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Page last updated
Thu 3 September 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

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Bivalvia | Unionida | Etherioidea | Iridinidae

subfamily ASPATHARIINAE Modell, 1942: cladomics
     2 genera

Cladomics Summary

number of species = 22 (23 total)
number of subtaxa (rank: genus) = 2
analyzed in 48 cladograms in 28 publications
maximum number of terminals in an analysis = 4 (mean = 1.67, median = 2)
maximum number of subtaxa (genus) = 2 (mean = 1.6, median = 2)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 30
maximum number of included terminals = 4, (mean = 2.07 median = 2)
maximum number of subtaxa (genus) = 2 (mean = 1.97, median = 2)
proportion of cladograms that support basic monophyly = 0.97

Rigorous Monophyly

number of cladograms that test rigorous monophyly = 21
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (genus) taxon and at least one rigorous outgroup (family).
maximum number of included terminals = 4 (mean = 2.1, median = 2)
maximum number of subtaxa (genus) = 2 (mean = 2, median = 2)
proportion of cladograms that support rigorous monophyly = 0.95

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
4432626310
  +-- Aspatharia **
--+             
  +-- Chambardia **

Combosch et al., 2017 [Publ 3806], fig. 1

iTtTbiriSoTroTroSiD
22021311--
  +-- Chambardia **
--+             
  +-- Aspatharia **

Bolotov et al., 2016 [Publ 3743], fig. 4

iTtTbiriSoTroTroSiD
22022800--
  +-- Chambardia **
--+             
  +-- Aspatharia **

Chen et al., 2025 [Publ 4455], fig. 1

iTtTbiriSoTroTroSiD
22022400--
  +-- Aspatharia **
--+             
  +-- Chambardia **

Whelan et al., 2011 [Publ 3380], fig. 1-ML

iTtTbiriSoTroTroSiD
22023411--
  +-- Aspatharia **
--+             
  +-- Chambardia **

Wu et al., 2020 [Publ 4121], fig. S1

iTtTbiriSoTroTroSiD
22023211--
  +-- Aspatharia **
--+             
  +-- Chambardia **

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iTtTbiriSoTroTroSiD
220212000--
  +-- Aspatharia **
--+             
  +-- Chambardia **

Bolotov et al., 2016 [Publ 3743], fig. 2

iTtTbiriSoTroTroSiD
22022800--
  +-- Aspatharia **
--+             
  +-- Chambardia **

Wu et al., 2021 [Publ 4366], fig. 2_I

iTtTbiriSoTroTroSiD
22028900--
  +-- Aspatharia **
--+             
  +-- Chambardia **

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
220219931--
  +-- Aspatharia **
--+             
  +-- Chambardia **

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
220219731--
  +-- Aspatharia **
--+             
  +-- Chambardia **

Sano et al., 2020 [Publ 4138], fig. 3

iTtTbiriSoTroTroSiD
22026711--
  +-- Aspatharia **
--+             
  +-- Chambardia **

Hoeh et al., 2009 [Publ 2716], fig. 3

iTtTbiriSoTroTroSiD
240245215
  +-- Chambardia **
  |             
--+-- Aspatharia **
  |             
  +-- Iridininae

Whelan et al., 2011 [Publ 3380], fig. 1-MP

iTtTbiriSoTroTroSiD
22023411--
  +-- Aspatharia **
--+             
  +-- Chambardia **

Galal-Khallaf et al., 2024 [Publ 4459], fig. 6

iTtTbiriSoTroTroSiD
22012621--
  +-- Iridininae
--+             
  +-- Chambardia **

Wu et al., 2020 [Publ 4121], fig. 2

iTtTbiriSoTroTroSiD
22023211--
  +-- Aspatharia **
--+             
  +-- Chambardia **

Huang et al., 2018 [Publ 3828], fig. 3_5

iTtTbiriSoTroTroSiD
22022711--
  +-- Aspatharia **
--+             
  +-- Chambardia **

Lopes-Lima et al., 2018 [Publ 3829], fig. 1

iTtTbiriSoTroTroSiD
22022611--
  +-- Aspatharia **
--+             
  +-- Chambardia **

Wu et al., 2021 [Publ 4366], fig. 6

iTtTbiriSoTroTroSiD
22028900--
  +-- Chambardia **
--+             
  +-- Aspatharia **

Sano et al., 2020 [Publ 4138], fig. 4

iTtTbiriSoTroTroSiD
22026711--
  +-- Aspatharia **
--+             
  +-- Chambardia **

