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Page last updated
Thu 17 July 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!


Bivalvia | Unionoida | Unionoidea | Unionidae | Unioninae | Anodontini

subtribe CRISTARIINA Lopes-Lima, Bogan & Froufe in Lopes-Lima et al., 2017: cladomics
     10 genera

Cladomics Summary

number of species = 36 (39 total)
number of subtaxa (rank: genus) = 10
analyzed in 185 cladograms in 107 publications
maximum number of terminals in an analysis = 33 (mean = 4.65, median = 4)
maximum number of subtaxa (genus) = 8 (mean = 2.61, median = 2)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 132
maximum number of included terminals = 33, (mean = 6.11 median = 4)
maximum number of subtaxa (genus) = 8 (mean = 3.25, median = 3)
proportion of cladograms that support basic monophyly = 0.85

Rigorous Monophyly

number of cladograms that test rigorous monophyly = 81
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (genus) taxon and at least one rigorous outgroup (family).
maximum number of included terminals = 27 (mean = 5.21, median = 4)
maximum number of subtaxa (genus) = 7 (mean = 3.42, median = 3)
proportion of cladograms that support rigorous monophyly = 0.83


Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.


PCoA Ordination


NJ Tree


Wu et al., 2025 [Publ 4442], fig. 2

** Top Monophyly Cladogram **

iTtTbiriSoTroTroSiD
33333182000
  +----------------- Cristaria **                  
  |                                             
  |           +----- Beringiana **                 
--+  +--------+                                 
  |  |        |  +-- Acudonta **                   
  |  |        +--+                              
  |  |           +-- Sinanodonta **                
  +--+                                          
     |        +----- Pletholophus_guangzhouensis **
     |  +-----+                                 
     |  |     |  +-- Pletholophus_tenuis **        
     |  |     +--+                              
     +--+        +-- Pletholophus_reinianus **     
        |                                       
        |  +-------- Pletholophus_honglinhensis ** 
        +--+                                    
           |  +----- Anemina **                    
           +--+                                 
              |  +-- Amuranodonta **               
              +--+                              
                 +-- Buldowskia **                 

Whelan et al., 2023 [Publ 4287], fig. 2

** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
44223737229
           +-- Anemina **             
  +--------+                       
  |        +-- Sinanodonta **         
--+                                
  |  +-------- Anodonta_cygnea     
  |  |                             
  +--+     +-- Anodonta_nuttalliana
     |  +--+                       
     |  |  +-- Pseudanodonta       
     +--+                          
        |  +-- Anodonta_anatina    
        +--+                       
           +-- Alasmidontina       

Xiang et al., 2025 [Publ 4433], fig. 6

iTtTbiriSoTroTroSiD
272725810020
  +-------------- Pletholophus **
  |                           
  |        +----- Beringiana **  
--+  +-----+                  
  |  |     |  +-- Mimo **        
  |  |     +--+               
  +--+        +-- Sinanodonta ** 
     |                        
     |  +-------- Cristaria **   
     +--+                     
        |  +----- Anemina **     
        +--+                  
           |  +-- Amuranodonta **
           +--+               
              +-- Buldowskia **  

Dai et al., 2024 [Publ 4385], fig. 3-ML

iTtTbiriSoTroTroSiD
272725740020
        +----- Amuranodonta **
  +-----+                  
  |     |  +-- Anemina **     
  |     +--+               
--+        +-- Buldowskia **  
  |                        
  |  +-------- Cristaria **   
  +--+                     
     |  +----- Pletholophus **
     +--+                  
        |  +-- Beringiana **  
        +--+               
           +-- Sinanodonta ** 

Dai et al., 2024 [Publ 4385], fig. 3-BI

iTtTbiriSoTroTroSiD
272724740021
        +----- Amuranodonta **
  +-----+                  
  |     |  +-- Anemina **     
  |     +--+               
--+        +-- Buldowskia **  
  |                        
  |  +-------- Cristaria **   
  +--+                     
     |  +----- Pletholophus **
     +--+                  
        |  +-- Beringiana **  
        +--+               
           +-- Sinanodonta ** 

Bogan et al., 2023 [Publ 4295], fig. 3

iTtTbiriSoTroTroSiD
272724741121
  +-------------- Anodontina  
  |                           
--+  +----------- Cristaria **   
  |  |                        
  |  |     +----- Amuranodonta **
  +--+  +--+                  
     |  |  |  +-- Anemina **     
     |  |  +--+               
     +--+     +-- Buldowskia **  
        |                     
        |  +----- Pletholophus **
        +--+                  
           |  +-- Beringiana **  
           +--+               
              +-- Sinanodonta ** 

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iTtTbiriSoTroTroSiD
2424197979220
           +-- Pseudanodonta
           |                
  +--------+-- Anodonta     
  |        |                
  |        +-- Alasmidontina
  |                         
--+  +-------- Cristaria **    
  |  |                      
  |  |  +----- Anemina **      
  |  +--+                   
  +--+  |  +-- Amuranodonta ** 
     |  +--+                
     |     +-- Buldowskia **   
     |                      
     |  +----- Pletholophus ** 
     +--+                   
        |  +-- Beringiana **   
        +--+                
           +-- Sinanodonta **  

Sano et al., 2022 [Publ 4311], fig. 3

iTtTbiriSoTroTroSiD
181816500021
        +-- Beringiana **  
  +-----+               
  |     +-- Sinanodonta ** 
--+                     
  |  +----- Pletholophus **
  +--+                  
     |  +-- Cristaria **   
     +--+               
        +-- Buldowskia **  

Wu et al., 2025 [Publ 4442], fig. 6

iTtTbiriSoTroTroSiD
151513820024
  +----------- Cristaria **   
  |                        
  |     +----- Amuranodonta **
--+  +--+                  
  |  |  |  +-- Anemina **     
  |  |  +--+               
  |  |     +-- Buldowskia **  
  +--+                     
     |     +-- Beringiana **  
     |  +--+               
     |  |  +-- Pletholophus **
     +--+                  
        |  +-- Acudonta **    
        +--+               
           +-- Sinanodonta ** 

Konopleva et al., 2023 [Publ 4269], fig. S1_4

iTtTbiriSoTroTroSiD
1313116433124
  +----------- Anodontina  
  |                        
  |     +----- Anemina **     
--+  +--+                  
  |  |  |  +-- Amuranodonta **
  |  |  +--+               
  +--+     +-- Buldowskia **  
     |                     
     |  +----- Cristaria **   
     +--+                  
        |  +-- Beringiana **  
        +--+               
           +-- Sinanodonta ** 

Douda et al., 2024 [Publ 4364], fig. 1

iTtTbiriSoTroTroSiD
131312210022
  +-- Beringiana ** 
--+              
  +-- Sinanodonta **

Sano et al., 2022 [Publ 4311], fig. 2

iTtTbiriSoTroTroSiD
121212500033
        +-- Beringiana **  
  +-----+               
  |     +-- Sinanodonta ** 
--+                     
  |  +----- Pletholophus **
  +--+                  
     |  +-- Cristaria **   
     +--+               
        +-- Buldowskia **  

Lopes-Lima et al., 2020 [Publ 4080], fig. 10

iTtTbiriSoTroTroSiD
12128220025
  +-- Beringiana ** 
--+              
  +-- Sinanodonta **

Bogan et al., 2023 [Publ 4295], fig. 6

iTtTbiriSoTroTroSiD
12129310028
  +----- Pletholophus **
--+                  
  |  +-- Cristaria **   
  +--+               
     +-- Sinanodonta ** 

Sano et al., 2020 [Publ 4138], fig. 3

iTtTbiriSoTroTroSiD
1135376578253
                                +-- Anemina **      
  +-----------------------------+                
  |                             +-- Buldowskia **   
  |                                              
--+                          +----- Sinanodonta **  
  |  +-----------------------+                   
  |  |                       |  +-- Pletholophus ** 
  |  |                       +--+                
  +--+                          +-- Beringiana **   
     |                                           
     |  +-------------------------- Cristaria **    
     |  |                                        
     +--+                       +-- Anodontina   
        |  +--------------------+                
        |  |                    +-- Alasmidontina
        +--+                                     
           |  +-------------------- Lanceolaria  
           +--+                                  
              |  +----------------- Lepidodesma  
              +--+                               
                 |  +-------------- Aculamprotula
                 +--+                            
                    |  +----------- Unio         
                    +--+                         
                       |  +-------- Inversiunio  
                       +--+                      
                          |  +----- Nodularia    
                          +--+                   
                             |  +-- Schistodesmus
                             +--+                
                                +-- Tchangsinaia 

