MUSSELpdb | subtribe Contradentina Modell, 1942 cladomics

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Page last updated
Thu 3 September 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

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subtribe CONTRADENTINA Modell, 1942: cladomics
6 genera

Cladomics Summary

number of species = 35 (35 total)
number of subtaxa (rank: genus) = 6
analyzed in 79 cladograms in 50 publications
maximum number of terminals in an analysis = 31 (mean = 6.43, median = 3)
maximum number of subtaxa (genus) = 5 (mean = 2.62, median = 2)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 70
maximum number of included terminals = 31, (mean = 7.13 median = 4.5)
maximum number of subtaxa (genus) = 5 (mean = 2.83, median = 3)
proportion of cladograms that support basic monophyly = 0.99

Rigorous Monophyly

number of cladograms that test rigorous monophyly = 62
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (genus) taxon and at least one rigorous outgroup (family).
maximum number of included terminals = 31 (mean = 6.69, median = 3.5)
maximum number of subtaxa (genus) = 5 (mean = 2.92, median = 3)
proportion of cladograms that support rigorous monophyly = 0.98

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination

NJ Tree

Pfeiffer et al., 2021 [Publ 4126], fig. 9

** Top Monophyly Cladogram **

tT	bi	riS	oT	roT	roS	iD
15	15	13	5	14	13	1	0

  +----------- Rectidentina
  |                        
--+        +-- Trapezoideus **
  |  +-----+               
  |  |     +-- Physunio **    
  +--+                     
     |  +----- Yaukthwa **    
     +--+                  
        |  +-- Solenaia **    
        +--+               
           +-- Lens **

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

** Top Sister-Group Cladogram **

tT	bi	riS	oT	roT	roS	iD
31	31	30	4	601	22	2	25

  +----------------------- Khairuloconcha     
  |                                           
--+  +-------------------- Ctenodesma         
  |  |                                        
  +--+  +----------------- Rectidentina       
     |  |                                     
     |  |              +-- Physunio_modelli **   
     +--+  +-----------+                      
        |  |           +-- Physunio_pellucidus **
        |  |                                  
        +--+           +-- Physunio_massini **   
           |  +--------+                      
           |  |        +-- Physunio_superbus **  
           +--+                               
              |  +-------- Physunio **           
              +--+                            
                 |  +----- Trapezoideus **       
                 +--+                         
                    |  +-- Yaukthwa **           
                    +--+                      
                       +-- Lens **

Bolotov et al., 2020 [Publ 4100], fig. S1

tT	bi	riS	oT	roT	roS	iD
23	23	21	4	88	12	1	18

  +----------- Rectidentina
--+                        
  |  +-------- Physunio **    
  +--+                     
     |  +----- Trapezoideus **
     +--+                  
        |  +-- Lens **        
        +--+               
           +-- Yaukthwa **

Bolotov et al., 2020 [Publ 4100], fig. 1

tT	bi	riS	oT	roT	roS	iD
23	23	21	4	94	12	1	16

  +-------- Rectidentina
  |                     
--+     +-- Trapezoideus **
  |  +--+               
  |  |  +-- Physunio **    
  +--+                  
     |  +-- Yaukthwa **    
     +--+               
        +-- Lens **

Konopleva et al., 2023 [Publ 4262], fig. 8

tT	bi	riS	oT	roT	roS	iD
19	19	17	4	63	16	1	11

  +-------- Rectidentina
  |                     
--+     +-- Physunio **    
  |  +--+               
  |  |  +-- Trapezoideus **
  +--+                  
     |  +-- Yaukthwa **    
     +--+               
        +-- Lens **

Konopleva et al., 2019 [Publ 4041], fig. 3

tT	bi	riS	oT	roT	roS	iD
19	19	17	4	15	12	1	14

  +----------- Rectidentina
--+                        
  |  +-------- Yaukthwa **    
  +--+                     
     |  +----- Lens **        
     +--+                  
        |  +-- Trapezoideus **
        +--+               
           +-- Physunio **

