- Unionini
Soroka, 2010 [Publ 3404], fig. 3b iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 9 | 5 | 2 | 4 | 4 | 1 | 43 | +----- Anodonta_anatina **
|
+----- Anodonta_cygnea **
--+
+----- Pseudanodonta **
|
| +-- Cristariina **
+--+
+-- Unionini
Soroka, 2010 [Publ 3404], fig. 5b iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 8 | 5 | 2 | 3 | 3 | 1 | 42 | +-------- Anodonta_anatina **
|
--+ +----- Cristariina **
| |
+--+----- Unionini
|
| +-- Anodonta_cygnea **
+--+
+-- Pseudanodonta **
Cyr et al., 2007 [Publ 2745], fig. 2B iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 4 | 2 | 0 | 0 | 0 | 42 | +-- Anodontina **
--+
+-- Alasmidontina **
Cyr et al., 2007 [Publ 2745], fig. 2C iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 4 | 2 | 0 | 0 | 0 | 42 | +-- Anodontina **
--+
+-- Alasmidontina **
Froufe et al., 2019 [Publ 3936], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 3 | 15 | 3 | 1 | 38 | +-------- Unionini
--+
| +----- Cristariina **
+--+
| +-- Anodontina **
+--+
+-- Alasmidontina **
Guerra et al., 2019 [Publ 4168], fig. 2-M iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 3 | 18 | 6 | 2 | 40 | +-- Lanceolariini
+-----+
| +-- Unionini
--+
| +----- Alasmidontina **
+--+
| +-- Anodontina **
+--+
+-- Cristariina **
Sayenko et al., 2017 [Publ 3760], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 1 | 0 | 0 | 0 | 39 | Soroka, 2010 [Publ 3404], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 7 | 5 | 2 | 3 | 3 | 1 | 42 | +-- Anodonta_anatina **
+-----+
| +-- Unionini
--+
| +----- Cristariina **
+--+
| +-- Anodonta_cygnea **
+--+
+-- Pseudanodonta **
Wu et al., 2022 [Publ 4238], fig. 2e iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 3 | 13 | 13 | 2 | 38 | +-- Lanceolariini
+-----+
| +-- Unionini
--+
| +----- Cristariina **
+--+
| +-- Anodontina **
+--+
+-- Alasmidontina **
Wu et al., 2023 [Publ 4296], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 2 | 19 | 17 | 3 | 39 | +----------- Acuticostini
--+
| +-------- Lanceolariini
+--+
| +----- Unionini
+--+
| +-- Anodontina **
+--+
+-- Alasmidontina **
Bulakhova et al., 2023 [Publ 4325], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 1 | 0 | 0 | 0 | 41 | Liu et al., 2023 [Publ 4310], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 2 | 20 | 18 | 3 | 39 | +-------------- Acuticostini
|
--+ +-- Lanceolaria_lanceolata
| +--------+
| | +-- Lanceolaria_grayii
+--+
| +-------- Lanceolaria_gladiola
+--+
| +----- Unionini
+--+
| +-- Anodontina **
+--+
+-- Alasmidontina **
Hoeh et al., 2009 [Publ 2716], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 1 | 42 | 2 | 1 | 39 | +-- Unionini
--+
+-- Alasmidontina **
Hoeh et al., 2009 [Publ 2716], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 1 | 43 | 2 | 1 | 39 | +-- Unionini
--+
+-- Alasmidontina **
Cyr et al., 2007 [Publ 2745], fig. 2D iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 1 | 0 | 0 | 0 | 42 | Soroka, 2010 [Publ 3404], fig. 9c iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 3 | 1 | 0 | 0 | 0 | 42 | Rosenberg et al., 1997 [Publ 2720], fig. 2A iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 18 | 1 | 2 | 18 | 1 | 1 | 54 | +----- Anodontina **
|
| +-- Gonideini
| |
| +-- Pyganodon_grandis **
| |
| +-- Utterbackia **
| |
| +-- Pyganodon_cataracta **
--+ |
| +-- Lampsilis_straminea
| |
| +-- Lampsilis_teres
| |
| +-- Megalonaias
| |
| +-- Obliquaria
| |
+--+-- Theliderma
|
+-- Quadrula
|
+-- Uniomerus
|
+-- Amblema
|
+-- Elliptio
|
+-- Fusconaia
|
+-- Plectomerus
|
+-- Pleurobema
|
+-- Unionini
Lopes-Lima et al., 2021 [Publ 4217], fig. 11 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 1 | 0 | 0 | 0 | 41 | Huang et al., 2018 [Publ 3828], fig. 6-M iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 3 | 13 | 4 | 1 | 38 | +-------- Unionini
--+
| +----- Cristariina **
+--+
| +-- Anodontina **
+--+
+-- Alasmidontina **
Mezhzherin et al., 2014 [Publ 3705], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 2 | 0 | 0 | 0 | 37 | +-- Cristariina **