Sano et al., 2020 [Publ 4138], fig. 4

iTtTbiriSoTroTroSiD
1135376578253
                                +-- Anemina **      
  +-----------------------------+                
  |                             +-- Buldowskia **   
  |                                              
--+                          +----- Sinanodonta **  
  |  +-----------------------+                   
  |  |                       |  +-- Pletholophus ** 
  |  |                       +--+                
  +--+                          +-- Beringiana **   
     |                                           
     |  +-------------------------- Cristaria **    
     |  |                                        
     +--+                       +-- Anodontina   
        |  +--------------------+                
        |  |                    +-- Alasmidontina
        +--+                                     
           |  +-------------------- Lepidodesma  
           +--+                                  
              |  +----------------- Lanceolaria  
              +--+                               
                 |  +-------------- Aculamprotula
                 +--+                            
                    |  +----------- Unio         
                    +--+                         
                       |  +-------- Nodularia    
                       +--+                      
                          |  +----- Inversiunio  
                          +--+                   
                             |  +-- Schistodesmus
                             +--+                
                                +-- Tchangsinaia 

Sano et al., 2020 [Publ 4138], fig. 2

iTtTbiriSoTroTroSiD
1135366578251
                                +-- Anemina **      
  +-----------------------------+                
  |                             +-- Buldowskia **   
  |                                              
--+                          +----- Pletholophus ** 
  |  +-----------------------+                   
  |  |                       |  +-- Beringiana **   
  |  |                       +--+                
  +--+                          +-- Sinanodonta **  
     |                                           
     |  +-------------------------- Cristaria **    
     |  |                                        
     +--+                       +-- Anodontina   
        |  +--------------------+                
        |  |                    +-- Alasmidontina
        +--+                                     
           |  +-------------------- Lanceolaria  
           +--+                                  
              |  +----------------- Lepidodesma  
              +--+                               
                 |  +-------------- Aculamprotula
                 +--+                            
                    |  +----------- Unio         
                    +--+                         
                       |  +-------- Inversiunio  
                       +--+                      
                          |  +----- Nodularia    
                          +--+                   
                             |  +-- Tchangsinaia 
                             +--+                
                                +-- Schistodesmus

Sano et al., 2020 [Publ 4138], fig. 1

iTtTbiriSoTroTroSiD
102256240047
  +----- Lanceolaria_grayii__1  
  |                             
  +----- Lanceolaria_grayii__2  
  |                             
  +----- Cristaria_plicata__1 **   
  |                             
  +----- Cristaria_plicata__2 **   
  |                             
  +----- Sinanodonta_lauta **      
  |                             
  +----- Beringiana_japonica **    
  |                             
  +----- Beringiana_fukuharai__1 **
  |                             
  +----- Beringiana_beringiana **  
  |                             
  +----- Beringiana_fukuharai__2 **
  |                             
  +----- Beringiana_fukuharai__3 **
  |                             
  +----- Buldowskia_iwakawai__1 ** 
  |                             
  +----- Buldowskia_iwakawai__2 ** 
  |                             
  +----- Sinanodonta_tumens__1 **  
--+                             
  +----- Sinanodonta_tumens__2 **  
  |                             
  +----- Pletholophus **           
  |                             
  +----- Anemina **                
  |                             
  +----- Lanceolaria_lanceolata 
  |                             
  +----- Cristaria_plicata__3 **   
  |                             
  +----- Lanceolaria_gladiola   
  |                             
  +----- Sinanodonta_angula **     
  |                             
  +----- Lepidodesma            
  |                             
  |  +-- Aculamprotula          
  +--+                          
  |  +-- Acuticosta             
  |                             
  |  +-- Cuneopsis              
  +--+                          
  |  +-- Nodularia              
  |                             
  |  +-- Schistodesmus          
  +--+                          
     +-- Inversiunio            

Froufe et al., 2017 [Publ 3802], fig. 6

iTtTbiriSoTroTroSiD
10107422126
  +----------- Anodontina 
--+                       
  |  +-------- Anemina **    
  +--+                    
     |  +----- Cristaria **  
     +--+                 
        |  +-- Beringiana ** 
        +--+              
           +-- Sinanodonta **

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
94522419011263
  +-------- Acuticosta           
  |                              
  +-------- Alasmidonta          
  |                              
  +-------- Anodonta_anatina     
  |                              
  +-------- Anemina **              
  |                              
  +-------- Anodonta_cygnea      
  |                              
  +-------- Cristaria **            
  |                              
  +-------- Anodonta_nuttalliana 
  |                              
  +-------- Lasmigona            
  |                              
  +-------- Lepidodesma          
  |                              
  +-------- Pseudanodonta        
  |                              
  +-------- Pyganodon            
  |                              
--+-------- Simpsonaias          
  |                              
  +-------- Strophitus           
  |                              
  +-------- Schistodesmus        
  |                              
  +-------- Utterbackia          
  |                              
  +-------- Lanceolaria          
  |                              
  +-------- Unio                 
  |                              
  +-------- Aculamprotula        
  |                              
  |     +-- Pletholophus **         
  +-----+                        
  |     +-- Sinanodonta **          
  |                              
  |  +----- Cuneopsis_celtiformis
  |  |                           
  |  +----- Tchangsinaia         
  |  |                           
  +--+----- Cuneopsis_rufescens  
     |                           
     +----- Nodularia            
     |                           
     |  +-- Cuneopsis_szechenyii 
     +--+                        
        +-- Cuneopsis_heudei     

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
997419011228
     +-------- Anemina **      
  +--+                      
  |  |  +----- Cristaria **    
  |  +--+                   
  |     |  +-- Pletholophus ** 
  |     +--+                
  |        +-- Sinanodonta **  
--+                         
  |     +----- Alasmidonta  
  |     |                   
  |     +----- Lasmigona    
  |     |                   
  |     +----- Pseudanodonta
  |     |                   
  +-----+----- Simpsonaias  
        |                   
        +----- Strophitus   
        |                   
        +----- Anodonta     
        |                   
        |  +-- Pyganodon    
        +--+                
           +-- Utterbackia  

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
997418811228
        +-------- Anemina **      
  +-----+                      
  |     |  +----- Cristaria **    
  |     +--+                   
  |        |  +-- Pletholophus ** 
  |        +--+                
--+           +-- Sinanodonta **  
  |                            
  |  +----------- Anodonta     
  |  |                         
  |  |        +-- Alasmidonta  
  |  +--------+                
  +--+        +-- Lasmigona    
     |                         
     |        +-- Pyganodon    
     |  +-----+                
     |  |     +-- Utterbackia  
     +--+                      
        |  +----- Pseudanodonta
        +--+                   
           |  +-- Simpsonaias  
           +--+                
              +-- Strophitus   

Lopes-Lima et al., 2020 [Publ 4080], fig. 6

iTtTbiriSoTroTroSiD
997420030
  +-------- Sinanodonta ** 
--+                     
  |  +----- Amuranodonta **
  +--+                  
     |  +-- Anemina **     
     +--+               
        +-- Buldowskia **  

Sano et al., 2017 [Publ 3799], fig. 2

iTtTbiriSoTroTroSiD
9975240032
        +-- Cristaria **   
  +-----+               
  |     +-- Buldowskia **  
--+                     
  |  +----- Beringiana **  
  +--+                  
     |  +-- Pletholophus **
     +--+               
        +-- Sinanodonta ** 

Bolotov et al., 2020 [Publ 4084], fig. 1

iTtTbiriSoTroTroSiD
9976273128
  +----------- Anodontina  
  |                        
  |     +----- Amuranodonta **
--+  +--+                  
  |  |  |  +-- Anemina **     
  |  |  +--+               
  +--+     +-- Buldowskia **  
     |                     
     |  +----- Cristaria **   
     +--+                  
        |  +-- Beringiana **  
        +--+               
           +-- Sinanodonta ** 

Bespalaya et al., 2018 [Publ 3811], fig. 3

iTtTbiriSoTroTroSiD
887220032
  +-- Anemina **    
--+              
  +-- Sinanodonta **

Kondakov et al., 2018 [Publ 3914], fig. 5

iTtTbiriSoTroTroSiD
886200025
  +-- Anemina **    
--+              
  +-- Sinanodonta **

Huang et al., 2019 [Publ 3843], fig. 1

iTtTbiriSoTroTroSiD
829273350050
  +----------------------------------- Anemina **              
  |                                                         
--+  +-------------------------------- Acuticosta           
  |  |                                                      
  +--+  +----------------------------- Sinanodonta **          
     |  |                                                   
     +--+  +-------------------------- Aculamprotula        
        |  |                                                
        +--+                       +-- Cristaria **            
           |  +--------------------+                        
           |  |                    +-- Lanceolaria          
           +--+                                             
              |  +-------------------- Lepidodesma          
              +--+                                          
                 |  +----------------- Schistodesmus        
                 +--+                                       
                    |  +-------------- Nodularia            
                    +--+                                    
                       |  +----------- Cuneopsis_rufescens  
                       +--+                                 
                          |  +-------- Tchangsinaia         
                          +--+                              
                             |  +----- Cuneopsis_celtiformis
                             +--+                           
                                |  +-- Cuneopsis_szechenyii 
                                +--+                        
                                   +-- Cuneopsis_heudei     

