The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)
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Bivalvia | Unionida | Unionoidea | Unionidae | Gonideinae tribe CHAMBERLAINIINI Bogan, Froufe & Lopes-Lima in Lopes-Lima et al., 2017: cladomics
2 genera Cladomics Summarynumber of species = 6 (6 total)
number of subtaxa (rank: genus) = 2
analyzed in 129 cladograms in 86 publications
maximum number of terminals in an analysis =
6 (mean = 2.14, median = 2)
maximum number of subtaxa (genus) =
2 (mean = 1.48, median = 1) Basic Monophylynumber of cladograms that test basic monophyly
(i.e., more than 1 included species) = 97
maximum number of included terminals = 6, (mean =
2.52 median = 2)
maximum number of subtaxa (genus) =
2 (mean = 1.64, median = 2)
proportion of cladograms that support basic monophyly
= 0.94 Rigorous Monophylynumber of cladograms that test rigorous monophyly = 61
(Rigorous monophyly refers to cladograms with terminal species representing
more than one subtaxon (genus) taxon and at least one rigorous outgroup (family).
maximum number of included terminals = 6 (mean = 2.79, median = 3)
maximum number of subtaxa (genus) =
2 (mean = 2, median = 2)
proportion of cladograms that support rigorous monophyly = 0.9
Pairwise Cladogram ClusteringThe graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.
Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14 ** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram ** iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 9 | 2 | 626 | 129 | 6 | 0 | +----------------------------- Indopseudodontina
|
| +-- Lannanaia
+--+ +-----------------------+
| | | +-- Pilsbryoconcha
| | |
| +--+ +----------------------- Songkhlanaia
| | |
| | | +-- Nyeinchanconcha
| +--+ +-----------------+
| | | +-- Sundadontina
| | |
| | | +-- Isannaia
| +--+ +--+
| | | +-- Namkongnaia
| | +-----------+
| | | | +-- Thaiconcha
| | | +--+
| +--+ +-- Bineurus
| |
| | +-------------- Pseudodon_walpolei
| | |
| +--+ +----------- Pseudodon_vagulus
| | |
| | | +----- Pseudodon_vondembuschianus
| +--+ +--+
--+ | | | +-- Pseudodon_mekongi
| | | +--+
| +--+ +-- Schepmaniini
| |
| | +----- Pseudodon_lenyanensis__2
| +--+
| | +-- Pseudodon_inoscularis
| +--+
| +-- Pseudodon_lenyanensis__1
|
| +-- Chamberlainia **
| +-----------------+
| | +-- Sinohyriopsis **
| +--+
| | | +----------------- Parvasolenaia
| | | |
| | +--+ +-------------- Pseudopostolata
| | | |
| | +--+ +----------- Koreosolenaia
| | | |
| | +--+ +-- Cosmopseudodon
| | | +-----+
| | | | +-- Guiunio
+--------+ +--+
| | +----- Ptychorhynchus
| +--+
| | +-- Obovalis
| +--+
| +-- Postolata
|
| +-- Caudiculatus
| +--------------+
| | +-- Lamprotulini
| |
+-----+ +-------------- Sinosolenaia_carinata
| |
| | +----- Gonidea
+--+ +-----+
| | | +-- Microcondylaea
| | +--+
+--+ +-- Leguminaia
|
| +-------- Inversidens
+--+
| +----- Sinosolenaia_recognita
+--+
| +-- Sinosolenaia_iridinea
+--+
+-- Sinosolenaia_oleivora
Kongim et al., 2023 [Publ 4290], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 4 | 2 | 9 | 9 | 3 | 7 | +-- Chamberlainia **
+-----+
| +-- Sinohyriopsis **
--+
| +----- Lamprotulini
+--+
| +-- Gonideini
+--+
+-- Pseudodontini
Goncalves et al., 2022 [Publ 4253], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 6 | 6 | 5 | 2 | 1 | 1 | 1 | 7 | +----- Lamprotulini
--+
| +-- Chamberlainia **
+--+
+-- Sinohyriopsis **
Lopes-Lima et al., 2020 [Publ 4080], fig. 17 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 3 | 2 | 2 | 0 | 0 | 8 | +-- Chamberlainia **
--+
+-- Sinohyriopsis **
Wu et al., 2021 [Publ 4366], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 2 | 66 | 10 | 3 | 8 | +-- Chamberlainia **
+-----+
| +-- Sinohyriopsis **
--+
| +----- Lamprotulini
+--+
| +-- Pseudodontini
+--+
+-- Gonideini
Froufe et al., 2019 [Publ 3936], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 2 | 41 | 10 | 3 | 8 | +-- Chamberlainia **
+-----+
| +-- Sinohyriopsis **
--+
| +----- Lamprotulini
+--+
| +-- Pseudodontini
+--+
+-- Gonideini
Konopleva et al., 2023 [Publ 4269], fig. S1_4 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 2 | 53 | 10 | 2 | 8 | +-------- Gonideini
--+
| +----- Lamprotulini
+--+
| +-- Chamberlainia **
+--+
+-- Sinohyriopsis **
Froufe et al., 2019 [Publ 3936], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 2 | 41 | 10 | 3 | 8 | +-- Chamberlainia **
+-----+
| +-- Sinohyriopsis **
--+
| +----- Lamprotulini