--+
+-- Anodontina **
Klishko et al., 2014 [Publ 3666], fig. 8 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 1 | 0 | 0 | 0 | 37 | Huang et al., 2019 [Publ 3843], fig. 3-M iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 3 | 19 | 6 | 2 | 38 | +-- Lanceolariini
+-----+
| +-- Unionini
--+
| +----- Cristariina **
+--+
| +-- Anodontina **
+--+
+-- Alasmidontina **
Cyr et al., 2007 [Publ 2745], fig. 2A iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 1 | 1 | 0 | 0 | 42 | Klishko et al., 2016 [Publ 3733], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 1 | 0 | 0 | 0 | 37 | Burzynski & Soroka, 2018 [Publ 3862], fig. 5a iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 1 | 3 | 16 | 6 | 2 | 39 | +-- Lanceolariini
+--+
| +-- Unionini
--+
| +-- Cristariina **
| |
+--+-- Anodontina **
|
+-- Alasmidontina **
Burzynski & Soroka, 2018 [Publ 3862], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 3 | 0 | 0 | 0 | 38 | +----- Cristariina **
--+
| +-- Anodontina **
+--+
+-- Alasmidontina **
Soroka, 2010 [Publ 3404], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 7 | 4 | 2 | 3 | 3 | 1 | 38 | +----- Cristariina **
--+
| +-- Unionini
+--+
+-- Anodontina **
Froufe et al., 2017 [Publ 3802], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 2 | 0 | 0 | 0 | 39 | +-- Cristariina **
--+
+-- Anodontina **
Lopes-Lima et al., 2020 [Publ 4080], fig. 9 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 1 | 2 | 0 | 0 | 38 | Hoeh et al., 2009 [Publ 2716], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 1 | 42 | 2 | 1 | 39 | +-- Unionini
--+
+-- Alasmidontina **
Amaro et al., 2017 [Publ 3791], fig. 3B iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 1 | 8 | 0 | 0 | 42 | Zieritz et al., 2012 [Publ 3673], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 8 | 4 | 2 | 4 | 3 | 1 | 39 | +----- Anodontina **
|
--+ +-- Cristariina **
| |
+--+-- Margaritifera
|
+-- Unionini
Froufe et al., 2019 [Publ 3936], fig. 3-M iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 3 | 15 | 3 | 1 | 38 | +-------- Unionini
--+
| +----- Cristariina **
+--+
| +-- Anodontina **
+--+
+-- Alasmidontina **
Amaro et al., 2017 [Publ 3791], fig. 2-M iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 1 | 2 | 6 | 0 | 0 | 39 | +----- Pyganodon **
|
--+ +-- Lasmigona **
| |
+--+-- Cristariina **
|
+-- Utterbackia **
Liu et al., 2023 [Publ 4310], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 2 | 20 | 18 | 3 | 39 | +----------------- Acuticostini
|
--+ +-------------- Lanceolaria_grayii
| |
| +-------------- Lanceolaria_gladiola
+--+
+-------------- Lanceolaria_lanceolata
|
| +----------- Unionini
+--+
| +-------- Lasmigona **
+--+
| +----- Anodontina **
+--+
| +-- Pyganodon **
+--+
+-- Strophitus **
Froufe et al., 2017 [Publ 3802], fig. 7 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 2 | 6 | 3 | 1 | 39 | +----- Unionini
--+
| +-- Cristariina **
+--+
+-- Anodontina **
Guerra et al., 2019 [Publ 4168], fig. 1-M iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 3 | 18 | 6 | 2 | 38 | +-- Lanceolariini
+-----+
| +-- Unionini
--+
| +----- Cristariina **
+--+
| +-- Anodontina **
+--+
+-- Alasmidontina **
Soroka, 2010 [Publ 3404], fig. 9b iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 1 | 0 | 0 | 0 | 41 | Zieritz et al., 2021 [Publ 4231], fig. 3-M_4 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 3 | 19 | 3 | 1 | 38 | +-------- Unionini
--+
| +----- Cristariina **
+--+
| +-- Anodontina **
+--+
+-- Alasmidontina **
Rosenberg et al., 1997 [Publ 2720], fig. 2B iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 18 | 1 | 2 | 18 | 2 | 1 | 53.5 | +----- Anodontina **
|
| +-- Gonideini
| |
| +-- Pyganodon_grandis **
| |
| +-- Utterbackia **
| |
| +-- Pyganodon_cataracta **
--+ |
| +-- Lampsilis_straminea
| |
| +-- Lampsilis_teres
| |
| +-- Megalonaias
| |
| +-- Obliquaria
| |
+--+-- Theliderma
|
+-- Quadrula
|
+-- Uniomerus
|
+-- Amblema