Huang et al., 2019 [Publ 3843], fig. S1

iTtTbiriSoTroTroSiD
831363370064
  +-------------------------- Anemina **      
  |                                        
--+  +----------------------- Sinanodonta **  
  |  |                                     
  +--+  +-------------------- Acuticosta   
     |  |                                  
     +--+                 +-- Lepidodesma  
        |  +--------------+                
        |  |              +-- Cristaria **    
        +--+                               
           |  +-------------- Lanceolaria  
           +--+                            
              |  +----------- Aculamprotula
              +--+                         
                 |  +-------- Nodularia    
                 +--+                      
                    |  +----- Tchangsinaia 
                    +--+                   
                       |  +-- Schistodesmus
                       +--+                
                          +-- Cuneopsis    

Vikhrev et al., 2017 [Publ 3810], fig. 4

iTtTbiriSoTroTroSiD
886220025
  +-- Anemina **    
--+              
  +-- Sinanodonta **

Bauer et al., 2021 [Publ 4365], fig. 2

iTtTbiriSoTroTroSiD
885200026
  +-- Anemina **    
--+              
  +-- Sinanodonta **

Sano et al., 2017 [Publ 3799], fig. 1

iTtTbiriSoTroTroSiD
8865130029
        +-- Cristaria **   
  +-----+               
  |     +-- Buldowskia **  
--+                     
  |  +----- Pletholophus **
  +--+                  
     |  +-- Sinanodonta ** 
     +--+               
        +-- Beringiana **  

Xiang et al., 2025 [Publ 4433], fig. 7

iTtTbiriSoTroTroSiD
775562128
  +-------------- Anodontina 
--+                          
  |  +----------- Cristaria **  
  +--+                       
     |  +-------- Anemina **    
     +--+                    
        |  +----- Beringiana ** 
        +--+                 
           |  +-- Mimo **       
           +--+              
              +-- Sinanodonta **

Dai et al., 2025 [Publ 4440], fig. 2

iTtTbiriSoTroTroSiD
636306394261
                                   +-- Anemina **               
  +--------------------------------+                         
  |                                +-- Amuranodonta **          
  |                                                          
  |                             +----- Pletholophus **          
--+  +--------------------------+                            
  |  |                          |  +-- Sinanodonta **           
  |  |                          +--+                         
  |  |                             +-- Beringiana **            
  +--+                                                       
     |                             +-- Alasmidontina         
     |  +--------------------------+                         
     |  |                          +-- Anodontina            
     |  |                                                    
     +--+  +-------------------------- Lanceolaria_gladiola  
        |  |                                                 
        |  +-------------------------- Lanceolaria_oxyrhyncha
        |  |                                                 
        +--+-------------------------- Cristaria **             
           |                                                 
           |  +----------------------- Acuticosta            
           |  |                                              
           |  +----------------------- Lepidodesma           
           +--+                                              
              |  +-------------------- Globunio              
              |  |                                           
              +--+  +----------------- Aculamprotula         
                 |  |                                        
                 |  |              +-- Middendorffinaia      
                 +--+  +-----------+                         
                    |  |           +-- Nodularia             
                    |  |                                     
                    +--+  +----------- Cuneopsis             
                       |  |                                  
                       |  |        +-- Diaurora              
                       +--+--------+                         
                          |        +-- Schistodesmus         
                          |                                  
                          |  +-------- Pseudocuneopsis       
                          +--+                               
                             |  +----- Inversiunio           
                             +--+                            
                                |  +-- Tchangsinaia          
                                +--+                         
                                   +-- Unio                  

Bolotov et al., 2017 [Publ 3761], fig. S1

iTtTbiriSoTroTroSiD
66431339227
           +----- Cristaria **           
  +--------+                          
  |        |  +-- Pletholophus **        
  |        +--+                       
--+           +-- Sinanodonta **         
  |                                   
  |  +----------- Anodonta_anatina    
  +--+                                
     |  +-------- Alasmidontina       
     +--+                             
        |  +----- Pseudanodonta       
        +--+                          
           |  +-- Anodonta_cygnea     
           +--+                       
              +-- Anodonta_nuttalliana

Dai et al., 2025 [Publ 4440], fig. 1

iTtTbiriSoTroTroSiD
639326394264
                                         +-- Anemina **         
  +--------------------------------------+                   
  |                                      +-- Amuranodonta **    
  |                                                          
  |                                   +----- Parreysia       
  +-----------------------------------+                      
  |                                   |  +-- Coelatura       
  |                                   +--+                   
--+                                      +-- Indonaia        
  |                                                          
  |                                   +----- Pletholophus **    
  |  +--------------------------------+                      
  |  |                                |  +-- Sinanodonta **     
  |  |                                +--+                   
  |  |                                   +-- Beringiana **      
  +--+                                                       
     |  +----------------------------------- Cristaria **       
     |  |                                                    
     |  |                                +-- Alasmidontina   
     |  +--------------------------------+                   
     +--+                                +-- Anodontina      
        |                                                    
        |  +-------------------------------- Lanceolaria     
        |  |                                                 
        +--+  +----------------------------- Lepidodesma     
           |  |                                              
           +--+  +-------------------------- Acuticosta      
              |  |                                           
              +--+  +----------------------- Globunio        
                 |  |                                        
                 +--+  +-------------------- Aculamprotula   
                    |  |                                     
                    +--+  +----------------- Middendorffinaia
                       |  |                                  
                       +--+  +-------------- Nodularia       
                          |  |                               
                          +--+  +----------- Cuneopsis       
                             |  |                            
                             |  |        +-- Diaurora        
                             +--+--------+                   
                                |        +-- Schistodesmus   
                                |                            
                                |  +-------- Pseudocuneopsis 
                                +--+                         
                                   |  +----- Unio            
                                   +--+                      
                                      |  +-- Tchangsinaia    
                                      +--+                   
                                         +-- Inversiunio     

Wu et al., 2018 [Publ 3836], fig. 2

iTtTbiriSoTroTroSiD
6633477228
        +-- Anemina **         
        |                   
  +-----+-- Cristaria **       
  |     |                   
--+     +-- Sinanodonta **     
  |                         
  |  +----- Anodonta_cygnea 
  +--+                      
     |  +-- Anodonta_anatina
     +--+                   
        +-- Alasmidontina   

Chen et al., 2023 [Publ 4302], fig. 2

iTtTbiriSoTroTroSiD
6646344229
              +-- Alasmidontina
  +-----------+                
  |           +-- Anodontina   
--+                            
  |  +----------- Cristaria **    
  |  |                         
  +--+        +-- Anemina **      
     |  +-----+                
     |  |     +-- Amuranodonta ** 
     +--+                      
        |  +----- Pletholophus ** 
        +--+                   
           |  +-- Sinanodonta **  
           +--+                
              +-- Beringiana **   

Bogan et al., 2023 [Publ 4295], fig. 4

iTtTbiriSoTroTroSiD
664310029
  +----- Cristaria **   
--+                  
  |  +-- Sinanodonta ** 
  +--+               
     +-- Pletholophus **

Bolotov et al., 2017 [Publ 3761], fig. S2

iTtTbiriSoTroTroSiD
66431339227
  +----- Cristaria **   
--+                  
  |  +-- Pletholophus **
  +--+               
     +-- Sinanodonta ** 

Chen et al., 2023 [Publ 4302], fig. 1

iTtTbiriSoTroTroSiD
631236344256
                          +-- Anemina **               
  +-----------------------+                         
  |                       +-- Amuranodonta **          
  |                                                 
  |                    +----- Pletholophus **          
  |  +-----------------+                            
--+  |                 |  +-- Sinanodonta **           
  |  |                 +--+                         
  |  |                    +-- Beringiana **            
  |  |                                              
  |  |  +-------------------- Lanceolaria_gladiola  
  +--+  |                                           
     |  +-------------------- Acuticosta            
     |  |                                           
     |  +-------------------- Lepidodesma           
     |  |                                           
     |  +-------------------- Lanceolaria_oxyrhyncha
     |  |                                           
     +--+-------------------- Cristaria **             
        |                                           
        |                 +-- Alasmidontina         
        +-----------------+                         
        |                 +-- Anodontina            
        |                                           
        |  +----------------- Aculamprotula         
        |  |                                        
        |  |              +-- Middendorffinaia      
        +--+  +-----------+                         
           |  |           +-- Nodularia             
           |  |                                     
           +--+  +----------- Cuneopsis             
              |  |                                  
              |  |        +-- Diaurora              
              +--+--------+                         
                 |        +-- Schistodesmus         
                 |                                  
                 |  +-------- Pseudocuneopsis       
                 +--+                               
                    |  +----- Inversiunio           
                    +--+                            
                       |  +-- Tchangsinaia          
                       +--+                         
                          +-- Unio                  