Konopleva et al., 2019 [Publ 3900], fig. 1

tT	bi	riS	oT	roT	roS	iD
16	16	14	4	35	11	1	17

  +----------- Rectidentina
--+                        
  |  +-------- Yaukthwa **    
  +--+                     
     |  +----- Physunio **    
     +--+                  
        |  +-- Trapezoideus **
        +--+               
           +-- Lens **

Bolotov et al., 2022 [Publ 4250], fig. S5

tT	bi	riS	oT	roT	roS	iD
15	15	12	3	0	0	0	22

  +----- Yaukthwa **    
--+                  
  |  +-- Physunio **    
  +--+               
     +-- Trapezoideus **

Zieritz et al., 2020 [Publ 4102], fig. 4

tT	bi	riS	oT	roT	roS	iD
14	14	12	4	17	12	2	15

           +-- Ctenodesmina
  +--------+               
  |        +-- Rectidentina
--+                        
  |  +-------- Physunio **    
  +--+                     
     |  +----- Yaukthwa **    
     +--+                  
        |  +-- Trapezoideus **
        +--+               
           +-- Lens **

Bolotov et al., 2017 [Publ 3761], fig. S1

tT	bi	riS	oT	roT	roS	iD
12	12	10	3	126	11	1	14

  +-------- Rectidentina
--+                     
  |  +----- Physunio **    
  +--+                  
     |  +-- Yaukthwa **    
     +--+               
        +-- Lens **

Huang et al., 2019 [Publ 3843], fig. S3

tT	bi	riS	oT	roT	roS	iD
12	12	10	3	186	11	1	14

  +-------- Rectidentina
--+                     
  |  +----- Physunio **    
  +--+                  
     |  +-- Yaukthwa **    
     +--+               
        +-- Lens **

Zieritz et al., 2024 [Publ 4381], fig. 7A

tT	bi	riS	oT	roT	roS	iD
12	12	10	2	0	0	0	13

  +-- Solenaia **
--+           
  +-- Lens **

Huang et al., 2019 [Publ 3843], fig. S4

tT	bi	riS	oT	roT	roS	iD
12	12	7	3	188	11	1	18

  +----- Rectidentina     
  |                       
  |  +-- Physunio_modelli ** 
--+  |                    
  |  +-- Physunio **         
  |  |                    
  +--+-- Physunio_superbus **
     |                    
     +-- Yaukthwa **         
     |                    
     +-- Lens **

Bolotov et al., 2022 [Publ 4250], fig. 8

tT	bi	riS	oT	roT	roS	iD
12	12	10	2	37	0	0	23

  +-- Trapezoideus **
--+               
  +-- Yaukthwa **

Bolotov et al., 2017 [Publ 3764], fig. S1

tT	bi	riS	oT	roT	roS	iD
12	12	10	3	112	11	1	14

  +-------- Rectidentina
--+                     
  |  +----- Physunio **    
  +--+                  
     |  +-- Yaukthwa **    
     +--+               
        +-- Lens **

Zieritz et al., 2026 [Publ 4812], fig. 2a

tT	bi	riS	oT	roT	roS	iD
12	12	10	2	0	0	0	13

  +-- Solenaia **
--+           
  +-- Lens **

Huang et al., 2019 [Publ 3843], fig. 2

tT	bi	riS	oT	roT	roS	iD
12	12	10	3	188	11	1	14

  +-------- Rectidentina
--+                     
  |  +----- Physunio **    
  +--+                  
     |  +-- Yaukthwa **    
     +--+               
        +-- Lens **

Bolotov et al., 2017 [Publ 3761], fig. S2

tT	bi	riS	oT	roT	roS	iD
11	11	9	3	127	11	1	17

  +-------- Rectidentina
--+                     
  |  +----- Lens **        
  +--+                  
     |  +-- Physunio **    
     +--+               
        +-- Yaukthwa **