|
+-- Elliptio
|
+-- Fusconaia
|
+-- Plectomerus
|
+-- Aculamprotula
|
+-- Unio
Pfeiffer et al., 2019 [Publ 3937], fig. 4-5 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 3 | 53 | 4 | 2 | 39 | +----------- Lanceolariini
--+
| +-------- Unionini
+--+
| +----- Cristariina **
+--+
| +-- Anodontina **
+--+
+-- Alasmidontina **
Mezhzherin et al., 2014 [Publ 3705], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 2 | 0 | 0 | 0 | 37 | +-- Cristariina **
--+
+-- Anodontina **
Pfeiffer et al., 2019 [Publ 3937], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 3 | 53 | 4 | 2 | 39 | +-- Lanceolariini
+-----+
| +-- Unionini
--+
| +----- Cristariina **
+--+
| +-- Anodontina **
+--+
+-- Alasmidontina **
Graf & Cummings, 2006 [Publ 1968], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 1 | 43 | 2 | 1 | 39 | +-- Unionini
--+
+-- Alasmidontina **
Amaro et al., 2017 [Publ 3791], fig. 3A iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 1 | 8 | 0 | 0 | 42 | Burzynski & Soroka, 2018 [Publ 3862], fig. 5b iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 3 | 6 | 6 | 2 | 38 | +----------- Lanceolariini
--+
| +-------- Unionini
+--+
| +----- Cristariina **
+--+
| +-- Anodontina **
+--+
+-- Alasmidontina **
Walker et al., 2006 [Publ 2666], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 2 | 12 | 0 | 0 | 39.5 | +-- Anodontina **
--+
+-- Alasmidontina **
Kano et al., 2019 [Publ 4194], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 7 | 3 | 2 | 41 | Lopes-Lima et al., 2016 [Publ 3757], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 0 | 0 | 0 | 38 | Moyer & Díaz-Ferguson, 2012 [Publ 3375], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 3 | 0 | 0 | 40 | Soroka, 2010 [Publ 3404], fig. 8a iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 2 | 3 | 3 | 1 | 38 | +----- Unionini
--+
| +-- Cristariina **
+--+
+-- Anodontina **
Soroka, 2008 [Publ 3695], fig. 1-M iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 1 | 0 | 0 | 37 | +----- Cristariina **
--+
| +-- Anodontina **
+--+
+-- Alasmidontina **
Lopes-Lima et al., 2017 [Publ 3792], fig. 2-M iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 2 | 7 | 1 | 1 | 39 | +----- Unionini
--+
| +-- Anodontina **
+--+
+-- Alasmidontina **
Hoeh et al., 2001 [Publ 2652], fig. 14f2 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 2 | 28 | 2 | 1 | 40 | +----- Pyganodon **
--+
| +-- Anodontina **
+--+
+-- Strophitus **
Froufe et al., 2015 [Publ 3664], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 2 | 9 | 0 | 0 | 39 | +-- Anodontina **
--+
+-- Alasmidontina **
Chase et al., 2018 [Publ 3861], fig. 1-M iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 2 | 8 | 2 | 1 | 39 | +----- Unionini
--+
| +-- Anodontina **
+--+
+-- Alasmidontina **
Araujo et al., 2018 [Publ 3779], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 2 | 15 | 14 | 1 | 38 | +----- Unionini
--+
| +-- Anodontina **
+--+
+-- Alasmidontina **
Bogan & Hoeh, 2000 [Publ 2650], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 2 | 31 | 2 | 1 | 40 | +-- Anodontina **
--+
+-- Alasmidontina **
Reis et al., 2013 [Publ 3580], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 0 | 1 | 12 | 7 | 1 | 38 | +-- Anodontina **
--+
+-- Unionini
Nagel et al., 1998 [Publ 1946], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 6 | 4 | 1 | 38 | +-- Anodontina **
--+
+-- Unionini
Hoeh et al., 2002 [Publ 2653], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 2 | 28 | 2 | 1 | 40 | +-------- Unio_pictorum
--+
| +----- Unio_tumidus
+--+
| +-- Anodontina **
+--+
+-- Alasmidontina **
Chase et al., 2018 [Publ 3861], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 2 | 8 | 2 | 1 | 39 | +----- Unionini
--+
| +-- Anodontina **
+--+
+-- Alasmidontina **
Gustafson & Iwamoto, 2005 [Publ 2741], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 2 | 2 | 0 | 0 | 39 | +-- Cristariina **
--+
+-- Anodontina **