Bolotov et al., 2016 [Publ 3732], fig. 2

iTtTbiriSoTroTroSiD
665120032
1

Wu et al., 2024 [Publ 4379], fig. 5

iTtTbiriSoTroTroSiD
66444712227
           +-------- Cristaria **           
  +--------+                             
  |        |  +----- Anemina **             
  |        +--+                          
  |           |  +-- Beringiana **          
--+           +--+                       
  |              +-- Sinanodonta **         
  |                                      
  |  +-------------- Anodonta_nuttalliana
  +--+                                   
     |  +----------- Alasmidontina       
     +--+                                
        |  +-------- Pseudanodonta       
        +--+                             
           |  +----- Anodonta_anatina    
           +--+                          
              |  +-- Anodonta_cygnea     
              +--+                       
                 +-- Anodonta_exulcerata 

Hou et al., 2024 [Publ 4400], fig. 4

iTtTbiriSoTroTroSiD
66447012227
        +-------- Cristaria **           
  +-----+                             
  |     |  +----- Anemina **             
  |     +--+                          
  |        |  +-- Beringiana **          
  |        +--+                       
--+           +-- Sinanodonta **         
  |                                   
  |           +-- Anodonta_nuttalliana
  |  +--------+                       
  |  |        +-- Alasmidontina       
  +--+                                
     |  +-------- Pseudanodonta       
     +--+                             
        |  +----- Anodonta_anatina    
        +--+                          
           |  +-- Anodonta_cygnea     
           +--+                       
              +-- Anodonta_exulcerata 

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iTtTbiriSoTroTroSiD
66448912229
        +-- Anodontina   
  +-----+                
  |     +-- Alasmidontina
  |                      
--+     +-- Anemina **      
  |  +--+                
  |  |  +-- Cristaria **    
  +--+                   
     |  +-- Beringiana **   
     +--+                
        +-- Sinanodonta **  

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iTtTbiriSoTroTroSiD
66448512227
        +-------- Cristaria **           
  +-----+                             
  |     |  +----- Anemina **             
  |     +--+                          
  |        |  +-- Beringiana **          
  |        +--+                       
--+           +-- Sinanodonta **         
  |                                   
  |           +-- Anodonta_nuttalliana
  |  +--------+                       
  |  |        +-- Alasmidontina       
  +--+                                
     |  +-------- Pseudanodonta       
     +--+                             
        |  +----- Anodonta_anatina    
        +--+                          
           |  +-- Anodonta_cygnea     
           +--+                       
              +-- Anodonta_exulcerata 

Bolotov et al., 2017 [Publ 3764], fig. S1

iTtTbiriSoTroTroSiD
617031199242
  +-- Alasmidonta          
  |                        
  +-- Anodonta_anatina     
  |                        
  +-- Anodonta_cygnea      
  |                        
  +-- Anodonta_nuttalliana 
  |                        
  +-- Lasmigona            
  |                        
  +-- Pseudanodonta        
  |                        
  +-- Pyganodon            
  |                        
  +-- Simpsonaias          
  |                        
  +-- Strophitus           
--+                        
  +-- Sinanodonta_lauta **    
  |                        
  +-- Cristaria_plicata **    
  |                        
  +-- Cristaria_radiata **    
  |                        
  +-- Pletholophus **         
  |                        
  +-- Sinanodonta_lucida **   
  |                        
  +-- Sinanodonta_jourdyi **  
  |                        
  +-- Lanceolaria_gladiola 
  |                        
  +-- Lanceolaria_grayii__1
  |                        
  +-- Lanceolaria_grayii__2

Lopes-Lima et al., 2020 [Publ 4080], fig. 7

iTtTbiriSoTroTroSiD
664220027
  +-- Sinanodonta **
--+              
  +-- Beringiana ** 

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iTtTbiriSoTroTroSiD
66448512227
           +-------- Cristaria **           
  +--------+                             
  |        |  +----- Anemina **             
  |        +--+                          
  |           |  +-- Beringiana **          
--+           +--+                       
  |              +-- Sinanodonta **         
  |                                      
  |  +-------------- Anodonta_nuttalliana
  +--+                                   
     |  +----------- Alasmidontina       
     +--+                                
        |  +-------- Pseudanodonta       
        +--+                             
           |  +----- Anodonta_anatina    
           +--+                          
              |  +-- Anodonta_cygnea     
              +--+                       
                 +-- Anodonta_exulcerata 

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iTtTbiriSoTroTroSiD
66448912227
     +-------- Cristaria **           
  +--+                             
  |  |  +----- Anemina **             
  |  +--+                          
  |     |  +-- Beringiana **          
  |     +--+                       
--+        +-- Sinanodonta **         
  |                                
  |  +-------- Anodonta_nuttalliana
  |  |                             
  |  +-------- Pseudanodonta       
  +--+                             
     |  +----- Anodonta_cygnea     
     |  |                          
     +--+----- Alasmidontina       
        |                          
        |  +-- Anodonta_exulcerata 
        +--+                       
           +-- Anodonta_anatina    

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

iTtTbiriSoTroTroSiD
5533719228
        +-- Anodontina   
  +-----+                
  |     +-- Alasmidontina
--+                      
  |  +----- Cristaria **    
  +--+                   
     |  +-- Pletholophus ** 
     +--+                
        +-- Sinanodonta **  

Lopes-Lima et al., 2020 [Publ 4080], fig. 8

iTtTbiriSoTroTroSiD
553220028
  +-- Sinanodonta **
--+              
  +-- Cristaria **  

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

iTtTbiriSoTroTroSiD
5533719228
        +-- Anodontina   
  +-----+                
  |     +-- Alasmidontina
--+                      
  |  +----- Cristaria **    
  +--+                   
     |  +-- Pletholophus ** 
     +--+                
        +-- Sinanodonta **  

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iTtTbiriSoTroTroSiD
5534759230
           +-- Anodontina   
  +--------+                
  |        +-- Alasmidontina
--+                         
  |  +-------- Cristaria **    
  +--+                      
     |  +----- Anemina **      
     +--+                   
        |  +-- Pletholophus ** 
        +--+                
           +-- Sinanodonta **  

Bogan et al., 2023 [Publ 4295], fig. 5

iTtTbiriSoTroTroSiD
553310028
  +----- Cristaria **   
--+                  
  |  +-- Sinanodonta ** 
  +--+               
     +-- Pletholophus **

Soroka, 2010 [Publ 3404], fig. 9a

iTtTbiriSoTroTroSiD
553300028
  +----- Buldowskia ** 
--+                 
  |  +-- Beringiana ** 
  +--+              
     +-- Sinanodonta **

Mabuchi & Nishida, 2020 [Publ 4120], fig. 1

iTtTbiriSoTroTroSiD
555370035
  +-------------- Cristaria **               
  |                                       
  |  +----------- Sinanodonta_tumens **      
  |  |                                    
  |  |        +-- Sinanodonta_lauta__5 **    
  |  |        |                           
--+  |        +-- Sinanodonta_lauta__6 **    
  |  |        |                           
  |  |        +-- Beringiana_japonica__15 ** 
  |  |        |                           
  |  |        +-- Beringiana_japonica__16 ** 
  |  |  +-----+                           
  |  |  |     +-- Beringiana_japonica__17 ** 
  +--+  |     |                           
     |  |     +-- Beringiana_japonica__18 ** 
     |  |     |                           
     |  |     +-- Beringiana_japonica__19 ** 
     |  |     |                           
     |  |     +-- Beringiana_japonica__20 ** 
     |  |                                 
     |  |  +----- Beringiana_japonica__1 **  
     |  |  |                              
     |  |  +----- Beringiana_japonica__2 **  
     |  |  |                              
     +--+  +----- Beringiana_japonica__3 **  
        |  |                              
        |  +----- Beringiana_japonica__4 **  
        |  |                              
        |  +----- Beringiana_japonica__5 **  
        |  |                              
        |  +----- Sinanodonta_lauta__1 **    
        |  |                              
        |  +----- Beringiana_japonica__6 **  
        |  |                              
        |  +----- Sinanodonta_lauta__2 **    
        |  |                              
        |  +----- Sinanodonta_calipygos__1 **
        |  |                              
        +--+----- Beringiana_japonica__7 **  
           |                              
           +----- Sinanodonta_lauta__4 **    
           |                              
           +----- Beringiana_japonica__8 **  
           |                              
           +----- Beringiana_japonica__9 **  
           |                              
           +----- Beringiana_japonica__10 ** 
           |                              
           +----- Beringiana_japonica__11 ** 
           |                              
           +----- Beringiana_japonica__12 ** 
           |                              
           +----- Beringiana_japonica__13 ** 
           |                              
           +----- Beringiana_japonica__14 ** 
           |                              
           |  +-- Sinanodonta_lauta__3 **    
           +--+                           
              +-- Sinanodonta_calipygos__2 **