+--+
| +-- Pseudodontini
+--+
+-- Gonideini
Wu et al., 2021 [Publ 4215], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 2 | 21 | 16 | 3 | 8 | +-- Chamberlainia **
+-----------------+
| +-- Sinohyriopsis **
|
--+ +----------------- Pseudodontini
| |
| | +-- Sinosolenaia
+--+ +-----------+
| | +-- Lamprotulini
| |
+--+ +-- Ptychorhynchus
| +--------+
| | +-- Parvasolenaia
+--+
| +-------- Inversidens
+--+
| +----- Gonidea
+--+
| +-- Microcondylaea
+--+
+-- Leguminaia
Wu et al., 2023 [Publ 4339], fig. 4_S5-nt iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 2 | 93 | 17 | 4 | 8 | +-- Chamberlainia **
+--------+
| +-- Sinohyriopsis **
--+
| +-------- Rectidentini
+--+
| +----- Lamprotulini
+--+
| +-- Pseudodontini
+--+
+-- Gonideini
Wu et al., 2021 [Publ 4366], fig. 3-ML iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 2 | 71 | 11 | 3 | 8 | +-- Chamberlainia **
+-----+
| +-- Sinohyriopsis **
--+
| +----- Lamprotulini
+--+
| +-- Pseudodontini
+--+
+-- Gonideini
Huang et al., 2019 [Publ 3843], fig. S3 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 2 | 195 | 51 | 4 | 8 | +-- Chamberlainia **
--+
+-- Sinohyriopsis **
Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 2 | 119 | 30 | 4 | 8 | +-- Pseudodontini
+-----+
| +-- Gonideini
--+
| +----- Lamprotulini
+--+
| +-- Chamberlainia **
+--+
+-- Sinohyriopsis **
Dai et al., 2025 [Publ 4441], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 2 | 44 | 39 | 5 | 8 | +-------------- Rectidentini
|
--+ +-- Chamberlainia **
| +--------+
| | +-- Sinohyriopsis **
+--+
| +-------- Lamprotulini
+--+
| +----- Gonideini
+--+
| +-- Schepmaniini
+--+
+-- Pseudodontini
Dai et al., 2023 [Publ 4285], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 2 | 50 | 16 | 4 | 8 | +-- Chamberlainia **
+--------+
| +-- Sinohyriopsis **
--+
| +-------- Rectidentini
+--+
| +----- Lamprotulini
+--+
| +-- Pseudodontini
+--+
+-- Gonideini
Wu et al., 2022 [Publ 4238], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 2 | 71 | 11 | 3 | 8 | +-- Chamberlainia **
+-----+
| +-- Sinohyriopsis **
--+
| +----- Lamprotulini
+--+
| +-- Pseudodontini
+--+
+-- Gonideini
Matos et al., 2024 [Publ 4383], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 2 | 17 | 15 | 4 | 8 | +-- Chamberlainia **
+--------+
| +-- Sinohyriopsis **
--+
| +-------- Rectidentini
+--+
| +----- Lamprotulini
+--+
| +-- Pseudodontini
+--+
+-- Gonideini
Huang et al., 2019 [Publ 3843], fig. S4 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 127 | 72 | 2 | 197 | 52 | 4 | 131 | +----------- Chamberlainia **
|
+----------- Megalonaias
|
+----------- Uniomerus
|
+----------- Quadrulina
|
+----------- Sinohyriopsis **
|
+----------- Lamprotulini
|
+----------- Gonideini
|
+----------- Pseudodontini
|
+----------- Rectidentini
|
| +-------- Amblema
| |
| +-------- Toxolasma
| |
| +-------- Pleurobemini
| |
| | +-- Obliquaria
| +-----+
| | +-- Plectomerus
+--+
| | +----- Ellipsaria
| | |
| | +----- Potamilus_leptodon
| | |
| | +----- Medionidus
| | |
| | +----- Potamilus_alatus
| | |
--+ | +----- Ptychobranchus
| +--+
| +----- Truncilla
| |
| +----- Venustaconcha
| |
| +----- Cambarunio
| |
| +----- Leaunio
| |
| | +-- Ortmanniana
| +--+
| +-- Lampsilis
|
| +-------- Acuticostini
| |
| +-------- Alasmidonta
| |
| +-------- Anodonta_anatina
| |
| +-------- Anemina
| |
| +-------- Anodonta_cygnea
| |
| +-------- Cristaria
| |
| +-------- Brachyanodon
| |
| +-------- Lasmigona
| |
| +-------- Lepidodesmini
| |
| +-------- Pseudanodonta
| |
| +-------- Pyganodon
| |
+--+-------- Simpsonaias
|
+-------- Strophitus
|
+-------- Utterbackia
|
+-------- Lanceolariini
|
+-------- Schistodesmus
|
+-------- Unionina
|
+-------- Aculamprotulina
|
| +-- Pletholophus
+-----+
| +-- Sinanodonta
|
| +----- Cuneopsis_celtiformis
| |
| +----- Tchangsinaia
| |
+--+----- Cuneopsis_rufescens
|
+----- Nodularia
|
| +-- Cuneopsis_szechenyii
+--+
+-- Cuneopsis_heudei
Dai et al., 2024 [Publ 4389], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 2 | 43 | 38 | 5 | 8 | +-------------- Rectidentini
|
--+ +-- Chamberlainia **
| +--------+
| | +-- Sinohyriopsis **
+--+
| +-------- Lamprotulini
+--+
| +----- Gonideini
+--+
| +-- Schepmaniini
+--+
+-- Pseudodontini
Dai et al., 2023 [Publ 4285], fig. 3-ML iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 2 | 42 | 40 | 4 | 8 | +----------------------- Pseudodontini
|
| +-------------------- Rectidentini
