The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)
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Bivalvia | Unionida | Unionoidea | Unionidae | Parreysiinae tribe COELATURINI Modell, 1942: cladomics
10 genera Cladomics Summarynumber of species = 37 (41 total)
number of subtaxa (rank: genus) = 10
analyzed in 55 cladograms in 36 publications
maximum number of terminals in an analysis =
27 (mean = 3.91, median = 2)
maximum number of subtaxa (genus) =
7 (mean = 2.13, median = 2) Basic Monophylynumber of cladograms that test basic monophyly
(i.e., more than 1 included species) = 29
maximum number of included terminals = 27, (mean =
6.52 median = 4)
maximum number of subtaxa (genus) =
7 (mean = 3.14, median = 3)
proportion of cladograms that support basic monophyly
= 1 Rigorous Monophylynumber of cladograms that test rigorous monophyly = 28
(Rigorous monophyly refers to cladograms with terminal species representing
more than one subtaxon (genus) taxon and at least one rigorous outgroup (family).
maximum number of included terminals = 27 (mean = 6.36, median = 4)
maximum number of subtaxa (genus) =
7 (mean = 3.18, median = 3)
proportion of cladograms that support rigorous monophyly = 1
Pairwise Cladogram ClusteringThe graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.
Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14 ** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram ** iT| tT | bi | riS | oT | roT | roS | iD | | 27 | 27 | 31 | 7 | 605 | 58 | 4 | 0 | +-------------------------------------------------- Indochinellini
|
--+ +----------------------------------------------- Parreysiini
| |
| | +----- Nitia_monceti__2 **
+--+ +--------------------------------------+
| | | +-- Nitia_teretiuscula **
| | +--+
| | +-- Nitia_monceti__1 **
+--+
| +----------------------------------------- Grandidieria **
| |
| | +----- Pseudospatha_tanganyicensis__2 **
| | +--------------------------------+
+--+ | | +-- Moncetia **
| | +--+
| | +-- Pseudospatha_tanganyicensis__1 **
| |
| | +----- Coelatura_leopoldvillensis__2 **
+--+ +-----------------------------+
| | | +-- Coelatura_leopoldvillensis__1 **
| | +--+
| | +-- Coelatura_mesafricana **
| |
| | +-------------------------------- Coelatura_aegyptiaca__3 **
| | |
+--+ | +-------- Coelatura_hauttecoeuri__2 **
| | |
| | +-------- Nitia_acuminata **
| | |
| | | +----- Coelatura_aegyptiaca__2 **
| | +--------------------+ |
| | | | +----- Coelatura_alluaudi__2 **
+--+ | | |
| | | +----- Coelatura_hauttecoeuri__1 **
| | | |
| | +--+----- Coelatura_stuhlmanni__2 **
| | |
| | | +-- Coelatura_aegyptiaca__1 **
| | | |
| | | +-- Coelatura_stuhlmanni__1 **
| | | |
| | +--+-- Coelatura_alluaudi__1 **
+--+ |
| +-- Coelatura_bakeri__1 **
| |
| +-- Coelatura_bakeri__2 **
|
| +----------- Coelatura_horei__3 **
| |
| +--------------+ +-------- Coelatura_ratidota **
| | | |
| | +--+ +----- Coelatura__2 **
| | | |
| | +--+ +-- Coelatura_horei__1 **
| | | |
+--+ +--+-- Coelatura__1 **
| |
| +-- Coelatura_horei__2 **
|
| +----------------------- Coelatura_gabonensis **
| |