Wang et al., 2018 [Publ 3846], fig. 2

iTtTbiriSoTroTroSiD
4423273229
        +-- Anodontina   
  +-----+                
  |     +-- Alasmidontina
--+                      
  |  +----- Cristaria **    
  +--+                   
     |  +-- Anemina **      
     +--+                
        +-- Sinanodonta **  

Wu et al., 2020 [Publ 4121], fig. 3

iTtTbiriSoTroTroSiD
4423455229
        +-- Anodontina   
  +-----+                
  |     +-- Alasmidontina
--+                      
  |  +----- Cristaria **    
  +--+                   
     |  +-- Anemina **      
     +--+                
        +-- Sinanodonta **  

Froufe et al., 2019 [Publ 3936], fig. 3

iTtTbiriSoTroTroSiD
4423405229
        +-- Anodontina   
  +-----+                
  |     +-- Alasmidontina
--+                      
  |  +----- Cristaria **    
  +--+                   
     |  +-- Anemina **      
     +--+                
        +-- Sinanodonta **  

Wu et al., 2021 [Publ 4366], fig. 6

iTtTbiriSoTroTroSiD
4423876231
           +----- Anemina **         
  +--------+                      
  |        |  +-- Cristaria **       
  |        +--+                   
--+           +-- Sinanodonta **     
  |                               
  |  +----------- Anodonta_cygnea 
  |  |                            
  +--+        +-- Lasmigona       
     |  +-----+                   
     |  |     +-- Anodonta_anatina
     +--+                         
        |  +----- Pseudanodonta   
        +--+                      
           |  +-- Pyganodon       
           +--+                   
              +-- Strophitus      

Cha et al., 2018 [Publ 3916], fig. 1

iTtTbiriSoTroTroSiD
4423345229
        +-- Anodontina   
  +-----+                
  |     +-- Alasmidontina
--+                      
  |  +----- Cristaria **    
  +--+                   
     |  +-- Anemina **      
     +--+                
        +-- Sinanodonta **  

Aunins et al., 2018 [Publ 3863], fig. 1

iTtTbiriSoTroTroSiD
4423367229
        +-- Anodontina   
  +-----+                
  |     +-- Alasmidontina
--+                      
  |  +----- Cristaria **    
  +--+                   
     |  +-- Anemina **      
     +--+                
        +-- Sinanodonta **  

Riccardi et al., 2020 [Publ 4089], fig. 2

iTtTbiriSoTroTroSiD
4423159229
           +----- Cristaria **           
  +--------+                          
  |        |  +-- Anemina **             
  |        +--+                       
--+           +-- Sinanodonta **         
  |                                   
  |  +----------- Anodonta_anatina    
  |  |                                
  +--+        +-- Anodonta_nuttalliana
     |  +-----+                       
     |  |     +-- Alasmidontina       
     +--+                             
        |  +----- Pseudanodonta       
        +--+                          
           |  +-- Anodonta_exulcerata 
           +--+                       
              +-- Anodonta_cygnea     

Klishko et al., 2016 [Publ 3733], fig. 1

iTtTbiriSoTroTroSiD
442300029
  +----- Cristaria **  
--+                 
  |  +-- Sinanodonta **
  +--+              
     +-- Beringiana ** 

Huang et al., 2018 [Publ 3828], fig. 6

iTtTbiriSoTroTroSiD
4423385229
        +-- Anodontina   
  +-----+                
  |     +-- Alasmidontina
--+                      
  |  +----- Cristaria **    
  +--+                   
     |  +-- Anemina **      
     +--+                
        +-- Sinanodonta **  

Wu et al., 2019 [Publ 3864], fig. 3

iTtTbiriSoTroTroSiD
4423425229
        +-- Anodontina   
  +-----+                
  |     +-- Alasmidontina
--+                      
  |  +----- Cristaria **    
  +--+                   
     |  +-- Anemina **      
     +--+                
        +-- Sinanodonta **  

Wu et al., 2022 [Publ 4238], fig. 3

iTtTbiriSoTroTroSiD
4423696229
        +-- Anodontina   
  +-----+                
  |     +-- Alasmidontina
--+                      
  |  +----- Cristaria **    
  +--+                   
     |  +-- Anemina **      
     +--+                
        +-- Sinanodonta **  

Sayenko et al., 2017 [Publ 3760], fig. 3

iTtTbiriSoTroTroSiD
442200029
  +-- Beringiana ** 
--+              
  +-- Sinanodonta **

Huang et al., 2019 [Publ 3843], fig. 3

iTtTbiriSoTroTroSiD
4423526229
     +----- Cristaria **       
  +--+                      
  |  |  +-- Anemina **         
  |  +--+                   
--+     +-- Sinanodonta **     
  |                         
  |  +----- Anodonta_cygnea 
  +--+                      
     |  +-- Anodonta_anatina
     +--+                   
        +-- Alasmidontina   

Wu et al., 2024 [Publ 4388], fig. 6

iTtTbiriSoTroTroSiD
4424503229
           +-- Anodontina   
  +--------+                
  |        +-- Alasmidontina
--+                         
  |  +-------- Cristaria **    
  +--+                      
     |  +----- Anemina **      
     +--+                   
        |  +-- Beringiana **   
        +--+                
           +-- Sinanodonta **  

Wu et al., 2021 [Publ 4366], fig. 4

iTtTbiriSoTroTroSiD
4423648229
     +----- Cristaria **       
  +--+                      
  |  |  +-- Anemina **         
  |  +--+                   
--+     +-- Sinanodonta **     
  |                         
  |  +----- Anodonta_cygnea 
  +--+                      
     |  +-- Anodonta_anatina
     +--+                   
        +-- Alasmidontina   

Wu et al., 2021 [Publ 4366], fig. 3-ML

iTtTbiriSoTroTroSiD
4423699229
     +----- Cristaria **       
  +--+                      
  |  |  +-- Anemina **         
  |  +--+                   
--+     +-- Sinanodonta **     
  |                         
  |  +----- Anodonta_cygnea 
  +--+                      
     |  +-- Anodonta_anatina
     +--+                   
        +-- Alasmidontina   

Soroka, 2010 [Publ 3404], fig. 9c

iTtTbiriSoTroTroSiD
443200032
  +-- Buldowskia ** 
--+              
  +-- Sinanodonta **

Lopes-Lima et al., 2020 [Publ 4080], fig. 9

iTtTbiriSoTroTroSiD
442320029
  +----- Cristaria **   
--+                  
  |  +-- Sinanodonta ** 
  +--+               
     +-- Pletholophus **

Bolotov et al., 2018 [Publ 3826], fig. 2

iTtTbiriSoTroTroSiD
4423796229
        +-- Anodontina   
  +-----+                
  |     +-- Alasmidontina
--+                      
  |  +----- Cristaria **    
  +--+                   
     |  +-- Pletholophus ** 
     +--+                
        +-- Sinanodonta **  

Wu et al., 2023 [Publ 4339], fig. 5

iTtTbiriSoTroTroSiD
4423155231
        +-- Alasmidontina
  +-----+                
  |     +-- Anodontina   
--+                      
  |  +----- Anemina **      
  +--+                   
     |  +-- Cristaria **    
     +--+                
        +-- Sinanodonta **  

Wu et al., 2021 [Publ 4366], fig. 3-BI

iTtTbiriSoTroTroSiD
4423699229
     +----- Cristaria **       
  +--+                      
  |  |  +-- Anemina **         
  |  +--+                   
--+     +-- Sinanodonta **     
  |                         
  |  +----- Anodonta_cygnea 
  +--+                      
     |  +-- Anodonta_anatina
     +--+                   
        +-- Alasmidontina   

Wu et al., 2022 [Publ 4238], fig. 2d

iTtTbiriSoTroTroSiD
4423346229
        +-- Anodontina   
  +-----+                
  |     +-- Alasmidontina
--+                      
  |  +----- Cristaria **    
  +--+                   
     |  +-- Anemina **      
     +--+                
        +-- Sinanodonta **  

Lopes-Lima et al., 2017 [Publ 3792], fig. 2

iTtTbiriSoTroTroSiD
4423285229
        +-- Anodontina   
  +-----+                
  |     +-- Alasmidontina
--+                      
  |  +----- Cristaria **    
  +--+                   
     |  +-- Anemina **      
     +--+                
        +-- Sinanodonta **  

Wu et al., 2021 [Publ 4366], fig. 2_I

iTtTbiriSoTroTroSiD
4423876231
           +----- Anemina **      
  +--------+                   
  |        |  +-- Cristaria **    
  |        +--+                
--+           +-- Sinanodonta **  
  |                            
  |  +----------- Lasmigona    
  +--+                         
     |  +-------- Anodonta     
     +--+                      
        |  +----- Pseudanodonta
        +--+                   
           |  +-- Pyganodon    
           +--+                
              +-- Strophitus   