Pfeiffer et al., 2021 [Publ 4126], fig. 12

tT	bi	riS	oT	roT	roS	iD
11	11	8	2	0	0	0	11

  +-- Solenaia **
--+           
  +-- Lens **

Jeratthitikul et al., 2021 [Publ 4212], fig. 4

tT	bi	riS	oT	roT	roS	iD
10	10	8	4	11	11	1	12

  +-------- Rectidentina
  |                     
--+     +-- Trapezoideus **
  |  +--+               
  |  |  +-- Physunio **    
  +--+                  
     |  +-- Yaukthwa **    
     +--+               
        +-- Lens **

Bolotov et al., 2018 [Publ 3826], fig. 2

tT	bi	riS	oT	roT	roS	iD
10	10	0	3	72	8	1	19

  +----- Rectidentina       
  |                         
  |  +-- Lens_eximius **       
  |  |                      
  |  +-- Physunio_modelli__1 **
  |  |                      
--+  +-- Yaukthwa_nesemanni ** 
  |  |                      
  |  +-- Yaukthwa_panhai **    
  |  |                      
  |  +-- Lens_comptus **       
  |  |                      
  +--+-- Lens_novoselovi **    
     |                      
     +-- Physunio_modelli__2 **
     |                      
     +-- Physunio **           
     |                      
     +-- Yaukthwa_inlenensis **
     |                      
     +-- Lens_contradens **    
     |                      
     +-- Lens_micropterus **

Pfeiffer et al., 2021 [Publ 4126], fig. 25

tT	bi	riS	oT	roT	roS	iD
9	9	7	2	0	0	0	20

  +-- Solenaia **
--+           
  +-- Yaukthwa **

Zieritz et al., 2024 [Publ 4381], fig. 2

tT	bi	riS	oT	roT	roS	iD
8	8	6	4	37	10	2	13

  +----------- Ctenodesmina
  |                        
--+  +-------- Rectidentina
  |  |                     
  +--+     +-- Physunio **    
     |  +--+               
     |  |  +-- Trapezoideus **
     +--+                  
        |  +-- Yaukthwa **    
        +--+               
           +-- Lens **

Pfeiffer et al., 2021 [Publ 4126], fig. 18

tT	bi	riS	oT	roT	roS	iD
8	8	7	3	0	0	0	12

  +----- Yaukthwa **    
--+                  
  |  +-- Trapezoideus **
  +--+               
     +-- Physunio **

Wu et al., 2024 [Publ 4350], fig. 6-7

tT	bi	riS	oT	roT	roS	iD
7	7	5	4	29	6	2	12

  +----------- Ctenodesmina
  |                        
--+  +-------- Rectidentina
  |  |                     
  +--+     +-- Physunio **    
     |  +--+               
     |  |  +-- Trapezoideus **
     +--+                  
        |  +-- Lens **        
        +--+               
           +-- Yaukthwa **

Wu et al., 2024 [Publ 4350], fig. 5c

tT	bi	riS	oT	roT	roS	iD
7	7	1	4	29	6	2	14

  +----- Ensidens           
  |                         
  +----- Hyriopsis_desowitzi
  |                         
  +----- Rectidens          
  |                         
  +----- Hyriopsis_bialata  
--+                         
  +----- Ctenodesmina       
  |                         
  |  +-- Physunio **           
  |  |                      
  |  +-- Yaukthwa_paiensis **  
  |  |                      
  |  +-- Yaukthwa_inlenensis **
  +--+                      
     +-- Trapezoideus **       
     |                      
     +-- Yaukthwa_elongatula **
     |                      
     +-- Lens **

Wu et al., 2024 [Publ 4388], fig. 5

tT	bi	riS	oT	roT	roS	iD
7	7	5	4	29	6	2	12

  +----------- Ctenodesmina
  |                        
--+  +-------- Rectidentina
  |  |                     
  +--+     +-- Physunio **    
     |  +--+               
     |  |  +-- Trapezoideus **
     +--+                  
        |  +-- Lens **        
        +--+               
           +-- Yaukthwa **

Zieritz et al., 2021 [Publ 4218], fig. 5

tT	bi	riS	oT	roT	roS	iD
7	7	5	4	24	7	2	14

  +-------------- Ctenodesmina
--+                           
  |  +----------- Rectidentina
  +--+                        
     |  +-------- Physunio **    
     +--+                     
        |  +----- Trapezoideus **
        +--+                  
           |  +-- Lens **        
           +--+               
              +-- Yaukthwa **