+--+ +-------------------- Coelatura_stanleyvillensis **
| |
| | +----------------- Coelatura_kunenensis **
+--+ |
| | +-------------- Nyassunio **
| | |
+--+ | +----------- Coelatura_hypsiprymna__2 **
| | |
| | +----------- Coelatura_mossambicensis__1 **
| | |
+--+ | +-- Coelatura_hypsiprymna__3 **
| +--------+
| | +-- Coelatura_mossambicensis__2 **
| |
| | +----- Coelatura_luapulaensis__2 **
| | |
| | +----- Coelatura_choziensis__2 **
+--+-----+
| +----- Coelatura_hypsiprymna__1 **
| |
| | +-- Coelatura_choziensis__1 **
| +--+
| +-- Coelatura_luapulaensis__1 **
|
| +----- Coelatura_briarti **
| +--+
| | | +-- Prisodontopsis_aviculaeformis__1 **
| | +--+
+--+ +-- Prisodontopsis_mweruensis__1 **
|
| +-- Prisodontopsis_aviculaeformis__2 **
| |
+-----+-- Prisodontopsis_aviculaeformis__3 **
|
+-- Prisodontopsis_mweruensis__2 **
Ortiz-Sepulveda et al., 2020 [Publ 4101], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 27 | 27 | 16 | 7 | 9 | 9 | 2 | 41 | +-------------------------------------- Leoparreysiini
|
--+ +----------------------------------- Indochinellini
| |
+--+ +-------------------------------- Grandidieria **
| |
| | +-- Pseudospatha **
+--+ +--------------------------+
| | +-- Moncetia **
| |
| | +-------------------------- Nitia **
+--+ |
| | +-- Coelatura_aegyptiaca **
| | |
| | +-- Coelatura_alluaudi **
+--+ |
| +--------------------+-- Coelatura_bakeri **
| | |
| | +-- Coelatura_hauttecoeuri **
| | |
| | +-- Coelatura_stuhlmanni **
+--+
| +-- Coelatura_leopoldvillensis **
| +-----------------+
| | +-- Coelatura_mesafricana **
| |
| | +-- Coelatura_horei **
+--+ |
| +--------------+-- Coelatura_ratidota **
| | |
| | +-- Coelatura **
+--+
| +-------------- Coelatura_gabonensis **
| |
| +-------------- Coelatura_stanleyvillensis **
+--+
| +----------- Coelatura_kunenensis **
| |
| | +----- Prisodontopsis_aviculaeformis **
+--+ +--+
| | | +-- Coelatura_briarti **
| | +--+
| | +-- Prisodontopsis_mweruensis **
+--+
| +-- Coelatura_choziensis **
| |
| +-- Coelatura_hypsiprymna **
| |
+-----+-- Coelatura_luapulaensis **
|
+-- Coelatura_mossambicensis **
|
+-- Nyassunio **
Ortiz-Sepulveda et al., 2020 [Publ 4101], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 27 | 27 | 16 | 7 | 9 | 9 | 2 | 41 | +-------------------------------- Leoparreysiini
|
| +----------------------------- Indochinellini
--+ |
| | +----- Grandidieria **
| | +--------------------+
+--+ | | +-- Pseudospatha **
| | +--+
| | +-- Moncetia **
| |
+--+ +----------------------- Nitia **
| |
| | +-- Coelatura_aegyptiaca **
| | |
| | +-- Coelatura_alluaudi **
+--+ |
| +-----------------+-- Coelatura_bakeri **
| | |
| | +-- Coelatura_hauttecoeuri **
| | |
| | +-- Coelatura_stuhlmanni **
+--+
| +-- Coelatura_leopoldvillensis **
| +--------------+
| | +-- Coelatura_mesafricana **
| |
| | +-- Coelatura_horei **
+--+ |
| +-----------+-- Coelatura_ratidota **
| | |
| | +-- Coelatura **
| |
+--+ +-- Coelatura_gabonensis **
| +--------+
| | +-- Coelatura_stanleyvillensis **
| |
+--+ +-------- Coelatura_kunenensis **
| |
| | +-- Coelatura_briarti **
| | |
+--+ +--+-- Prisodontopsis_mweruensis **