--+ |
| | +-------- Inversidens
| | +--------+
+--+ | | +----- Lamprotulini
| | +--+
| | | +-- Chamberlainia **
| | +--+
+--+ +-- Sinohyriopsis **
|
| +-------------- Sinosolenaia
| |
| | +----- Gonidea
+--+ +-----+
| | | +-- Leguminaia
| | +--+
| | +-- Microcondylaea
+--+
| +-- Koreosolenaia
| +-----+
| | +-- Parvasolenaia
+--+
| +----- Postolata
+--+
| +-- Ptychorhynchus
+--+
+-- Obovalis
Pavan-Kumar et al., 2022 [Publ 4263], fig. S5 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 2 | 36 | 9 | 3 | 8 | +-- Chamberlainia **
+-----+
| +-- Sinohyriopsis **
--+
| +----- Gonideini
+--+
| +-- Pseudodontini
+--+
+-- Lamprotulini
Chen et al., 2025 [Publ 4663], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 2 | 45 | 40 | 5 | 8 | +-------------- Rectidentini
|
--+ +-- Chamberlainia **
| +--------+
| | +-- Sinohyriopsis **
+--+
| +-------- Lamprotulini
+--+
| +----- Gonideini
+--+
| +-- Schepmaniini
+--+
+-- Pseudodontini
Wu et al., 2023 [Publ 4339], fig. 4_S5-aa iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 2 | 93 | 17 | 4 | 8 | +-- Chamberlainia **
+--------+
| +-- Sinohyriopsis **
--+
| +-------- Rectidentini
+--+
| +----- Lamprotulini
+--+
| +-- Pseudodontini
+--+
+-- Gonideini
Wu et al., 2021 [Publ 4366], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 2 | 88 | 26 | 4 | 8 | +-- Chamberlainia **
+-----+
| +-- Sinohyriopsis **
--+
| +----- Lamprotulini
+--+
| +-- Pseudodontini
+--+
+-- Gonideini
Wu et al., 2026 [Publ 4811], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 2 | 93 | 25 | 4 | 8 | +-- Chamberlainia **
+--------+
| +-- Sinohyriopsis **
--+
| +-------- Rectidentini
+--+
| +----- Lamprotulini
+--+
| +-- Pseudodontini
+--+
+-- Gonideini
Zieritz et al., 2021 [Publ 4231], fig. 3-F_5 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 2 | 45 | 14 | 4 | 8 | +-- Chamberlainia **
+--------+
| +-- Sinohyriopsis **
--+
| +-------- Rectidentini
+--+
| +----- Lamprotulini
+--+
| +-- Pseudodontini
+--+
+-- Gonideini
Wu et al., 2024 [Publ 4350], fig. 8a_S1 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 2 | 89 | 18 | 4 | 8 | +-- Chamberlainia **
+--------+
| +-- Sinohyriopsis **
--+
| +-------- Rectidentini
+--+
| +----- Lamprotulini
+--+
| +-- Pseudodontini
+--+
+-- Gonideini
Wu et al., 2021 [Publ 4366], fig. 3-BI iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 2 | 71 | 11 | 3 | 8 | +-- Chamberlainia **
+-----+
| +-- Sinohyriopsis **
--+
| +----- Lamprotulini
+--+
| +-- Pseudodontini
+--+
+-- Gonideini
Zhang et al., 2024 [Publ 4397], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 2 | 24 | 22 | 4 | 8 | +-- Chamberlainia **
+--------+
| +-- Sinohyriopsis **
--+
| +-------- Rectidentini
+--+
| +----- Lamprotulini
+--+
| +-- Pseudodontini
+--+
+-- Gonideini
Chen et al., 2020 [Publ 4170], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 34 | 32 | 2 | 31 | 8 | 3 | 36 | +----- Prolasmidonta
+--+
| | +-- Alasmidonta
| +--+
+-----------------+ +-- Anemina
| |
| | +----- Pseudodontini
| +--+
| | +-- Lamprotula_caveata
| +--+
| +-- Chamberlainia **
|
| +----- Oxynaiina
| +-----+
| | | +-- Lepidodesmini
--+ | +--+
| | +-- Aculamprotulina
| +-----------+
| | | +----- Platynaias
| | | +--+
| | | | | +-- Utterbackia_imbecillis
| | | | +--+
| | +--+ +-- Pyganodon
| | |
| | | +----- Sinanodonta
| | +--+
| | | +-- Cristaria
+--+ +--+
| +-- Anodontina
|
| +-- Utterbackia_peninsularis
| +-----+
| | +-- Potamilus_alatus
| +-----------+
| | | +----- Lamprotula_leaii
| | +--+
| | | +-- Sinohyriopsis **
+--+ +--+
| +-- Gonideini
|
| +----------------- Gibbosula
+--+
| +-------------- Quadrulini
+--+
| +----------- Toxolasma
+--+
| +-------- Lampsilis_ornata
+--+
| +----- Potamilus_leptodon
+--+
| +-- Lampsilis_siliquoidea
+--+
+-- Lampsilis_powellii
Wu et al., 2024 [Publ 4350], fig. 8b_S2 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 2 | 89 | 18 | 4 | 8 | +-- Chamberlainia **
+--------+
| +-- Sinohyriopsis **
--+
| +-------- Rectidentini
+--+
| +----- Lamprotulini
+--+
| +-- Pseudodontini
+--+
+-- Gonideini
Wu et al., 2021 [Publ 4366], fig. 2_I iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 2 | 88 | 26 | 4 | 8 | +-- Chamberlainia **
+-----+
| +-- Sinohyriopsis **
--+
| +----- Lamprotulini
+--+
| +-- Pseudodontini
+--+
+-- Gonideini
Pavan-Kumar et al., 2022 [Publ 4263], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 2 | 34 | 8 | 3 | 8 | +-- Chamberlainia **
+-----+
| +-- Sinohyriopsis **