Liu et al., 2024 [Publ 4380], fig. 5

tT	bi	riS	oT	roT	roS	iD
7	7	5	4	29	6	2	12

  +----------- Ctenodesmina
  |                        
--+  +-------- Rectidentina
  |  |                     
  +--+     +-- Physunio **    
     |  +--+               
     |  |  +-- Trapezoideus **
     +--+                  
        |  +-- Lens **        
        +--+               
           +-- Yaukthwa **

Wu et al., 2024 [Publ 4350], fig. 5b

tT	bi	riS	oT	roT	roS	iD
7	7	5	4	29	6	2	14

  +----------------------- Hyriopsis_bialata  
  |                                           
--+                    +-- Rectidens          
  |  +-----------------+                      
  |  |                 +-- Ctenodesmina       
  +--+                                        
     |  +----------------- Ensidens           
     |  |                                     
     +--+  +-------------- Hyriopsis_desowitzi
        |  |                                  
        +--+  +----------- Yaukthwa_elongatula **
           |  |                               
           +--+  +-------- Trapezoideus **       
              |  |                            
              +--+     +-- Physunio **           
                 |  +--+                      
                 |  |  +-- Yaukthwa_inlenensis **
                 +--+                         
                    |  +-- Yaukthwa_paiensis **  
                    +--+                      
                       +-- Lens **

Wu et al., 2024 [Publ 4350], fig. 5a

tT	bi	riS	oT	roT	roS	iD
7	7	5	4	29	6	2	12

  +----------- Ctenodesmina
  |                        
--+  +-------- Rectidentina
  |  |                     
  +--+     +-- Physunio **    
     |  +--+               
     |  |  +-- Trapezoideus **
     +--+                  
        |  +-- Lens **        
        +--+               
           +-- Yaukthwa **

Jeratthitikul et al., 2019 [Publ 3911], fig. 4

tT	bi	riS	oT	roT	roS	iD
6	6	11	2	3	1	1	16

  +----- Rectidentina
--+                  
  |  +-- Physunio **    
  +--+               
     +-- Lens **

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

tT	bi	riS	oT	roT	roS	iD
5	5	3	5	52	3	1	10

  +----------- Rectidentina
  |                        
--+        +-- Trapezoideus **
  |  +-----+               
  |  |     +-- Physunio **    
  +--+                     
     |  +----- Yaukthwa **    
     +--+                  
        |  +-- Lens **        
        +--+               
           +-- Solenaia **

Pfeiffer et al., 2019 [Publ 3937], fig. 6

tT	bi	riS	oT	roT	roS	iD
5	5	3	5	52	3	1	10

  +----------- Rectidentina
  |                        
--+        +-- Trapezoideus **
  |  +-----+               
  |  |     +-- Physunio **    
  +--+                     
     |  +----- Yaukthwa **    
     +--+                  
        |  +-- Lens **        
        +--+               
           +-- Solenaia **

Bolotov et al., 2019 [Publ 3974], fig. 1

tT	bi	riS	oT	roT	roS	iD
5	5	3	1	13	0	0	19

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

tT	bi	riS	oT	roT	roS	iD
4	4	2	2	76	7	1	13

  +----- Rectidentina
--+                  
  |  +-- Lens **        
  +--+               
     +-- Physunio **

Pfeiffer et al., 2018 [Publ 3827], fig. 1

tT	bi	riS	oT	roT	roS	iD
4	4	1	3	3	3	1	13

  +----- Rectidentina
  |                  
--+  +-- Physunio **    
  |  |               
  +--+-- Trapezoideus **
     |               
     +-- Lens **

Wu et al., 2021 [Publ 4366], fig. 6

tT	bi	riS	oT	roT	roS	iD
4	4	2	3	87	4	1	14

  +-------- Rectidentina
--+                     
  |  +----- Physunio **    
  +--+                  
     |  +-- Lens **        
     +--+               
        +-- Yaukthwa **