| | |
| | +-- Prisodontopsis_aviculaeformis **
| |
+--+ +-- Coelatura_choziensis **
| |
| +-- Coelatura_hypsiprymna **
| |
+--+-- Coelatura_luapulaensis **
|
+-- Coelatura_mossambicensis **
|
+-- Nyassunio **
Galal-Khallaf et al., 2024 [Publ 4459], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 11 | 11 | 12 | 2 | 17 | 0 | 0 | 40 | +----------------- Coelatura_horei **
|
| +-- Coelatura_gabonensis **
| +--------+
| | +-- Coelatura_aegyptiaca__4 **
| |
| +--+ +-------- Coelatura_kunenensis **
--+ | | |
| | | | +----- Coelatura_luapulaensis__1 **
| | +--+ |
| | | +----- Coelatura_luapulaensis__3 **
| | | |
| | +--+----- Coelatura_luapulaensis__4 **
| | |
| | +----- Coelatura_choziensis__2 **
+--+ |
| | +-- Coelatura_luapulaensis__2 **
| +--+
| +-- Coelatura_choziensis__1 **
|
| +----------- Coelatura_alluaudi__2 **
| |
| +----------- Nitia **
| |
| | +-------- Coelatura_aegyptiaca__1 **
+--+ |
| +-------- Coelatura_bakeri__2 **
| |
| | +-- Coelatura_aegyptiaca__2 **
| +-----+
+--+ +-- Coelatura_stuhlmanni__1 **
|
| +-- Coelatura_stuhlmanni__2 **
+-----+
| +-- Coelatura_bakeri__1 **
|
| +----- Coelatura_aegyptiaca__3 **
+--+
| +-- Coelatura_alluaudi__1 **
+--+
+-- Coelatura_hauttecoeuri **
Graf et al., 2014 [Publ 3591], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 8 | 8 | 11 | 3 | 3 | 1 | 1 | 34 | +----------------------- Leoparreysiini
|
--+ +-------------------- Nitia **
| |
+--+ +----------------- Coelatura_aegyptiaca__2 **
| |
+--+ +-------------- Coelatura_aegyptiaca__1 **
| |
+--+ +----------- Coelatura_gabonensis **
| |
+--+ +-------- Coelatura_kunenensis **
| |
| | +----- Prisodontopsis **
+--+ |
| | +-- Coelatura_choziensis__1 **
| | |
+--+ +-- Coelatura_choziensis__2 **
| |
| +-- Coelatura_luapulaensis__1 **
| |
+--+-- Coelatura_luapulaensis__2 **
|
+-- Coelatura_luapulaensis__3 **
|
+-- Coelatura_luapulaensis__4 **
|
+-- Coelatura_luapulaensis__5 **
Bolotov et al., 2017 [Publ 3764], fig. S1 iT| tT | bi | riS | oT | roT | roS | iD | | 8 | 8 | 0 | 3 | 117 | 30 | 4 | 45 | +-------------- Trapezidens
|
+--------+ +----------- Lamellidens_lamellatus
| | |
| +--+ +-------- Lamellidens_jenkinsianus
| | |
| | | +----- Lamellidens_generosus
| +--+ |
| | +----- Lamellidens_mainwaringi
| | |
| +--+ +-- Lamellidens_ferrugineus
| +--+
--+ | +-- Lamellidens_marginalis
| |
| | +-- Lamellidens_savadiensis
| +--+
| +-- Lamellidens_corrianus__1
|
| +-------------------- Parreysiini
| |
| | +----------------- Leoparreysiini
| | |
| | | +-- Nitia **
+--+ | |
| | +-- Coelatura_gabonensis **
| | |
| | +-- Prisodontopsis_mweruensis **
| | |
+--+ +-- Coelatura_kunenensis **
| |
| +-----------+-- Coelatura_aegyptiaca__1 **
| | |
| | +-- Coelatura_aegyptiaca__2 **
| | |
| | +-- Prisodontopsis_aviculaeformis **
| | |
| | +-- Coelatura_luapulaensis **
+--+ |
| +-- Coelatura_choziensis **
|
| +----- Indochinella
| +-----+
| | | +-- Scabies
| | +--+
+--+ +-- Radiatula
|
| +-------- Indonaia_shurtleffiana
| |
+--+ +----- Indonaia_rugosa
| |
| | +-- Indonaia_andersoniana
+--+--+
| +-- Indonaia_caerulea
|
| +-- Indonaia_bonneaudii