Dai et al., 2025 [Publ 4440], fig. 3

iTtTbiriSoTroTroSiD
4423846229
        +-- Anodontina   
  +-----+                
  |     +-- Alasmidontina
--+                      
  |  +----- Cristaria **    
  +--+                   
     |  +-- Anemina **      
     +--+                
        +-- Sinanodonta **  

Soroka & Burzynski, 2017 [Publ 3837], fig. 1

iTtTbiriSoTroTroSiD
4423196229
        +-- Anodontina   
  +-----+                
  |     +-- Alasmidontina
--+                      
  |  +----- Cristaria **    
  +--+                   
     |  +-- Anemina **      
     +--+                
        +-- Sinanodonta **  

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

iTtTbiriSoTroTroSiD
4423445229
        +-- Anodontina   
  +-----+                
  |     +-- Alasmidontina
--+                      
  |  +----- Cristaria **    
  +--+                   
     |  +-- Anemina **      
     +--+                
        +-- Sinanodonta **  

Klishko et al., 2014 [Publ 3666], fig. 8

iTtTbiriSoTroTroSiD
442300029
  +----- Cristaria **  
--+                 
  |  +-- Beringiana ** 
  +--+              
     +-- Sinanodonta **

Bulakhova et al., 2023 [Publ 4325], fig. 5

iTtTbiriSoTroTroSiD
442100031
1

Wang et al., 2018 [Publ 3865], fig. 2

iTtTbiriSoTroTroSiD
4423273229
        +-- Anodontina   
  +-----+                
  |     +-- Alasmidontina
--+                      
  |  +----- Cristaria **    
  +--+                   
     |  +-- Anemina **      
     +--+                
        +-- Sinanodonta **  

Fonseca et al., 2016 [Publ 3775], fig. 1

iTtTbiriSoTroTroSiD
4423275229
        +-- Anodontina   
  +-----+                
  |     +-- Alasmidontina
--+                      
  |  +----- Cristaria **    
  +--+                   
     |  +-- Anemina **      
     +--+                
        +-- Sinanodonta **  

Froufe et al., 2019 [Publ 3936], fig. 6

iTtTbiriSoTroTroSiD
4423405229
        +-- Anodontina   
  +-----+                
  |     +-- Alasmidontina
--+                      
  |  +----- Cristaria **    
  +--+                   
     |  +-- Anemina **      
     +--+                
        +-- Sinanodonta **  

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

iTtTbiriSoTroTroSiD
4423342229
        +-- Anodontina   
  +-----+                
  |     +-- Alasmidontina
--+                      
  |  +----- Cristaria **    
  +--+                   
     |  +-- Anemina **      
     +--+                
        +-- Sinanodonta **  

Fukata & Iigo, 2020 [Publ 4106], fig. 1

iTtTbiriSoTroTroSiD
4423496229
        +-- Anodontina   
  +-----+                
  |     +-- Alasmidontina
--+                      
  |  +----- Cristaria **    
  +--+                   
     |  +-- Anemina **      
     +--+                
        +-- Sinanodonta **  

Soroka, 2010 [Publ 3404], fig. 9b

iTtTbiriSoTroTroSiD
442200029
  +-- Buldowskia ** 
--+              
  +-- Sinanodonta **

Xue et al., 2016 [Publ 3659], fig. 1

iTtTbiriSoTroTroSiD
4533111130
     +-- Cristaria **    
  +--+                
  |  +-- Alasmidontina
--+                   
  |  +-- Anemina **      
  +--+                
     +-- Sinanodonta **  

Dai et al., 2024 [Publ 4389], fig. 3

iTtTbiriSoTroTroSiD
3313440032
  +----- Sinanodonta **
--+                 
  |  +-- Anemina **    
  +--+              
     +-- Cristaria **  

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

iTtTbiriSoTroTroSiD
3313332230
        +-- Anodontina   
  +-----+                
  |     +-- Alasmidontina
--+                      
  |  +----- Cristaria **    
  +--+                   
     |  +-- Sinanodonta **  
     +--+                
        +-- Anemina **      

Kano et al., 2019 [Publ 4194], fig. 6

iTtTbiriSoTroTroSiD
331370032
  +----- Sinanodonta ** 
--+                  
  |  +-- Pletholophus **
  +--+               
     +-- Beringiana **  

Dai et al., 2023 [Publ 4285], fig. 5

iTtTbiriSoTroTroSiD
3313504230
        +-- Anodontina   
  +-----+                
  |     +-- Alasmidontina
--+                      
  |  +----- Cristaria **    
  +--+                   
     |  +-- Anemina **      
     +--+                
        +-- Sinanodonta **  

Chen et al., 2020 [Publ 4170], fig. 1

iTtTbiriSoTroTroSiD
334323318263
                       +----- Prolasmidonta           
                    +--+                              
                    |  |  +-- Alasmidonta             
                    |  +--+                           
  +-----------------+     +-- Anemina **                 
  |                 |                                 
  |                 |  +----- Pseudodontina           
  |                 +--+                              
  |                    |  +-- Lamprotula_caveata      
  |                    +--+                           
  |                       +-- Chamberlainia           
  |                                                   
  |                    +----- Schistodesmus           
  |              +-----+                              
  |              |     |  +-- Lepidodesma             
--+              |     +--+                           
  |              |        +-- Aculamprotula           
  |  +-----------+                                    
  |  |           |     +----- Platynaias              
  |  |           |  +--+                              
  |  |           |  |  |  +-- Utterbackia_imbecillis  
  |  |           |  |  +--+                           
  |  |           +--+     +-- Pyganodon               
  |  |              |                                 
  |  |              |  +----- Sinanodonta **             
  |  |              +--+                              
  |  |                 |  +-- Cristaria **               
  +--+                 +--+                           
     |                    +-- Anodontina              
     |                                                
     |                    +-- Utterbackia_peninsularis
     |           +--------+                           
     |           |        +-- Potamilus_alatus        
     |  +--------+                                    
     |  |        |  +-------- Lamprotula_leaii        
     |  |        +--+                                 
     |  |           |  +----- Sinohyriopsis           
     |  |           +--+                              
     +--+              |  +-- Parvasolenaia           
        |              +--+                           
        |                 +-- Sinosolenaia            
        |                                             
        |  +----------------- Gibbosula               
        +--+                                          
           |  +-------------- Quadrulina              
           +--+                                       
              |  +----------- Toxolasma               
              +--+                                    
                 |  +-------- Lampsilis_ornata        
                 +--+                                 
                    |  +----- Potamilus_leptodon      
                    +--+                              
                       |  +-- Lampsilis_siliquoidea   
                       +--+                           
                          +-- Lampsilis_powellii      

Ouyang et al., 2011 [Publ 3408], fig. 1

iTtTbiriSoTroTroSiD
3312100030
  +-- Anemina **    
--+              
  +-- Sinanodonta **

Wu et al., 2020 [Publ 4147], fig. 3

iTtTbiriSoTroTroSiD
3313264230
        +-- Anodontina   
  +-----+                
  |     +-- Alasmidontina
--+                      
  |  +----- Cristaria **    
  +--+                   
     |  +-- Sinanodonta **  
     +--+                
        +-- Anemina **      

Feng et al., 2016 [Publ 3800], fig. 2

iTtTbiriSoTroTroSiD
3312113230
  +----- Alasmidontina
--+                   
  |  +-- Anemina **      
  +--+                
     +-- Sinanodonta **  

Wu et al., 2016 [Publ 3711], fig. 1

iTtTbiriSoTroTroSiD
3753184236
  +----------- Sinanodonta **  
--+                         
  |  +-------- Anemina **      
  +--+                      
     |  +----- Cristaria **    
     +--+                   
        |  +-- Anodontina   
        +--+                
           +-- Alasmidontina

Gomes-dos-Santos et al., 2019 [Publ 4042], fig. 8

iTtTbiriSoTroTroSiD
331333130
  +-------- Anodontina 
--+                    
  |  +----- Cristaria **  
  +--+                 
     |  +-- Anemina **    
     +--+              
        +-- Sinanodonta **

Dai et al., 2025 [Publ 4441], fig. 1

iTtTbiriSoTroTroSiD
3313450032
  +----- Sinanodonta **
--+                 
  |  +-- Anemina **    
  +--+              
     +-- Cristaria **  

Wu et al., 2018 [Publ 4143], fig. 2

iTtTbiriSoTroTroSiD
3753194236
  +----------- Sinanodonta **  
--+                         
  |  +-------- Anemina **      
  +--+                      
     |  +----- Cristaria **    
     +--+                   
        |  +-- Anodontina   
        +--+                
           +-- Alasmidontina

Wu et al., 2024 [Publ 4379], fig. 4_6

iTtTbiriSoTroTroSiD
3312190030
  +-- Cristaria **  
--+              
  +-- Sinanodonta **