Dai et al., 2024 [Publ 4386], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 2 | 37 | 23 | 4 | 39 | +-------------- Lepidodesmini
--+
| +----------- Unionini
+--+
| +-------- Lanceolariini
+--+
| +----- Acuticostini
+--+
| +-- Cristariina **
+--+
+-- Alasmidontina **
Ouyang et al., 2011 [Publ 3408], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 10 | 6 | 3 | 39 | +----------- Cristariina **
|
--+ +-- Nodularia
| +-----+
| | +-- Cuneopsis
+--+
| +----- Lanceolaria_grayii
| |
+--+----- Acuticostini
|
| +-- Lanceolaria_gladiola
+--+
+-- Aculamprotula
Burzynski et al., 2017 [Publ 3774], fig. 1-top iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 2 | 10 | 3 | 1 | 39 | +----- Unionini
--+
| +-- Anodontina **
+--+
+-- Alasmidontina **
Soroka, 2010 [Publ 3404], fig. 3a-M iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 6 | 4 | 2 | 3 | 3 | 1 | 41 | +----- Anodontina **
--+
| +-- Cristariina **
+--+
+-- Unionini
Krebs et al., 2019 [Publ 4092], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 0 | 0 | 0 | 41 | Wu et al., 2024 [Publ 4379], fig. 4_6 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 19 | 14 | 1 | 38 | +-------- Aculamprotula
--+
| +----- Cristariina **
+--+
| +-- Schistodesmus
+--+
+-- Nodularia
Reis et al., 2013 [Publ 3580], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 11 | 6 | 1 | 38 | +-- Anodontina **
--+
+-- Unionini
Prié & Puillandre, 2014 [Publ 3529], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 14 | 11 | 1 | 38 | +-- Anodontina **
--+
+-- Unionini
Rosenberg et al., 1994 [Publ 2719], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 10 | 0 | 1 | 16 | 1 | 1 | 48 | +-- Pyganodon_cataracta **
|
+-- Pyganodon_grandis **
|
+-- Utterbackia **
|
+-- Lampsilis_teres
|
+-- Lampsilis_straminea
--+
+-- Megalonaias
|
+-- Obliquaria
|
+-- Theliderma
|
+-- Quadrula
|
+-- Uniomerus
Fonseca et al., 2016 [Publ 3775], fig. 1-M iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 2 | 6 | 1 | 1 | 39 | +----- Unionini
--+
| +-- Anodontina **
+--+
+-- Alasmidontina **
Soroka, 2008 [Publ 3695], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 1 | 0 | 0 | 37 | +----- Cristariina **
--+
| +-- Anodontina **
+--+
+-- Alasmidontina **
Walker et al., 2006 [Publ 2666], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 2 | 12 | 0 | 0 | 39.5 | +-- Anodontina **
--+
+-- Alasmidontina **
Soroka, 2010 [Publ 3404], fig. 8b iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 6 | 4 | 2 | 3 | 3 | 1 | 39 | +----- Cristariina **
--+
| +-- Anodontina **
+--+
+-- Unionini
Roe & Hoeh, 2003 [Publ 2655], fig. 4f3 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 2 | 28 | 2 | 1 | 40 | +----- Unionini
--+
| +-- Anodontina **
+--+
+-- Alasmidontina **
Froufe et al., 2014 [Publ 3592], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 0 | 0 | 0 | 38 | Froufe et al., 2015 [Publ 3664], fig. 1-M iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 2 | 5 | 0 | 0 | 39 | +-- Anodontina **
--+
+-- Alasmidontina **
Roe & Hoeh, 2003 [Publ 2655], fig. 4f2 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 2 | 28 | 2 | 1 | 40 | +-------- Unionini
--+
| +----- Pyganodon **
+--+
| +-- Anodontina **
+--+
+-- Strophitus **
Hoeh et al., 2001 [Publ 2652], fig. 14f4 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 2 | 28 | 2 | 1 | 40 | +----- Unio_pictorum
|
--+----- Unio_tumidus
|
| +-- Anodontina **
+--+
+-- Alasmidontina **
Burzynski et al., 2017 [Publ 3774], fig. 1-bottom iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 2 | 10 | 3 | 1 | 39 | +----- Unionini
--+
| +-- Anodontina **
+--+
+-- Alasmidontina **
Soroka & Burzynski, 2018 [Publ 3762], fig. 3-M iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 2 | 4 | 2 | 1 | 39 | +----- Unionini
--+
| +-- Anodontina **
+--+
+-- Alasmidontina **
Dai et al., 2024 [Publ 4389], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 44 | 0 | 0 | 38 | Källersjö et al., 2005 [Publ 2684], fig. 1A iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 0 | 1 | 4 | 3 | 1 | 38 | Smith, 2021 [Publ 4331], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 2 | 17 | 0 | 0 | 39 | +-- Cristariina **