--+
| +----- Lamprotulini
+--+
| +-- Pseudodontini
+--+
+-- Gonideini
Wu et al., 2024 [Publ 4388], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 2 | 50 | 19 | 4 | 8 | +-- Chamberlainia **
+--------+
| +-- Sinohyriopsis **
--+
| +-------- Rectidentini
+--+
| +----- Lamprotulini
+--+
| +-- Pseudodontini
+--+
+-- Gonideini
Dai et al., 2025 [Publ 4440], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 2 | 85 | 17 | 4 | 8 | +-- Chamberlainia **
+--------+
| +-- Sinohyriopsis **
--+
| +-------- Rectidentini
+--+
| +----- Lamprotulini
+--+
| +-- Pseudodontini
+--+
+-- Gonideini
Dai et al., 2023 [Publ 4285], fig. 3-BI iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 2 | 42 | 40 | 4 | 8 | +----------------------- Pseudodontini
|
--+ +-------------------- Rectidentini
| |
| | +----------------- Inversidens
+--+ |
| | +----- Lamprotulini
| +-----------+
| | | +-- Chamberlainia **
+--+ +--+
| +-- Sinohyriopsis **
|
| +-------------- Sinosolenaia
| |
| | +----- Gonidea
+--+ +-----+
| | | +-- Leguminaia
| | +--+
| | +-- Microcondylaea
+--+
| +-- Koreosolenaia
| +-----+
| | +-- Parvasolenaia
+--+
| +----- Postolata
+--+
| +-- Ptychorhynchus
+--+
+-- Obovalis
Huang et al., 2019 [Publ 3843], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 2 | 197 | 52 | 4 | 8 | +-- Chamberlainia **
+-----+
| +-- Sinohyriopsis **
--+
| +----- Lamprotulini
+--+
| +-- Gonideini
+--+
+-- Pseudodontini
Chen et al., 2025 [Publ 4448], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 2 | 83 | 15 | 4 | 8 | +-- Chamberlainia **
+--------+
| +-- Sinohyriopsis **
--+
| +-------- Rectidentini
+--+
| +----- Lamprotulini
+--+
| +-- Pseudodontini
+--+
+-- Gonideini
Matos et al., 2024 [Publ 4384], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 2 | 17 | 15 | 4 | 8 | +-- Chamberlainia **
+--------+
| +-- Sinohyriopsis **
--+
| +-------- Rectidentini
+--+
| +----- Lamprotulini
+--+
| +-- Pseudodontini
+--+
+-- Gonideini
Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 2 | 2 | 138 | 23 | 4 | 8 | +-- Chamberlainia **
+--------+
| +-- Sinohyriopsis **
--+
| +-------- Rectidentini
+--+
| +----- Lamprotulini
+--+
| +-- Pseudodontini
+--+
+-- Gonideini
Huang et al., 2019 [Publ 3843], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 43 | 10 | 2 | -- | Bolotov et al., 2017 [Publ 3761], fig. S1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 136 | 46 | 4 | -- | +-------- Gonideini
--+
| +----- Lamprotulini
+--+
| +-- Sinohyriopsis **
+--+
+-- Chamberlainia **
Fonseca et al., 2016 [Publ 3775], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 29 | 5 | 2 | -- | +----- Sinohyriopsis **
--+
| +-- Gonideini
+--+
+-- Lamprotulini
Sano et al., 2020 [Publ 4138], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 66 | 6 | 2 | -- | Wu et al., 2018 [Publ 4143], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 20 | 4 | 2 | -- | +----- Sinohyriopsis **
--+
| +-- Gonideini
+--+
+-- Lamprotulini
Pfeiffer et al., 2019 [Publ 3937], fig. 4-5 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 55 | 15 | 4 | -- | +-- Chamberlainia **
+--------+
| +-- Sinohyriopsis **
--+
| +-------- Rectidentini
+--+
| +----- Lamprotulini
+--+
| +-- Gonideini
+--+
+-- Pseudodontini
Wang et al., 2018 [Publ 3865], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 29 | 6 | 2 | -- | +----- Sinohyriopsis **
--+
| +-- Gonideini
+--+
+-- Lamprotulini
Wu et al., 2019 [Publ 3864], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 44 | 6 | 2 | -- | +----- Sinohyriopsis **
--+
| +-- Lamprotulini
+--+
+-- Gonideini
Lopes-Lima et al., 2017 [Publ 3792], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 30 | 5 | 2 | -- | +----- Sinohyriopsis **
--+
| +-- Gonideini
+--+
+-- Lamprotulini
Bolotov et al., 2017 [Publ 3761], fig. S2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 136 | 0 | 2 | 136 | 46 | 4 | 142 | +-------- Chamberlainia **
|
+--------------+ +----- Cumberlandia
| | |
| +--+ +-- Gibbosula
| | |
| +--+-- Pseudunio
| |
| +-- Margaritifera
|
| +-- Tamsiella
| +--+
| | +-- Etheria
| +--------+
| | | +-- Aspatharia
| | +--+
| | +-- Chambardia
| |
+--------+ +----- Rhipidodontini
| | +-----+
| | | | +-- Castaliini
| | | +--+
| | | +-- Hyriini
| +--+
| | +----- Velesunio
--+ | +--+
| | | | +-- Alathyria_jacksoni
| | | +--+
| +--+ +-- Alathyria_profuga
| |
| | +----- Microdontia
| +--+
| | +-- Alathyria_pertexta
| +--+
| +-- Lortiella
|
| +-- Parreysiini