+--+
+-- Lamellidens_corrianus__2
Bolotov et al., 2017 [Publ 3761], fig. S1 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 4 | 3 | 134 | 24 | 4 | 44 | +-------------------------- Lamellidentini
--+
| +----------------------- Parreysiini
+--+
| +-------------------- Leoparreysiini
+--+
| +----------------- Indochinellini
+--+
| +-------------- Nitia **
+--+
| +----------- Coelatura_aegyptiaca__2 **
+--+
| +-------- Coelatura_aegyptiaca__1 **
+--+
| +----- Prisodontopsis **
+--+
| +-- Coelatura_gabonensis **
+--+
+-- Coelatura_kunenensis **
Huang et al., 2019 [Publ 3843], fig. S3 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 4 | 3 | 194 | 29 | 4 | 46 | +----------------------- Lamellidentini
|
--+ +-------------------- Parreysiini
| |
+--+ +----------------- Leoparreysiini
| |
+--+ +-------------- Indochinellini
| |
+--+ +----------- Coelatura_aegyptiaca__2 **
| |
+--+ +-- Coelatura_aegyptiaca__1 **
| +-----+
| | +-- Nitia **
+--+
| +----- Coelatura_kunenensis **
+--+
| +-- Coelatura_gabonensis **
+--+
+-- Prisodontopsis **
Huang et al., 2019 [Publ 3843], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 2 | 3 | 196 | 29 | 4 | 44 | +-------------------- Lamellidentini
|
--+ +----------------- Parreysiini
| |
+--+ +-------------- Leoparreysiini
| |
+--+ +----------- Indochinellini
| |
+--+ +-------- Coelatura_aegyptiaca__2 **
| |
+--+ +----- Coelatura_aegyptiaca__1 **
| |
| +----- Nitia **
+--+
| +-- Coelatura_gabonensis **
| |
+--+-- Coelatura_kunenensis **
|
+-- Prisodontopsis **
Huang et al., 2019 [Publ 3843], fig. S4 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 2 | 3 | 196 | 29 | 4 | 43 | +----------------- Parreysiini
|
+----------------- Lamellidentini
--+
| +-------------- Leoparreysiini
| |
+--+ +----------- Indochinellini
| |
+--+ +-------- Coelatura_aegyptiaca__2 **
| |
| +-------- Nitia **
+--+
| +----- Coelatura_aegyptiaca__1 **
| |
+--+ +-- Coelatura_gabonensis **
| |
+--+-- Coelatura_kunenensis **
|
+-- Prisodontopsis **
Bolotov et al., 2017 [Publ 3761], fig. S2 iT| tT | bi | riS | oT | roT | roS | iD | | 5 | 5 | 4 | 3 | 134 | 24 | 4 | 42 | +-- Parreysiini
+--------------------+
| +-- Lamellidentini
--+
| +-------------------- Leoparreysiini
+--+
| +----------------- Indochinellini
+--+
| +-------------- Nitia **
+--+
| +----------- Coelatura_aegyptiaca__1 **
+--+
| +-------- Coelatura_aegyptiaca__2 **
+--+
| +----- Coelatura_gabonensis **
+--+
| +-- Coelatura_kunenensis **
+--+
+-- Prisodontopsis **
Whelan et al., 2011 [Publ 3380], fig. 1-ML iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 3 | 3 | 32 | 5 | 3 | 43 | +-------------------- Lamellidentini
--+
| +----------------- Leoparreysiini
+--+
| +-------------- Indochinellini
+--+
| +----------- Nitia **
+--+
| +-------- Coelatura_aegyptiaca__2 **
+--+
| +----- Coelatura_aegyptiaca__1 **
+--+
| +-- Coelatura_gabonensis **
+--+
+-- Prisodontopsis **
Whelan et al., 2011 [Publ 3380], fig. 1-MP iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 3 | 32 | 5 | 3 | 43 | +----------------- Lamellidentini
|
--+ +-------------- Leoparreysiini