--+
+-- Alasmidontina **
Källersjö et al., 2005 [Publ 2684], fig. 1B-F iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 4 | 3 | 1 | 38 | Soroka & Burzynski, 2018 [Publ 3762], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 2 | 4 | 2 | 1 | 39 | +----- Unionini
--+
| +-- Anodontina **
+--+
+-- Alasmidontina **
Soroka, 2010 [Publ 3404], fig. 3b-M iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 6 | 4 | 2 | 3 | 3 | 1 | 41 | +----- Anodontina **
--+
| +-- Cristariina **
+--+
+-- Unionini
Hinzmann et al., 2013 [Publ 3541], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 0 | 0 | 0 | 38 | Wen et al., 2017 [Publ 3795], fig. 3-M iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 2 | 3 | 0 | 0 | 39 | +-- Anodontina **
--+
+-- Alasmidontina **
Lopes-Lima et al., 2021 [Publ 4217], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 22 | 16 | 1 | 40 | +-- Anodontina **
--+
+-- Unionini
Dai et al., 2025 [Publ 4441], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 45 | 0 | 0 | 38 | Lopes-Lima et al., 2017 [Publ 3734], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 1 | 14 | 9 | 1 | 38 | +-- Anodontina **
--+
+-- Unionini
Wen et al., 2017 [Publ 3795], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 2 | 13 | 4 | 2 | 38 | +-- Anodontina **
+-----+
| +-- Cristariina **
--+
| +----- Aculamprotula
+--+
| +-- Lepidodesmini
+--+
+-- Unio
Walker et al., 2006 [Publ 2666], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 2 | 12 | 0 | 0 | 39.5 | +-- Anodontina **
--+
+-- Alasmidontina **
Lopes-Lima et al., 2019 [Publ 3901], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 26 | 0 | 0 | -- | +-- Anodontina **
--+
+-- Alasmidontina **
Nagel et al., 1998 [Publ 1946], fig. 9 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 4 | 1 | 1 | -- | +-- Unionini
--+
+-- Anodontina **
Chase et al., 2018 [Publ 3847], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 8 | 2 | 1 | -- | +----- Unionini
--+
| +-- Anodontina **
+--+
+-- Alasmidontina **
Soroka, 2010 [Publ 3404], fig. 5a-M iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 0 | 0 | 0 | -- | +-- Anodontina **
--+
+-- Cristariina **
Lopes-Lima et al., 2024 [Publ 4376], fig. 4-5_MtG iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 20 | 20 | 2 | -- | +----- Unionini
--+
| +-- Lanceolariini
+--+
+-- Anodontina **
Zhou et al., 2007 [Publ 2658], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 18 | 0 | 1 | 16 | 6 | 2 | 45 | +----------------- Sinanodonta **
|
--+ +-------------- Cristaria **
| |
+--+ +-- Lamprotulini
| +--------+
| | +-- Quadrulini
+--+
| +-------- Aculamprotula
+--+
| +----- Acuticostini
+--+
| +-- Nodularia
+--+
+-- Cuneopsis
Zhou et al., 2007 [Publ 2658], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 19 | 11 | 3 | -- | Huang et al., 2013 [Publ 3629], fig. 3-M iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 4 | 0 | 0 | -- | Dai et al., 2024 [Publ 4386], fig. 5-ML iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 16 | 0 | 2 | 19 | 14 | 4 | 47 | +----------- Anodontina **
|
--+ +-- Acuticostini
| +-----+
| | +-- Cristariina **
+--+
| +----- Lepidodesmini
+--+
| +-- Lanceolariini
+--+
+-- Unionini
Lopes-Lima et al., 2024 [Publ 4376], fig. 4-5_AHE iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 20 | 20 | 2 | -- | +----- Unionini
--+
| +-- Lanceolariini
+--+
+-- Anodontina **
Lopes-Lima et al., 2019 [Publ 3901], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 24 | 0 | 0 | -- | +-- Anodontina **
--+
+-- Alasmidontina **
Guerra et al., 2017 [Publ 3772], fig. 2a-M iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 6 | 0 | 0 | -- | Soroka, 2010 [Publ 3404], fig. 4-M iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 0 | 0 | 0 | -- | +-- Anodontina **
--+
+-- Cristariina **