| +-----+
| | +-- Lamellidentini
| +-----------+
| | | +----- Leoparreysiini
| | +--+
| | | +-- Coelaturini
| | +--+
| | +-- Indochinellini
| |
| | +----------- Unionini
| | |
+--+ +-----+ +-------- Lanceolariini
| | | |
| | | +-------- Cristariina
| | +--+
| | | +----- Anodonta_anatina
| | | |
| | | +----- Pseudanodonta
| | +--+
| | +----- Alasmidontina
+--+ |
| | +-- Anodonta_cygnea
| +--+
| +-- Brachyanodon
|
| +-------------- Rectidentini
| |
| | +-------- Pseudodontini
| | +--+
+--+ | | +----- Gonideini
| | +--+
| | | +-- Sinohyriopsis **
| | +--+
+--+ +-- Lamprotulini
|
| +-- Amblemini
| +--+
| | +-- Lampsilini
+-----+
| +-- Quadrulini
+--+
+-- Pleurobemini
Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 78 | 26 | 4 | -- | +----- Lamprotulini
+--------+
| | +-- Chamberlainia **
| +--+
| +-- Sinohyriopsis **
--+
| +-- Parvasolenaia
| +--------+
| | +-- Pseudodontini
+--+
| +-------- Sinosolenaia
+--+
| +----- Gonidea
+--+
| +-- Microcondylaea
+--+
+-- Leguminaia
Wu et al., 2020 [Publ 4121], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 47 | 6 | 2 | -- | +----- Sinohyriopsis **
--+
| +-- Lamprotulini
+--+
+-- Gonideini
Wang et al., 2016 [Publ 3714], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 1 | 0 | 0 | -- | Lopes-Lima et al., 2020 [Publ 4080], fig. 16 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 6 | 0 | 2 | 6 | 4 | 1 | 11 | +----- Chamberlainia **
--+
| +-- Sinohyriopsis **
+--+
+-- Lamprotulini
Huang et al., 2019 [Publ 3843], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 54 | 8 | 2 | -- | +----- Sinohyriopsis **
--+
| +-- Lamprotulini
+--+
+-- Gonideini
Klymus et al., 2025 [Publ 4814], fig. 3B iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 34 | 12 | 4 | -- | +-- Sinohyriopsis **
+--------+
| +-- Chamberlainia **
--+
| +-------- Rectidentini
+--+
| +----- Pseudodontini
+--+
| +-- Lamprotulini
+--+
+-- Gonideini
Pfeiffer & Graf, 2013 [Publ 3531], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 29 | 9 | 4 | -- | +-------- Sinohyriopsis **
|
--+ +-- Gonideini
| +--+
| | +-- Pseudodontini
+--+
| +-- Rectidentini
+--+
+-- Lamprotulini
Zieritz et al., 2021 [Publ 4231], fig. 3-M_4 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 21 | 10 | 4 | -- | +-- Chamberlainia **
+--------+
| +-- Sinohyriopsis **
--+
| +-------- Rectidentini
+--+
| +----- Pseudodontini
+--+
| +-- Lamprotulini
+--+
+-- Gonideini
Sano et al., 2017 [Publ 3799], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 31 | 6 | 2 | -- | +-- Obovalis
+-----+
| +-- Ptychorhynchus
|
--+ +-- Inversidens
| +--+
| | +-- Sinosolenaia
+--+
| +-- Lamprotulini
+--+
+-- Sinohyriopsis **
Zhou et al., 2016 [Publ 3710], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 18 | 4 | 2 | -- | +----- Sinohyriopsis **
--+
| +-- Lamprotulini
+--+
+-- Gonideini
Zhou et al., 2017 [Publ 3788], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 14 | 5 | 2 | -- | +----- Sinohyriopsis **
--+
| +-- Lamprotulini
+--+
+-- Gonideini
Dai et al., 2024 [Publ 4358], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 51 | 9 | 3 | -- | +-- Chamberlainia **
+-----+
| +-- Sinohyriopsis **
--+
| +----- Lamprotulini
+--+
| +-- Pseudodontini
+--+
+-- Gonideini
Bolotov et al., 2017 [Publ 3764], fig. S1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 13 | 0 | 2 | 122 | 48 | 4 | 16 | +-- Chamberlainia **
|
+-- Sinohyriopsis **
|
+-- Lamprotula_caveata
|
+-- Lamprotula_leaii
|
+-- Potomida
|
+-- Pronodularia
|
--+-- Gonidea
|
+-- Leguminaia_wheatleyi
|
+-- Leguminaia
|
+-- Microcondylaea
|
+-- Sinosolenaia_carinata
|
+-- Sinosolenaia_oleivora
|
+-- Sinosolenaia_recognita
Huang et al., 2019 [Publ 3843], fig. S1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 45 | 10 | 2 | -- | +----------------- Sinohyriopsis **
|
| +----- Parvasolenaia_rivularis
--+ +--------+
| | | +-- Parvasolenaia_triangularis
| | +--+
| | +-- Ptychorhynchus
+--+
| +-- Sinosolenaia_carinata
| +--------+
| | +-- Lamprotulini
+--+
| +-------- Sinosolenaia_recognita
| |
+--+ +----- Sinosolenaia_oleivora__2
| |
+--+ +-- Sinosolenaia_oleivora__1
| |
+--+-- Sinosolenaia_iridinea
|
+-- Sinosolenaia_oleivora__3
Fukata & Iigo, 2020 [Publ 4106], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 52 | 10 | 3 | -- | +-------- Sinohyriopsis **
--+
| +----- Pseudodontini
+--+
| +-- Lamprotulini
+--+
+-- Gonideini
Cha et al., 2018 [Publ 3916], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 36 | 7 | 2 | -- | +----- Sinohyriopsis **