| |
+--+ +----------- Indochinellini
| |
+--+ +-------- Nitia **
| |
+--+ +----- Coelatura_aegyptiaca__1 **
| |
+--+----- Coelatura_aegyptiaca__2 **
|
| +-- Coelatura_gabonensis **
+--+
+-- Prisodontopsis **
Bolotov et al., 2024 [Publ 4369], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 3 | 58 | 58 | 4 | 43 | +----------------- Lamellidentini
|
--+ +-- Leoparreysiini
| +-----------+
| | +-- Parreysiini
+--+
| +----------- Indochinellini
+--+
| +-------- Nitia **
+--+
| +----- Coelatura_aegyptiaca **
+--+
| +-- Coelatura_kunenensis **
+--+
+-- Nyassunio **
Bolotov et al., 2022 [Publ 4252], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 3 | 57 | 53 | 4 | 43 | +-------------------- Lamellidentini
--+
| +----------------- Parreysiini
+--+
| +-------------- Leoparreysiini
+--+
| +----------- Indochinellini
+--+
| +-------- Nitia **
+--+
| +----- Coelatura_aegyptiaca **
+--+
| +-- Nyassunio **
+--+
+-- Coelatura_kunenensis **
Bolotov et al., 2022 [Publ 4239], fig. S2 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 3 | 54 | 50 | 4 | 43 | +-------------------- Parreysiini
--+
| +----------------- Lamellidentini
+--+
| +-------------- Leoparreysiini
+--+
| +----------- Indochinellini
+--+
| +-------- Nitia **
+--+
| +----- Coelatura_aegyptiaca **
+--+
| +-- Coelatura_kunenensis **
+--+
+-- Nyassunio **
Bolotov et al., 2024 [Publ 4369], fig. S1 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 3 | 59 | 58 | 4 | 43 | +-------------------- Parreysia
|
--+ +-- Leoparreysiini
| +--------------+
| | +-- Lamellidentini
+--+
| +-------------- Assamnaia
+--+
| +----------- Indochinellini
+--+
| +-------- Nitia **
+--+
| +----- Coelatura_aegyptiaca **
+--+
| +-- Nyassunio **
+--+
+-- Coelatura_kunenensis **
Bolotov et al., 2022 [Publ 4239], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 3 | 54 | 50 | 4 | 43 | +----------------- Lamellidentini
|
--+ +-- Parreysiini
| +-----------+
| | +-- Leoparreysiini
+--+
| +----------- Indochinellini
+--+
| +-------- Nitia **
+--+
| +----- Coelatura_aegyptiaca **
+--+
| +-- Coelatura_kunenensis **
+--+
+-- Nyassunio **
Bolotov et al., 2022 [Publ 4252], fig. S1 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 0 | 3 | 57 | 53 | 4 | 45 | +-------------- Parreysiini
|
--+ +----------- Lamellidentini
| |
+--+ +-------- Leoparreysiini
| |
+--+ +----- Indochinellini
| |
+--+ +-- Coelatura_aegyptiaca **
| |
| +-- Coelatura_kunenensis **
+--+
+-- Nitia **
|
+-- Nyassunio **
Bolotov et al., 2022 [Publ 4239], fig. S1 iT| tT | bi | riS | oT | roT | roS | iD | | 4 | 4 | 2 | 3 | 54 | 50 | 4 | 43 | +-------------------- Lamellidentini
--+
| +----------------- Parreysiini
+--+
| +-------------- Leoparreysiini
+--+
| +----------- Indochinellini
+--+
| +-------- Nitia **
+--+
| +----- Coelatura_aegyptiaca **
+--+
| +-- Coelatura_kunenensis **
+--+
+-- Nyassunio **
Chen et al., 2026 [Publ 4833], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 24 | 21 | 4 | 44 | +----- Nitia **
--+
| +-- Coelatura **
+--+
+-- Prisodontopsis **