Wu et al., 2021 [Publ 4366], fig. 2_I

tT	bi	riS	oT	roT	roS	iD
4	4	2	3	87	4	1	14

  +-------- Rectidentina
--+                     
  |  +----- Physunio **    
  +--+                  
     |  +-- Lens **        
     +--+               
        +-- Yaukthwa **

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

tT	bi	riS	oT	roT	roS	iD
3	5	0	3	119	2	1	16

  +-- Lens **     
  |            
  +-- Ensidens 
  |            
--+-- Physunio ** 
  |            
  +-- Rectidens
  |            
  +-- Yaukthwa **

Dai et al., 2025 [Publ 4441], fig. 1

tT	bi	riS	oT	roT	roS	iD
3	3	1	2	44	4	2	12

  +-------- Ctenodesmina
--+                     
  |  +----- Rectidentina
  +--+                  
     |  +-- Physunio **    
     +--+               
        +-- Lens **

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

tT	bi	riS	oT	roT	roS	iD
3	3	1	2	37	1	1	14

  +----- Rectidentina
--+                  
  |  +-- Lens **        
  +--+               
     +-- Physunio **

Dai et al., 2024 [Publ 4389], fig. 3

tT	bi	riS	oT	roT	roS	iD
3	3	1	2	43	4	2	12

  +-------- Ctenodesmina
--+                     
  |  +----- Rectidentina
  +--+                  
     |  +-- Physunio **    
     +--+               
        +-- Lens **

Dai et al., 2023 [Publ 4285], fig. 3-ML

tT	bi	riS	oT	roT	roS	iD
3	3	1	2	42	2	1	12

  +----- Rectidentina
--+                  
  |  +-- Physunio **    
  +--+               
     +-- Lens **

Chen et al., 2025 [Publ 4663], fig. 1

tT	bi	riS	oT	roT	roS	iD
3	3	1	2	45	4	2	12

  +-------- Ctenodesmina
--+                     
  |  +----- Rectidentina
  +--+                  
     |  +-- Physunio **    
     +--+               
        +-- Lens **

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	73	3	1	12

  +-------- Rectidentina
--+                     
  |  +----- Lens **        
  +--+                  
     |  +-- Trapezoideus **
     +--+               
        +-- Physunio **

Jeratthitikul et al., 2021 [Publ 4219], fig. 1

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	31	3	1	12

  +-------- Rectidentina
--+                     
  |  +----- Lens **        
  +--+                  
     |  +-- Trapezoideus **
     +--+               
        +-- Physunio **

Jeratthitikul et al., 2021 [Publ 4219], fig. 2

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	32	3	1	12

  +-------- Rectidentina
--+                     
  |  +----- Lens **        
  +--+                  
     |  +-- Trapezoideus **
     +--+               
        +-- Physunio **

Konopleva et al., 2017 [Publ 3807], fig. 6

tT	bi	riS	oT	roT	roS	iD
3	3	1	2	28	1	1	14

  +----- Rectidentina
--+                  
  |  +-- Lens **        
  +--+               
     +-- Physunio **

Jeratthitikul et al., 2024 [Publ 4387], fig. 1

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	56	6	2	14

  +----------- Ctenodesmina
--+                        
  |  +-------- Rectidentina
  +--+                     
     |  +----- Physunio **    
     +--+                  
        |  +-- Lens **        
        +--+               
           +-- Trapezoideus **

Dai et al., 2023 [Publ 4285], fig. 3-BI

tT	bi	riS	oT	roT	roS	iD
3	3	1	2	42	2	1	12

  +----- Rectidentina
--+                  
  |  +-- Physunio **    
  +--+               
     +-- Lens **

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	73	3	1	12

  +-------- Rectidentina
--+                     
  |  +----- Lens **        
  +--+                  
     |  +-- Trapezoideus **
     +--+               
        +-- Physunio **

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	94	2	1	--

  +----- Rectidentina
--+                  
  |  +-- Lens **        
  +--+               
     +-- Physunio **