--+
| +-- Gonideini
+--+
+-- Lamprotulini
Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 74 | 29 | 4 | -- | +----- Lamprotulini
--+
| +-- Chamberlainia **
+--+
+-- Sinohyriopsis **
Peng et al., 2012 [Publ 3689], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 1 | 0 | 0 | -- | Bolotov et al., 2020 [Publ 4100], fig. S1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 109 | 69 | 4 | -- | +-------------- Rectidentini
--+
| +----------- Pseudodontini
+--+
| +-------- Gonideini
+--+
| +----- Lamprotulini
+--+
| +-- Sinohyriopsis **
+--+
+-- Chamberlainia **
Plazzi et al., 2011 [Publ 3676], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 16 | 1 | 1 | -- | Aunins et al., 2018 [Publ 3863], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 38 | 5 | 2 | -- | +----- Sinohyriopsis **
--+
| +-- Lamprotulini
+--+
+-- Gonideini
Wu et al., 2020 [Publ 4147], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 28 | 6 | 2 | -- | +----- Sinohyriopsis **
--+
| +-- Gonideini
+--+
+-- Lamprotulini
Wang et al., 2018 [Publ 3846], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 29 | 6 | 2 | -- | +----- Sinohyriopsis **
--+
| +-- Lamprotulini
+--+
+-- Gonideini
Jeratthitikul et al., 2021 [Publ 4219], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 32 | 32 | 4 | -- | +-------------- Rectidentini
--+
| +----------- Pseudodontini
+--+
| +-------- Gonideini
+--+
| +----- Lamprotulini
+--+
| +-- Chamberlainia **
+--+
+-- Sinohyriopsis **
Huang et al., 2018 [Publ 3828], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 40 | 6 | 2 | -- | +----- Sinohyriopsis **
--+
| +-- Gonideini
+--+
+-- Lamprotulini
Guerra et al., 2017 [Publ 3772], fig. 2b iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 23 | 3 | 2 | -- | +----- Sinohyriopsis **
--+
| +-- Lamprotulini
+--+
+-- Gonideini
Sano et al., 2020 [Publ 4138], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 66 | 6 | 2 | -- | +----------- Sinohyriopsis **
--+
| +-------- Inversidens
+--+
| +----- Lamprotulini
+--+
| +-- Ptychorhynchus
+--+
+-- Obovalis
Jeratthitikul et al., 2021 [Publ 4219], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 33 | 33 | 4 | -- | +-------------- Rectidentini
--+
| +----------- Pseudodontini
+--+
| +-------- Gonideini
+--+
| +----- Lamprotulini
+--+
| +-- Chamberlainia **
+--+
+-- Sinohyriopsis **
Froufe et al., 2019 [Publ 3936], fig. 3-M iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 17 | 6 | 3 | -- | +-- Chamberlainia **
+-----+
| +-- Sinohyriopsis **
--+
| +----- Lamprotulini
+--+
| +-- Gonideini
+--+
+-- Pseudodontini
Guerra et al., 2017 [Publ 3772], fig. 2a iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 23 | 3 | 2 | -- | +----- Sinohyriopsis **
--+
| +-- Lamprotulini
+--+
+-- Gonideini
Sheng et al., 2014 [Publ 3704], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 6 | 1 | 1 | -- | +-- Lamprotulini
--+
+-- Sinohyriopsis **
Pfeiffer et al., 2019 [Publ 3937], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 55 | 15 | 4 | -- | +-- Chamberlainia **
+--------+
| +-- Sinohyriopsis **
--+
| +-------- Rectidentini
+--+
| +----- Pseudodontini
+--+
| +-- Gonideini
+--+
+-- Lamprotulini
Dai et al., 2024 [Publ 4386], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 40 | 8 | 3 | -- | +-- Chamberlainia **
+-----+
| +-- Sinohyriopsis **
--+
| +----- Lamprotulini
+--+
| +-- Pseudodontini
+--+
+-- Gonideini
Wei et al., 2016 [Publ 3665], fig. 0 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 4 | 0 | 0 | -- | Jeratthitikul et al., 2024 [Publ 4387], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 57 | 57 | 5 | -- | +----- Rectidentini
+-----+
| | +-- Chamberlainia **
| +--+
--+ +-- Sinohyriopsis **
|
| +-------- Lamprotulini
+--+
| +----- Gonideini
+--+
| +-- Schepmaniini
+--+
+-- Pseudodontini
Wu et al., 2016 [Publ 3711], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 19 | 4 | 2 | -- | +----- Sinohyriopsis **
--+
| +-- Gonideini
+--+
+-- Lamprotulini
Sano et al., 2020 [Publ 4138], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 32 | 6 | 2 | -- | +----- Gonideini
--+
| +-- Lamprotulini
+--+
+-- Sinohyriopsis **
Campbell & Lydeard, 2026 [Publ 4810], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 45 | 13 | 4 | -- | +-- Chamberlainia **
+--------+
| +-- Sinohyriopsis **
--+
| +-------- Rectidentini
+--+
| +----- Pseudodontini
+--+
| +-- Lamprotulini
+--+