Dai et al., 2024 [Publ 4358], fig. 4

iTtTbiriSoTroTroSiD
3313494230
        +-- Anodontina   
  +-----+                
  |     +-- Alasmidontina
--+                      
  |  +----- Cristaria **    
  +--+                   
     |  +-- Anemina **      
     +--+                
        +-- Sinanodonta **  

Chong et al., 2008 [Publ 2668], fig. 3

iTtTbiriSoTroTroSiD
220216152--
  +-- Beringiana ** 
--+              
  +-- Sinanodonta **

Guerra et al., 2017 [Publ 3772], fig. 2b

iTtTbiriSoTroTroSiD
22022362--
     +-- Cristaria **    
  +--+                
  |  +-- Sinanodonta **  
--+                   
  |  +-- Anodontina   
  +--+                
     +-- Alasmidontina

Riccardi et al., 2020 [Publ 4089], fig. 1

iTtTbiriSoTroTroSiD
2202441--
  +----- Anodontina 
--+                 
  |  +-- Sinanodonta **
  +--+              
     +-- Anemina **    

Zhou et al., 2016 [Publ 3710], fig. 1

iTtTbiriSoTroTroSiD
2602184135
  +----- Sinanodonta **  
--+                   
  |  +-- Cristaria **    
  +--+                
     +-- Alasmidontina

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iTtTbiriSoTroTroSiD
22025522--
     +-- Cristaria **    
  +--+                
  |  +-- Sinanodonta **  
--+                   
  |  +-- Anodontina   
  +--+                
     +-- Alasmidontina

Soroka, 2010 [Publ 3404], fig. 3b-M

iTtTbiriSoTroTroSiD
2201411--
1

Soroka, 2010 [Publ 3404], fig. 3b

iTtTbiriSoTroTroSiD
2201731--
1

Xia et al., 2016 [Publ 3712], fig. 7B

iTtTbiriSoTroTroSiD
230210032
  +----- Sinanodonta **  
--+                   
  |  +-- Cristaria **    
  +--+                
     +-- Sinohyriopsis

Pfarrer et al., 2022 [Publ 4305], fig. 2-5

iTtTbiriSoTroTroSiD
22021031--
  +----- Anodontina 
--+                 
  |  +-- Cristaria **  
  +--+              
     +-- Sinanodonta **

Zhou et al., 2007 [Publ 2658], fig. 2

iTtTbiriSoTroTroSiD
21802160047
  +----------------- Sinanodonta **  
  |                               
--+  +-------------- Cristaria **    
  |  |                            
  +--+           +-- Lamprotula   
     |  +--------+                
     |  |        +-- Quadrulina   
     +--+                         
        |  +-------- Aculamprotula
        +--+                      
           |  +----- Acuticosta   
           +--+                   
              |  +-- Nodularia    
              +--+                
                 +-- Cuneopsis    

Zhou et al., 2007 [Publ 2658], fig. 1

iTtTbiriSoTroTroSiD
22021900--
  +-- Sinanodonta **
--+              
  +-- Cristaria **  

Soroka, 2010 [Publ 3404], fig. 3a-M

iTtTbiriSoTroTroSiD
2201411--
1

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iTtTbiriSoTroTroSiD
22025522--
     +-- Cristaria **    
  +--+                
  |  +-- Sinanodonta **  
--+                   
  |  +-- Anodontina   
  +--+                
     +-- Alasmidontina

Soroka, 2010 [Publ 3404], fig. 5b

iTtTbiriSoTroTroSiD
2201631--
1

Zhang et al., 2004 [Publ 2735], fig. 1

iTtTbiriSoTroTroSiD
22021200--
  +-- Sinanodonta **
--+              
  +-- Cristaria **  

Tomilova et al., 2020 [Publ 4097], fig. 3

iTtTbiriSoTroTroSiD
2201641--
1

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

iTtTbiriSoTroTroSiD
22022931--
  +----- Alasmidontina
--+                   
  |  +-- Cristaria **    
  +--+                
     +-- Sinanodonta **  

Wen et al., 2017 [Publ 3795], fig. 3

iTtTbiriSoTroTroSiD
22021411--
  +----- Anodontina 
--+                 
  |  +-- Sinanodonta **
  +--+              
     +-- Cristaria **  

Huang et al., 2002 [Publ 2657], fig. 2

iTtTbiriSoTroTroSiD
22023251--
  +-- Sinanodonta **
--+              
  +-- Cristaria **  

Huang et al., 2002 [Publ 2657], fig. 1

iTtTbiriSoTroTroSiD
22023251--
  +----- Alasmidontina
--+                   
  |  +-- Sinanodonta **  
  +--+                
     +-- Cristaria **    

Soroka, 2010 [Publ 3404], fig. 3a

iTtTbiriSoTroTroSiD
2201731--
1

Soroka, 2010 [Publ 3404], fig. 5a

iTtTbiriSoTroTroSiD
2201631--
  +-------- Anodonta_anatina
--+                         
  |  +----- Sinanodonta **     
  +--+                      
     |  +-- Anodonta_cygnea 
     +--+                   
        +-- Pseudanodonta   

Huang et al., 2013 [Publ 3629], fig. 3

iTtTbiriSoTroTroSiD
22021441--
  +----- Alasmidontina
--+                   
  |  +-- Sinanodonta **  
  +--+                
     +-- Cristaria **    

Zhang et al., 2013 [Publ 4214], fig. 5

iTtTbiriSoTroTroSiD
240220033
  +-------- Cristaria **    
--+                      
  |  +----- Sinanodonta **  
  +--+                   
     |  +-- Margaritifera
     +--+                
        +-- Lamprotula   

Guerra et al., 2017 [Publ 3772], fig. 2a

iTtTbiriSoTroTroSiD
2802236237
  +-------- Sinanodonta **  
--+                      
  |  +----- Cristaria **    
  +--+                   
     |  +-- Anodontina   
     +--+                
        +-- Alasmidontina

Huang et al., 2019 [Publ 3843], fig. 3-M

iTtTbiriSoTroTroSiD
11012232--
  +----- Sinanodonta **  
--+                   
  |  +-- Anodontina   
  +--+                
     +-- Alasmidontina

Zieritz et al., 2021 [Publ 4231], fig. 3-M_4

iTtTbiriSoTroTroSiD
11012232--
  +----- Sinanodonta **  
--+                   
  |  +-- Anodontina   
  +--+                
     +-- Alasmidontina

Soroka, 2010 [Publ 3404], fig. 5b-M

iTtTbiriSoTroTroSiD
1101111--
  +-- Anodontina 
--+              
  +-- Sinanodonta **

Plazzi & Passamonti, 2010 [Publ 3354], fig. 2-3

iTtTbiriSoTroTroSiD
1101300--
-- Sinanodonta_woodiana **

Burzynski & Soroka, 2018 [Publ 3862], fig. 5a

iTtTbiriSoTroTroSiD
11011932--
  +-- Sinanodonta **  
  |                
--+-- Anodontina   
  |                
  +-- Alasmidontina

Sheng et al., 2014 [Publ 3704], fig. 2

iTtTbiriSoTroTroSiD
1101711--
-- Cristaria_plicata **

Soroka, 2008 [Publ 3695], fig. 1

iTtTbiriSoTroTroSiD
1101322--
  +----- Sinanodonta **  
--+                   
  |  +-- Anodontina   
  +--+                
     +-- Alasmidontina

Soroka, 2010 [Publ 3404], fig. 5a-M

iTtTbiriSoTroTroSiD
1101111--
  +-- Anodontina 
--+              
  +-- Sinanodonta **

Soroka, 2010 [Publ 3404], fig. 2

iTtTbiriSoTroTroSiD
1101631--
-- Sinanodonta_woodiana **

Burzynski & Soroka, 2018 [Publ 3862], fig. 3

iTtTbiriSoTroTroSiD
1101332--
  +----- Sinanodonta **  
--+                   
  |  +-- Anodontina   
  +--+                
     +-- Alasmidontina

Smith, 2021 [Publ 4331], fig. 1

iTtTbiriSoTroTroSiD
11011921--
  +-- Cristaria **    
--+                
  +-- Alasmidontina

Froufe et al., 2019 [Publ 3936], fig. 4

iTtTbiriSoTroTroSiD
11011832--
  +----- Sinanodonta **  
--+                   
  |  +-- Anodontina   
  +--+                
     +-- Alasmidontina

Froufe et al., 2017 [Publ 3802], fig. 2

iTtTbiriSoTroTroSiD
1101331--
  +-- Cristaria ** 
--+             
  +-- Anodontina

Gustafson & Iwamoto, 2005 [Publ 2741], fig. 2

iTtTbiriSoTroTroSiD
1101421--
  +-- Beringiana **
--+             
  +-- Anodontina