Pfeiffer et al., 2018 [Publ 3827], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 3 | 3 | 1 | 3 | 25 | 24 | 4 | 44 | +----------------- Lamellidentini
--+
| +-------------- Parreysiini
+--+
| +----------- Leoparreysiini
+--+
| +-------- Indochinellini
+--+
| +----- Nitia **
+--+
| +-- Coelatura **
+--+
+-- Prisodontopsis **
Bolotov et al., 2018 [Publ 3826], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 80 | 22 | 4 | -- | +-------------- Lamellidentini
--+
| +----------- Parreysiini
+--+
| +-------- Leoparreysiini
+--+
| +----- Indochinellini
+--+
| +-- Coelatura **
+--+
+-- Nitia **
Zieritz et al., 2024 [Publ 4381], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 43 | 3 | 3 | -- | +----------- Lamellidentini
--+
| +-------- Leoparreysiini
+--+
| +----- Indochinellini
+--+
| +-- Coelatura **
+--+
+-- Nitia **
Pfeiffer et al., 2019 [Publ 3937], fig. 4-5 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 55 | 8 | 3 | -- | +----------- Lamellidentini
--+
| +-------- Leoparreysiini
+--+
| +----- Indochinellini
+--+
| +-- Coelatura **
+--+
+-- Prisodontopsis **
Bolotov et al., 2020 [Publ 4100], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 115 | 39 | 4 | -- | +----------- Lamellidentini
|
--+ +-- Parreysiini
| +-----+
| | +-- Leoparreysiini
+--+
| +----- Indochinellini
+--+
| +-- Coelatura **
+--+
+-- Nitia **
Pfeiffer et al., 2019 [Publ 3937], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 55 | 8 | 3 | -- | +----------- Lamellidentini
--+
| +-------- Leoparreysiini
+--+
| +----- Indochinellini
+--+
| +-- Coelatura **
+--+
+-- Prisodontopsis **
Bolotov et al., 2019 [Publ 3974], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 16 | 11 | 2 | -- | +-------- Leoparreysiini
--+
| +----- Indochinellini
+--+
| +-- Nitia **
+--+
+-- Coelatura **
Bolotov et al., 2020 [Publ 4100], fig. S1 iT| tT | bi | riS | oT | roT | roS | iD | | 2 | 2 | 0 | 2 | 109 | 38 | 4 | -- | +----------- Lamellidentini
|
--+ +-- Parreysiini
| +-----+
| | +-- Leoparreysiini
+--+
| +----- Indochinellini
+--+
| +-- Coelatura **
+--+
+-- Nitia **
Dai et al., 2025 [Publ 4440], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 43 | 2 | 2 | -- | +----- Parreysiini
--+
| +-- Coelatura **
+--+
+-- Indochinellini
Hoeh et al., 2009 [Publ 2716], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 46 | 0 | 0 | -- | -- Coelatura_aegyptiaca **
Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 121 | 2 | 2 | -- | +-- Coelatura **
|
--+-- Leoparreysiini
|
+-- Indochinellini
Roe & Hoeh, 2003 [Publ 2655], fig. 4f3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 30 | 0 | 0 | -- | -- Coelatura_aegyptiaca **
Hoeh et al., 2002 [Publ 2653], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 30 | 0 | 0 | -- | -- Coelatura_aegyptiaca **
Hoeh et al., 2009 [Publ 2716], fig. 4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 45 | 0 | 0 | -- | -- Coelatura_aegyptiaca **
Chen et al., 2025 [Publ 4487], fig. 3 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 40 | 2 | 2 | -- | +----- Parreysiini
--+
| +-- Coelatura **
+--+
+-- Indochinellini