+-- Gonideini
Shirai et al., 2010 [Publ 3353], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 1 | 1 | 1 | -- | +-- Lamprotulini
--+
+-- Sinohyriopsis **
Froufe et al., 2019 [Publ 3936], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 17 | 6 | 3 | -- | +-- Chamberlainia **
+-----+
| +-- Sinohyriopsis **
--+
| +----- Lamprotulini
+--+
| +-- Gonideini
+--+
+-- Pseudodontini
Bolotov et al., 2020 [Publ 4100], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 16 | 0 | 2 | 115 | 69 | 4 | 20 | +-------- Rectidentini
|
| +----- Pseudodontini
| |
| | +-- Sinosolenaia_iridinea
--+ | |
| | +-- Sinosolenaia_recognita__1
| | |
| | +-- Sinosolenaia_carinata
| | |
| | +-- Leguminaia
+--+ |
| +-- Microcondylaea
| |
| +-- Gonidea
| |
| +-- Ptychorhynchus
| |
| +-- Parvasolenaia
| |
+--+-- Lamprotula_leaii
|
+-- Lamprotula
|
+-- Lamprotula_caveata
|
+-- Potomida
|
+-- Pronodularia
|
+-- Sinohyriopsis **
|
+-- Chamberlainia **
|
+-- Sinosolenaia_recognita__2
|
+-- Sinosolenaia_oleivora
Huang et al., 2013 [Publ 3629], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 14 | 2 | 2 | -- | +----- Sinohyriopsis **
--+
| +-- Gonideini
+--+
+-- Lamprotulini
Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 74 | 29 | 4 | -- | +----- Lamprotulini
--+
| +-- Chamberlainia **
+--+
+-- Sinohyriopsis **
Froufe et al., 2015 [Publ 3664], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 10 | 5 | 2 | -- | Klymus et al., 2025 [Publ 4814], fig. 3A iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 34 | 12 | 4 | -- | +-- Chamberlainia **
+--------+
| +-- Sinohyriopsis **
--+
| +-------- Rectidentini
+--+
| +----- Pseudodontini
+--+
| +-- Lamprotulini
+--+
+-- Gonideini
Zhou et al., 2007 [Publ 2658], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 19 | 3 | 1 | -- | Sano et al., 2020 [Publ 4138], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 1 | 66 | 6 | 2 | -- | +----------- Sinohyriopsis **
--+
| +-------- Inversidens
+--+
| +----- Lamprotulini
+--+
| +-- Obovalis
+--+
+-- Ptychorhynchus
Huang et al., 2019 [Publ 3843], fig. 3-M iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 22 | 6 | 2 | -- | +----- Sinohyriopsis **
--+
| +-- Lamprotulini
+--+
+-- Gonideini
Wu et al., 2024 [Publ 4350], fig. 6-7 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 35 | 33 | 5 | -- | +----------- Chamberlainia **
|
--+ +-------- Rectidentini
| |
+--+ +-- Schepmaniini
| +--+
| | +-- Pseudodontini
+--+
| +-- Lamprotulini
+--+
+-- Gonideini
Campbell et al., 2005 [Publ 2660], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 101 | 3 | 2 | -- | -- Sinohyriopsis_cumingii **
Zieritz et al., 2024 [Publ 4381], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 44 | 33 | 6 | -- | +-- Chamberlainia **
--+
+-- Lamprotulini
Teiga-Teixeira et al., 2020 [Publ 4173], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 18 | 1 | 1 | -- | +-- Chamberlainia **
--+
+-- Lamprotulini
Burzynski et al., 2017 [Publ 3774], fig. 1-top iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 12 | 4 | 2 | -- | +----- Sinohyriopsis **
--+
| +-- Gonideini
+--+
+-- Lamprotulini
Burzynski et al., 2017 [Publ 3774], fig. 1-bottom iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 12 | 4 | 2 | -- | +----- Sinohyriopsis **
--+
| +-- Gonideini
+--+
+-- Lamprotulini
Wu et al., 2024 [Publ 4350], fig. 5c iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 35 | 33 | 5 | -- | +----------------------------- Schepmania_nieuwenhuisi__1
|
+----------------------------- Schepmania_nieuwenhuisi__2
--+
| +-------------------------- Pseudodontini
| |
+--+ +----------------------- Rectidentini
| |
+--+ +-------------------- Inversidens
| |
| | +----- Lamprotula
+--+ +-----------+
| | | +-- Chamberlainia **
| | +--+
+--+ +-- Potomida
|
| +-------------- Sinosolenaia
| |
+--+ +----------- Gonidea
| |
+--+----------- Microcondylaea
|
| +-------- Koreosolenaia
+--+
| +----- Parvasolenaia
+--+
| +-- Postolata
+--+
+-- Ptychorhynchus
Zhang et al., 2004 [Publ 2735], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 13 | 1 | 1 | -- | +-- Lamprotulini
--+
+-- Sinohyriopsis **
Xue et al., 2016 [Publ 3659], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 14 | 1 | 1 | -- | +-- Sinohyriopsis **
--+
+-- Lamprotulini
Wu et al., 2024 [Publ 4388], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 35 | 33 | 5 | -- | +-------------- Chamberlainia **
--+
| +----------- Rectidentini
+--+