Bolotov et al., 2017 [Publ 3761], fig. S2

iTtTbiriSoTroTroSiD
220213700--
  +-- Aspatharia **
--+             
  +-- Chambardia **

Bolotov et al., 2017 [Publ 3761], fig. S1

iTtTbiriSoTroTroSiD
220213700--
  +-- Chambardia **
--+             
  +-- Aspatharia **

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iTtTbiriSoTroTroSiD
22027911--
  +-- Aspatharia **
--+             
  +-- Chambardia **

Combosch et al., 2017 [Publ 3806], fig. 2

iTtTbiriSoTroTroSiD
22021311--
  +-- Chambardia **
--+             
  +-- Aspatharia **

Hoeh et al., 2009 [Publ 2716], fig. 5

iTtTbiriSoTroTroSiD
22024421--
  +----- Iridininae
--+                
  |  +-- Chambardia **
  +--+             
     +-- Aspatharia **

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
220219931--
  +-- Aspatharia **
--+             
  +-- Chambardia **

Hoeh et al., 2009 [Publ 2716], fig. 4

iTtTbiriSoTroTroSiD
22024421--
  +----- Iridininae
--+                
  |  +-- Chambardia **
  +--+             
     +-- Aspatharia **

Graf & Cummings, 2006 [Publ 1968], fig. 2

iTtTbiriSoTroTroSiD
22024521--
  +----- Iridininae
--+                
  |  +-- Chambardia **
  +--+             
     +-- Aspatharia **

Sano et al., 2020 [Publ 4138], fig. 2

iTtTbiriSoTroTroSiD
22026711--
  +-- Aspatharia **
--+             
  +-- Chambardia **

Lopes-Lima et al., 2018 [Publ 3829], fig. 3

iTtTbiriSoTroTroSiD
22022611--
  +-- Chambardia **
--+             
  +-- Aspatharia **

Graf et al., 2015 [Publ 3619], fig. 1

iTtTbiriSoTroTroSiD
11011900--
-- Chambardia_wahlbergi **

Elderkin et al., 2016 [Publ 3693], fig. 2

iTtTbiriSoTroTroSiD
1101200--
-- Aspatharia_pfeifferiana **

Walker et al., 2006 [Publ 2666], fig. 6

iTtTbiriSoTroTroSiD
11013521--
-- Chambardia_rubens **

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

iTtTbiriSoTroTroSiD
11013900--
-- Chambardia_wahlbergi **

González & Giribet, 2015 [Publ 3632], fig. 6

iTtTbiriSoTroTroSiD
1101500--
-- Aspatharia_pfeifferiana **

Lemer et al., 2019 [Publ 4093], fig. 1

iTtTbiriSoTroTroSiD
1101500--
-- Aspatharia_pfeifferiana **

González & Giribet, 2015 [Publ 3632], fig. 3

iTtTbiriSoTroTroSiD
1101500--
-- Aspatharia_pfeifferiana **

Guerra et al., 2019 [Publ 4168], fig. 2

iTtTbiriSoTroTroSiD
11014011--
-- Chambardia_rubens **

Bieler et al., 2014 [Publ 3622], fig. 28

iTtTbiriSoTroTroSiD
1101400--
-- Aspatharia_pfeifferiana **

Bieler et al., 2014 [Publ 3622], fig. 26_27_29_30

iTtTbiriSoTroTroSiD
1101400--
-- Aspatharia_pfeifferiana **

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

iTtTbiriSoTroTroSiD
110113911--
-- Chambardia_rubens **

González & Giribet, 2015 [Publ 3632], fig. 4

iTtTbiriSoTroTroSiD
1101500--
-- Aspatharia_pfeifferiana **

Smith, 2021 [Publ 4331], fig. 1

iTtTbiriSoTroTroSiD
11011900--
-- Aspatharia_pfeifferiana **

Guerra et al., 2019 [Publ 4168], fig. 1

iTtTbiriSoTroTroSiD
11014011--
-- Chambardia_rubens **

Konopleva et al., 2017 [Publ 3807], fig. 6

iTtTbiriSoTroTroSiD
11013000--
-- Chambardia_wahlbergi **

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iTtTbiriSoTroTroSiD
11015600--
-- Aspatharia_pfeifferiana **

González & Giribet, 2015 [Publ 3632], fig. 2

iTtTbiriSoTroTroSiD
1101500--
-- Aspatharia_pfeifferiana **

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iTtTbiriSoTroTroSiD
11015600--
-- Aspatharia_pfeifferiana **
 
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