Dai et al., 2025 [Publ 4440], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 43 | 2 | 2 | -- | +-- Coelatura **
|
--+-- Indochinellini
|
+-- Parreysiini
Pfeiffer & Graf, 2013 [Publ 3531], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 30 | 1 | 1 | -- | +-- Lamellidentini
--+
+-- Coelatura **
Jeratthitikul et al., 2019 [Publ 3922], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 22 | 22 | 4 | -- | +----------- Lamellidentini
--+
| +-------- Leoparreysiini
+--+
| +----- Parreysiini
+--+
| +-- Coelatura **
+--+
+-- Indochinellini
Bogan & Hoeh, 2000 [Publ 2650], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 33 | 0 | 0 | -- | -- Coelatura_aegyptiaca **
Graf & Cummings, 2006 [Publ 1968], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 46 | 0 | 0 | -- | -- Coelatura_aegyptiaca **
Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 80 | 3 | 3 | -- | +-------- Lamellidentini
--+
| +----- Leoparreysiini
+--+
| +-- Coelatura **
+--+
+-- Indochinellini
Hoeh et al., 2001 [Publ 2652], fig. 14f4 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 30 | 0 | 0 | -- | -- Coelatura_aegyptiaca **
Araujo et al., 2017 [Publ 3801], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 13 | 1 | 1 | -- | +-- Lamellidentini
--+
+-- Coelatura **
Hoeh et al., 2009 [Publ 2716], fig. 5 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 45 | 0 | 0 | -- | -- Coelatura_aegyptiaca **
Wu et al., 2021 [Publ 4366], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 90 | 12 | 3 | -- | +-------- Lamellidentini
--+
| +----- Leoparreysiini
+--+
| +-- Coelatura **
+--+
+-- Indochinellini
Chen et al., 2023 [Publ 4302], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 40 | 2 | 2 | -- | +-- Coelatura **
|
--+-- Indochinellini
|
+-- Parreysiini
Konopleva et al., 2017 [Publ 3807], fig. 6 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 30 | 10 | 3 | -- | +-------- Lamellidentini
--+
| +----- Leoparreysiini
+--+
| +-- Coelatura **
+--+
+-- Indochinellini
Wu et al., 2021 [Publ 4366], fig. 2_I iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 90 | 12 | 3 | -- | +-------- Lamellidentini
--+
| +----- Leoparreysiini
+--+
| +-- Coelatura **
+--+
+-- Indochinellini
Roe & Hoeh, 2003 [Publ 2655], fig. 4f2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 30 | 0 | 0 | -- | -- Coelatura_aegyptiaca **
Graf, 2000 [Publ 2647], fig. 1 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 16 | 1 | 1 | -- | Pfeiffer & Graf, 2015 [Publ 3620], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 39 | 7 | 3 | -- | +-------- Lamellidentini
--+
| +----- Leoparreysiini
+--+
| +-- Coelatura **
+--+
+-- Indochinellini
Chen et al., 2023 [Publ 4302], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 40 | 2 | 2 | -- | +----- Parreysiini
--+
| +-- Coelatura **
+--+
+-- Indochinellini
Chen et al., 2025 [Publ 4487], fig. 2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 40 | 2 | 2 | -- | +----- Parreysiini
--+
| +-- Coelatura **
+--+
+-- Indochinellini
Hoeh et al., 2001 [Publ 2652], fig. 14f2 iT| tT | bi | riS | oT | roT | roS | iD | | 1 | 1 | 0 | 1 | 30 | 0 | 0 | -- | -- Coelatura_aegyptiaca **
|