Walker et al., 2006 [Publ 2666], fig. 6-M iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 12 | 0 | 0 | -- | Hoeh et al., 2002 [Publ 2673], fig. 1-F iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 7 | 0 | 0 | -- | Amaro et al., 2017 [Publ 3791], fig. 3A-M iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 5 | 0 | 0 | -- | Amaro et al., 2017 [Publ 3791], fig. 3B-M iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 5 | 0 | 0 | -- | Mock et al., 2004 [Publ 2686], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 0 | 0 | 0 | -- | +-- Anodontina **
--+
+-- Cristariina **
Hoeh et al., 1996 [Publ 2694], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 2 | 0 | 0 | -- | Zhang et al., 2004 [Publ 2735], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 12 | 10 | 3 | -- | +-------- Cristariina **
--+
| +----- Unionini
+--+
| +-- Acuticostini
+--+
+-- Lanceolariini
Dai et al., 2024 [Publ 4386], fig. 5-BI iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 16 | 0 | 2 | 19 | 14 | 4 | 45 | +----------------- Acuticostini
|
+----------------- Cristariina **
|
+----------------- Anodontina **
--+
| +-------------- Lanceolariini
| |
| | +-- Aculamprotula
| +-----------+
+--+ +-- Lepidodesmini
|
| +----------- Unio
| |
+--+ +-------- Middendorffinaia
| |
+--+ +----- Cuneopsis
| |
+--+----- Nodularia
|
| +-- Inversiunio
+--+
+-- Schistodesmus
Hoeh et al., 2002 [Publ 2673], fig. 1-M iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 7 | 0 | 0 | -- | Soroka, 2010 [Publ 3404], fig. 5b-M iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 0 | 0 | 0 | -- | +-- Anodontina **
--+
+-- Cristariina **
Sheng et al., 2014 [Publ 3704], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 8 | 0 | 2 | 6 | 0 | 0 | 44 | +-------------- Cristariina **
|
--+ +----------- Alasmidontina **
| |
+--+ +-------- Lampsilis
| |
+--+ +-- Venustaconcha
| +--+
| | +-- Quadrulini
+--+
| +-- Lamprotulini
+--+
+-- Chamberlainiini
Klishko et al., 2019 [Publ 3886], fig. 9 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 9 | 8 | 1 | -- | +-- Anodontina **
--+
+-- Unionini
Graf, 2002 [Publ 2651], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 16 | 2 | 1 | -- | +-- Unionini
--+
+-- Alasmidontina **
Bogatov et al., 2018 [Publ 3859], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 4 | 3 | 1 | -- | +-------- Unio_crassus
--+
| +----- Unio_pictorum
+--+
| +-- Unio_tumidus
+--+
+-- Anodontina **
Bogatov et al., 2018 [Publ 3859], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 4 | 3 | 1 | -- | +-------- Unio_tumidus
--+
| +----- Anodontina **
+--+
| +-- Unio_crassus
+--+
+-- Unio_pictorum
Krebs, 2004 [Publ 2692], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 3 | 0 | 0 | -- | Whelan et al., 2011 [Publ 3380], fig. 1-MP iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 34 | 2 | 1 | -- | +-- Alasmidontina **
--+
+-- Unionini
Mock et al., 2004 [Publ 2686], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 0 | 0 | 0 | -- | +-- Anodontina **
--+
+-- Cristariina **
Wei et al., 2016 [Publ 3665], fig. 0 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 4 | 0 | 2 | 4 | 0 | 0 | 40 | +-- Cristariina **
|
+-- Lampsilini
--+
+-- Alasmidontina **
|
+-- Quadrulini
Whelan et al., 2011 [Publ 3380], fig. 1-ML iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 34 | 2 | 1 | -- | +-- Alasmidontina **
--+
+-- Unionini
Pfeiffer & Graf, 2015 [Publ 3620], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 38 | 1 | 1 | -- | +-- Unionini
--+
+-- Alasmidontina **
Chen et al., 2012 [Publ 3624], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 5 | 0 | 0 | -- | +-- Alasmidontina **
--+
+-- Cristariina **
Walker et al., 2006 [Publ 2666], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 34 | 0 | 0 | -- | Zhang et al., 2013 [Publ 4214], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 4 | 0 | 1 | 2 | 0 | 0 | 40 | +-------- Cristaria **
--+
| +----- Sinanodonta **
+--+
| +-- Margaritifera
+--+
+-- Lamprotulini