Dai et al., 2023 [Publ 4285], fig. 5

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	51	2	1	--

  +----- Rectidentina
--+                  
  |  +-- Lens **        
  +--+               
     +-- Physunio **

Matos et al., 2024 [Publ 4383], fig. 3

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	18	2	1	--

  +----- Rectidentina
--+                  
  |  +-- Physunio **    
  +--+               
     +-- Lens **

Klymus et al., 2025 [Publ 4814], fig. 3B

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	34	2	1	--

  +----- Rectidentina
--+                  
  |  +-- Lens **        
  +--+               
     +-- Physunio **

Zieritz et al., 2021 [Publ 4231], fig. 3-M_4

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	21	2	1	--

  +----- Rectidentina
--+                  
  |  +-- Lens **        
  +--+               
     +-- Physunio **

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	94	2	1	--

  +----- Rectidentina
--+                  
  |  +-- Lens **        
  +--+               
     +-- Physunio **

Wu et al., 2026 [Publ 4811], fig. 4

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	94	2	1	--

  +----- Rectidentina
--+                  
  |  +-- Lens **        
  +--+               
     +-- Physunio **

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	46	2	1	--

  +----- Rectidentina
--+                  
  |  +-- Lens **        
  +--+               
     +-- Physunio **

Wu et al., 2024 [Publ 4350], fig. 8a_S1

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	90	2	1	--

  +----- Rectidentina
--+                  
  |  +-- Lens **        
  +--+               
     +-- Physunio **

Zhang et al., 2024 [Publ 4397], fig. 5

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	25	2	1	--

  +----- Rectidentina
--+                  
  |  +-- Lens **        
  +--+               
     +-- Physunio **

Wu et al., 2024 [Publ 4350], fig. 8b_S2

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	90	2	1	--

  +----- Rectidentina
--+                  
  |  +-- Lens **        
  +--+               
     +-- Physunio **

Wu et al., 2024 [Publ 4388], fig. 6

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	51	2	1	--

  +----- Rectidentina
--+                  
  |  +-- Lens **        
  +--+               
     +-- Physunio **

Dai et al., 2025 [Publ 4440], fig. 3

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	86	2	1	--

  +----- Rectidentina
--+                  
  |  +-- Lens **        
  +--+               
     +-- Physunio **

Campbell & Lydeard, 2026 [Publ 4810], fig. 4

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	45	2	1	--

  +----- Rectidentina
--+                  
  |  +-- Lens **        
  +--+               
     +-- Physunio **

Chen et al., 2025 [Publ 4448], fig. 3

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	84	2	1	--

  +----- Rectidentina
--+                  
  |  +-- Lens **        
  +--+               
     +-- Physunio **

Matos et al., 2024 [Publ 4384], fig. 3

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	18	2	1	--

  +----- Rectidentina
--+                  
  |  +-- Physunio **    
  +--+               
     +-- Lens **

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	139	2	1	--

  +----- Rectidentina
--+                  
  |  +-- Lens **        
  +--+               
     +-- Physunio **

Klymus et al., 2025 [Publ 4814], fig. 3A

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	34	2	1	--

  +----- Rectidentina
--+                  
  |  +-- Lens **        
  +--+               
     +-- Physunio **

Hoeh et al., 2009 [Publ 2716], fig. 5

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	45	0	0	--

-- Lens_contradens **

Hoeh et al., 2009 [Publ 2716], fig. 4

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	45	0	0	--

-- Lens_contradens **

Hoeh et al., 2009 [Publ 2716], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	46	0	0	--

-- Lens_contradens **

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	30	0	0	--

-- Physunio **

Graf & Cummings, 2006 [Publ 1968], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	46	0	0	--

-- Lens_contradens **

Smith, 2021 [Publ 4331], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	19	0	0	--

-- Lens_contradens **

Whelan et al., 2011 [Publ 3380], fig. 1-ML

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	35	0	0	--

-- Lens_contradens **

Graf, 2002 [Publ 2651], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	17	0	0	--

-- Lens_contradens **

Whelan et al., 2011 [Publ 3380], fig. 1-MP

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	35	0	0	--

-- Lens_contradens **

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