| +-------- Lamprotulini
+--+
| +----- Gonideini
+--+
| +-- Schepmaniini
+--+
+-- Pseudodontini
Xia et al., 2016 [Publ 3712], fig. 7B iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 2 | 0 | 0 | -- | -- Sinohyriopsis_cumingii **
Huang et al., 2002 [Publ 2657], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 33 | 3 | 2 | -- | -- Sinohyriopsis_cumingii **
Burzynski & Soroka, 2018 [Publ 3862], fig. 5a iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 19 | 4 | 2 | -- | +----- Sinohyriopsis **
--+
| +-- Gonideini
+--+
+-- Lamprotulini
Chen et al., 2012 [Publ 3624], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 6 | 1 | 1 | -- | +-- Sinohyriopsis **
--+
+-- Lamprotulini
Sano et al., 2017 [Publ 3799], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 20 | 3 | 2 | -- | +-------- Obovalis
--+
| +----- Inversidens
+--+
| +-- Lamprotulini
+--+
+-- Sinohyriopsis **
Mabuchi & Nishida, 2020 [Publ 4120], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 11 | 3 | 2 | -- | +-------- Obovalis
--+
| +----- Inversidens
+--+
| +-- Sinohyriopsis **
+--+
+-- Lamprotulini
Huang et al., 2002 [Publ 2657], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 33 | 3 | 2 | -- | -- Sinohyriopsis_cumingii **
Smith, 2021 [Publ 4331], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 19 | 2 | 2 | -- | +----- Sinohyriopsis **
--+
| +-- Rectidentini
+--+
+-- Lamprotulini
Zieritz et al., 2021 [Publ 4218], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 30 | 26 | 5 | -- | +-------------- Chamberlainia **
--+
| +----------- Lamprotulini
+--+
| +-------- Rectidentini
+--+
| +----- Gonideini
+--+
| +-- Schepmaniini
+--+
+-- Pseudodontini
Ouyang et al., 2015 [Publ 4493], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 30 | 5 | 2 | -- | -- Sinohyriopsis_cumingii **
Jiang et al., 2010 [Publ 3438], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 3 | 1 | 1 | -- | +-- Lamprotulini
--+
+-- Sinohyriopsis **
Liu et al., 2024 [Publ 4380], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 35 | 33 | 5 | -- | +-- Schepmaniini
+--------+
| +-- Pseudodontini
--+
| +-------- Gonideini
+--+
| +----- Rectidentini
+--+
| +-- Chamberlainia **
+--+
+-- Lamprotulini
Kim et al., 2021 [Publ 4197], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 17 | 0 | 0 | -- | -- Sinohyriopsis_schlegelii **
Wu et al., 2018 [Publ 3836], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 52 | 8 | 2 | -- | -- Sinohyriopsis_cumingii **
Zieritz et al., 2020 [Publ 4102], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 30 | 30 | 4 | -- | +----------- Chamberlainia **
--+
| +-------- Lamprotulini
+--+
| +----- Gonideini
+--+
| +-- Pseudodontini
+--+
+-- Rectidentini
Ouyang et al., 2015 [Publ 4493], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 30 | 5 | 2 | -- | +----- Lamprotulini
--+
| +-- Sinohyriopsis **
+--+
+-- Gonideini
Wu et al., 2024 [Publ 4350], fig. 5b iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 35 | 33 | 5 | -- | +----------------------------- Schepmaniini
|
--+ +-------------------------- Pseudodontini
| |
+--+ +----------------------- Rectidentini
| |
+--+ +-------------------- Inversidens
| |
| | +----- Lamprotula
+--+ +-----------+
| | | +-- Chamberlainia **
| | +--+
+--+ +-- Potomida
|
| +-------------- Sinosolenaia
| |
+--+ +-- Gonidea
| +--------+
| | +-- Microcondylaea
+--+
| +-------- Koreosolenaia
+--+
| +----- Parvasolenaia
+--+
| +-- Postolata
+--+
+-- Ptychorhynchus
Breton et al., 2011 [Publ 3376], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 37 | 1 | 1 | -- | -- Sinohyriopsis_cumingii **
Wu et al., 2024 [Publ 4350], fig. 5a iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 35 | 33 | 5 | -- | +-------------------------- Chamberlainia **
|
--+ +----------------------- Rectidentini
| |
| | +-- Schepmaniini
+--+ +-----------------+
| | +-- Pseudodontini
| |
+--+ +-- Inversidens
| +--------------+
| | +-- Lamprotulini
+--+
| +-------------- Sinosolenaia
| |
+--+ +-- Gonidea
| +--------+
| | +-- Microcondylaea
+--+
| +-------- Parvasolenaia
+--+
| +----- Koreosolenaia
+--+
| +-- Postolata
+--+
+-- Ptychorhynchus
Plazzi & Passamonti, 2010 [Publ 3354], fig. 2-3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 3 | 1 | 1 | -- | +-- Sinohyriopsis **
--+
+-- Lamprotulini
Campbell et al., 2005 [Publ 2660], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 101 | 3 | 2 | -- | -- Sinohyriopsis_cumingii **
|