Soroka, 2010 [Publ 3404], fig. 2-M iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 5 | 0 | 2 | 3 | 3 | 1 | 38 | +----- Cristariina **
--+
| +-- Anodontina **
+--+
+-- Unionini
Krebs, 2004 [Publ 2692], fig. 2-M iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 2 | 0 | 0 | -- | Curole & Kocher, 2005 [Publ 2682], fig. 1-M iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 10 | 0 | 0 | -- | Dyduch-Falniowska & Koziol, 1989 [Publ 3814], fig. 5_7 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 3 | 3 | 1 | -- | +-- Anodontina **
--+
+-- Unionini
Xia et al., 2016 [Publ 3712], fig. 7B iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 3 | 0 | 1 | 1 | 0 | 0 | 38 | +----- Sinanodonta **
--+
| +-- Cristaria **
+--+
+-- Chamberlainiini
Guerra et al., 2017 [Publ 3772], fig. 2b-M iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 6 | 0 | 0 | -- | Lopes-Lima et al., 2019 [Publ 3901], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 27 | 0 | 0 | -- | +-- Anodontina **
--+
+-- Alasmidontina **
Huff et al., 2004 [Publ 2697], fig. 5-6 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 11 | 1 | 1 | -- | Graf & Ó Foighil, 2000 [Publ 2649], fig. 1_2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 10 | 1 | 1 | -- | +-- Unionini
--+
+-- Alasmidontina **
Hewitt et al., 2019 [Publ 3962], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 9 | 0 | 0 | -- | Inoue et al., 2013 [Publ 3528], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 16 | 0 | 0 | -- | Araujo et al., 2009 [Publ 2589], fig. 7B iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 10 | 7 | 1 | -- | +-- Anodontina **
--+
+-- Unionini
Araujo et al., 2017 [Publ 3801], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 13 | 3 | 1 | -- | +-- Schistodesmus
+--+
| +-- Aculamprotula
--+
| +-- Anodontina **
+--+
+-- Unio
Lydeard et al., 2000 [Publ 2654], fig. 2d iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 13 | 0 | 0 | -- | Sonowal et al., 2021 [Publ 4221], fig. 3a iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 14 | 3 | 1 | -- | +-- Anodontina **
|
+-- Unio_ravoisieri
--+
+-- Unio_pictorum
|
+-- Aculamprotula
Taylor et al., 2007 [Publ 2696], fig. 1-3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 3 | 1 | 1 | -- | +-- Anodontina **
--+
+-- Unionini
Zanatta & Murphy, 2006 [Publ 2672], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 56 | 0 | 0 | -- | Zanatta & Murphy, 2006 [Publ 2672], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 55 | 0 | 0 | -- | Sonowal et al., 2021 [Publ 4221], fig. 3b iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 14 | 3 | 1 | -- | Peng et al., 2012 [Publ 3689], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 2 | 0 | 0 | -- | Pfeiffer et al., 2016 [Publ 3660], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 37 | 0 | 0 | -- | -- Utterbackiana_suborbiculata **
Lydeard et al., 2000 [Publ 2654], fig. 2c iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 13 | 0 | 0 | -- | Graf, 2000 [Publ 2647], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 16 | 1 | 1 | -- | Gomes-dos-Santos et al., 2019 [Publ 3943], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 9 | 1 | 1 | -- | +-- Unionini
--+
+-- Anodontina **
Huff et al., 2004 [Publ 2697], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 7 | 0 | 0 | -- | Bolotov et al., 2016 [Publ 3743], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 29 | 2 | 1 | -- | +-- Anodontina **
--+
+-- Unionini
Pieri et al., 2018 [Publ 3867], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 19 | 0 | 0 | -- | -- Arcidens_confragosus **
Huff et al., 2004 [Publ 2697], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 11 | 1 | 1 | -- | +-- Unionini
--+
+-- Anodontina **
Zhou et al., 2017 [Publ 3788], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 15 | 5 | 2 | -- | -- Anemina_arcaeformis **
Plazzi & Passamonti, 2010 [Publ 3354], fig. 2-3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 3 | 0 | 0 | -- | -- Sinanodonta_woodiana **
Campbell, 2000 [Publ 2724], fig. 1-5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 2 | 0 | 0 | -- | -- Utterbackia_imbecillis **
Sonowal et al., 2021 [Publ 4221], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 15 | 3 | 1 | -- | Soroka, 2010 [Publ 3404], fig. 8b-M iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 2 | 2 | 1 | -- | +-- Cristariina **
--+
+-- Unionini
Dai et al., 2024 [Publ 4386], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 12 | 11 | 2 | -- | +-------------- Unio
|
| +----- Aculamprotula
--+ +-----+
| | | +-- Anodontina **
| | +--+
+--+ +-- Lepidodesmini
|
| +-------- Middendorffinaia
+--+
| +----- Inversiunio
+--+
| +-- Cuneopsis
+--+
+-- Nodularia
Liu et al., 2017 [Publ 3939], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 14 | 13 | 3 | -- | -- Anemina_arcaeformis **
Araujo et al., 2009 [Publ 2589], fig. 7A iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 10 | 7 | 1 | -- | +----------- Nodularia
|
| +-- Unio_gibbus
--+ |
| +-----+-- Unio_tumidus
| | |
| | +-- Anodontina **
+--+
| +----- Unio_pictorum__1
| |
| +----- Unio_pictorum__2
+--+
+----- Unio_mancus
|
| +-- Unio_tumidiformis
+--+
+-- Unio_crassus
Inoue et al., 2020 [Publ 3941], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 41 | 0 | 0 | -- | -- Arcidens_confragosus **
Hoeh et al., 1998 [Publ 2695], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 4 | 0 | 0 | -- | Araujo et al., 2009 [Publ 2589], fig. 7C iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 9 | 6 | 1 | -- | +-- Anodontina **
--+
+-- Unionini
Mulvey et al., 1997 [Publ 3390], fig. 1_3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 4 | 0 | 0 | -- | -- Utterbackia_imbecillis **
Turner et al., 2000 [Publ 2722], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 13 | 0 | 0 | -- | -- Utterbackiana_couperiana **
Lydeard et al., 2000 [Publ 2654], fig. 2a iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 13 | 0 | 0 | -- | Huff et al., 2004 [Publ 2697], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 10 | 1 | 1 | -- | +-- Unionini
--+
+-- Anodontina **
Soroka, 2010 [Publ 3404], fig. 8a-M iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 2 | 2 | 1 | -- | +-- Cristariina **
--+
+-- Unionini
Curole & Kocher, 2002 [Publ 2712], fig. 1a-M iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 3 | 0 | 0 | -- | -- Utterbackiana_implicata **
Curole & Kocher, 2002 [Publ 2712], fig. 1b-M iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 3 | 0 | 0 | -- | -- Utterbackiana_implicata **
Inoue et al., 2013 [Publ 3579], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 8 | 0 | 0 | -- | Zanatta & Harris, 2013 [Publ 3530], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 4 | 0 | 0 | -- | Bolotov et al., 2016 [Publ 3743], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 29 | 2 | 1 | -- | +-- Anodontina **
--+
+-- Unionini
Zieritz et al., 2024 [Publ 4381], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 45 | 1 | 1 | -- | +-- Anodontina **
--+
+-- Unionini
Lydeard et al., 2000 [Publ 2654], fig. 2b iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 13 | 0 | 0 | -- | Metzger et al., 2018 [Publ 3952], fig. 1C iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 2 | 0 | 0 | -- | Inoue et al., 2020 [Publ 3941], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 40 | 0 | 0 | -- | -- Arcidens_confragosus **
Hoeh et al., 1998 [Publ 2695], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 4 | 0 | 0 | -- | Curole & Kocher, 2005 [Publ 2682], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 10 | 0 | 0 | -- | Gomes-dos-Santos et al., 2019 [Publ 3943], fig. 1-M iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 5 | 1 | 1 | -- | +-- Unionini
--+
+-- Anodontina **
Jiang et al., 2010 [Publ 3438], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 3 | 0 | 0 | -- | Marshall et al., 2014 [Publ 3593], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 13 | 1 | 1 | -- | +-- Alasmidontina **
--+
+-- Unionini
|