Guerra et al., 2019 [Publ 4168], fig. 2-M

iTtTbiriSoTroTroSiD
11012132--
  +----- Alasmidontina
--+                   
  |  +-- Anodontina   
  +--+                
     +-- Sinanodonta **  

Mock et al., 2004 [Publ 2686], fig. 3

iTtTbiriSoTroTroSiD
1101111--
  +-- Anodontina 
--+              
  +-- Sinanodonta **

Dai et al., 2024 [Publ 4386], fig. 6

iTtTbiriSoTroTroSiD
11013921--
  +-- Sinanodonta **  
--+                
  +-- Alasmidontina

Chase et al., 2018 [Publ 3847], fig. 4

iTtTbiriSoTroTroSiD
110137132--
  +----- Sinanodonta **  
--+                   
  |  +-- Anodontina   
  +--+                
     +-- Alasmidontina

Peng et al., 2012 [Publ 3689], fig. 1

iTtTbiriSoTroTroSiD
1101200--
-- Cristaria_plicata **

Soroka, 2010 [Publ 3404], fig. 8b-M

iTtTbiriSoTroTroSiD
1101200--
-- Sinanodonta_woodiana **

Soroka, 2010 [Publ 3404], fig. 8b

iTtTbiriSoTroTroSiD
1101521--
-- Sinanodonta_woodiana **

Soroka, 2010 [Publ 3404], fig. 8a

iTtTbiriSoTroTroSiD
1101521--
  +-- Sinanodonta **
--+              
  +-- Anodontina 

Soroka, 2010 [Publ 3404], fig. 4

iTtTbiriSoTroTroSiD
1101631--
  +-- Sinanodonta **
--+              
  +-- Anodontina 

Soroka, 2008 [Publ 3695], fig. 1-M

iTtTbiriSoTroTroSiD
1101322--
  +----- Sinanodonta **  
--+                   
  |  +-- Anodontina   
  +--+                
     +-- Alasmidontina

Liu et al., 2017 [Publ 3939], fig. 2

iTtTbiriSoTroTroSiD
11011400--
-- Anemina_arcaeformis **

Mock et al., 2004 [Publ 2686], fig. 4

iTtTbiriSoTroTroSiD
1101111--
  +-- Anodontina 
--+              
  +-- Sinanodonta **

Nagel et al., 1998 [Publ 1946], fig. 7

iTtTbiriSoTroTroSiD
1101441--
  +-------------- Pseudanodonta   
--+                               
  |  +----------- Anodonta_cygnea 
  +--+                            
     |  +-------- Sinanodonta **     
     +--+                         
        |  +----- Anodonta__2     
        +--+                      
           |  +-- Anodonta_anatina
           +--+                   
              +-- Anodonta__1     

Guerra et al., 2019 [Publ 4168], fig. 2

iTtTbiriSoTroTroSiD
11014062--
  +----- Sinanodonta **  
--+                   
  |  +-- Anodontina   
  +--+                
     +-- Alasmidontina

Guerra et al., 2019 [Publ 4168], fig. 1

iTtTbiriSoTroTroSiD
11014062--
  +----- Sinanodonta **  
--+                   
  |  +-- Anodontina   
  +--+                
     +-- Alasmidontina

Mezhzherin et al., 2014 [Publ 3705], fig. 2

iTtTbiriSoTroTroSiD
1101331--
  +-- Sinanodonta **
--+              
  +-- Anodontina 

Wu et al., 2022 [Publ 4238], fig. 2e

iTtTbiriSoTroTroSiD
11011632--
  +----- Sinanodonta **  
--+                   
  |  +-- Anodontina   
  +--+                
     +-- Alasmidontina

Wei et al., 2016 [Publ 3665], fig. 0

iTtTbiriSoTroTroSiD
1101511--
-- Cristaria_plicata **

Plazzi et al., 2011 [Publ 3676], fig. 5

iTtTbiriSoTroTroSiD
11011742--
-- Cristaria_plicata **

Zieritz et al., 2012 [Publ 3673], fig. 4

iTtTbiriSoTroTroSiD
1101731--
-- Sinanodonta_woodiana **

Breton et al., 2011 [Publ 3376], fig. 3

iTtTbiriSoTroTroSiD
110136111--
  +-- Cristaria **    
--+                
  +-- Alasmidontina

Huang et al., 2018 [Publ 3828], fig. 6-M

iTtTbiriSoTroTroSiD
11011632--
  +----- Sinanodonta **  
--+                   
  |  +-- Anodontina   
  +--+                
     +-- Alasmidontina

Smith et al., 2018 [Publ 3790], fig. 1

iTtTbiriSoTroTroSiD
110117171--
  +-- Cristaria **    
--+                
  +-- Alasmidontina

Nagel et al., 1996 [Publ 3400], fig. 351

iTtTbiriSoTroTroSiD
1101441--
  +-------------- Pseudanodonta   
--+                               
  |  +----------- Anodonta_cygnea 
  +--+                            
     |  +-------- Sinanodonta **     
     +--+                         
        |  +----- Anodonta__2     
        +--+                      
           |  +-- Anodonta_anatina
           +--+                   
              +-- Anodonta__1     

Chen et al., 2012 [Publ 3624], fig. 4

iTtTbiriSoTroTroSiD
1101611--
  +-- Alasmidontina
--+                
  +-- Cristaria **    

Dai et al., 2024 [Publ 4386], fig. 5-ML

iTtTbiriSoTroTroSiD
11012011--
-- Anemina_arcaeformis **

Mezhzherin et al., 2014 [Publ 3705], fig. 4

iTtTbiriSoTroTroSiD
1101331--
  +-- Sinanodonta **
--+              
  +-- Anodontina 

Froufe et al., 2019 [Publ 3936], fig. 3-M

iTtTbiriSoTroTroSiD
11011832--
  +----- Sinanodonta **  
--+                   
  |  +-- Anodontina   
  +--+                
     +-- Alasmidontina

Soroka, 2010 [Publ 3404], fig. 8a-M

iTtTbiriSoTroTroSiD
1101200--
-- Sinanodonta_woodiana **

Inoue et al., 2018 [Publ 3858], fig. 1_3a-c

iTtTbiriSoTroTroSiD
1101103192--
  +-- Alasmidonta_marginata      
  |                              
  +-- Prolasmidonta              
  |                              
  +-- Alasmidonta_undulata       
  |                              
  +-- Beringiana **                 
  |                              
  +-- Anodonta_kennerlyi         
  |                              
  +-- Anodonta_nuttalliana       
  |                              
  +-- Anodonta_oregonensis       
  |                              
  +-- Arcidens_confragosus       
  |                              
  +-- Arcidens_wheeleri          
  |                              
  +-- Lasmigona_complanata       
  |                              
--+-- Platynaias_compressa       
  |                              
  +-- Lasmigona_costata          
  |                              
  +-- Platynaias_decorata        
  |                              
  +-- Platynaias_subviridis      
  |                              
  +-- Pyganodon_grandis          
  |                              
  +-- Pyganodon_lacustris        
  |                              
  +-- Pseudodontoideus           
  |                              
  +-- Strophitus                 
  |                              
  +-- Utterbackia_imbecillis     
  |                              
  +-- Utterbackia_peninsularis__1
  |                              
  +-- Utterbackia_peninsularis__2

Jiang et al., 2010 [Publ 3438], fig. 5

iTtTbiriSoTroTroSiD
1101300--
-- Cristaria_plicata **

Amaro et al., 2017 [Publ 3791], fig. 2-M

iTtTbiriSoTroTroSiD
1101931--
  +----- Pyganodon  
  |                 
--+  +-- Lasmigona  
  |  |              
  +--+-- Sinanodonta **
     |              
     +-- Utterbackia

Froufe et al., 2017 [Publ 3802], fig. 7

iTtTbiriSoTroTroSiD
1101931--
  +-- Sinanodonta **
--+              
  +-- Anodontina 

Zhou et al., 2017 [Publ 3788], fig. 2

iTtTbiriSoTroTroSiD
11011500--
-- Anemina_arcaeformis **

Dai et al., 2024 [Publ 4386], fig. 5-BI

iTtTbiriSoTroTroSiD
11012011--
-- Anemina_arcaeformis **

Burzynski & Soroka, 2018 [Publ 3862], fig. 5b

iTtTbiriSoTroTroSiD
1101932--
  +----- Sinanodonta **  
--+                   
  |  +-- Anodontina   
  +--+                
     +-- Alasmidontina

Guerra et al., 2019 [Publ 4168], fig. 1-M

iTtTbiriSoTroTroSiD
11012132--
  +----- Sinanodonta **  
--+                   
  |  +-- Anodontina   
  +--+                
     +-- Alasmidontina

Soroka, 2010 [Publ 3404], fig. 4-M

iTtTbiriSoTroTroSiD
1101111--
  +-- Anodontina 
--+              
  +-- Sinanodonta **

Soroka, 2010 [Publ 3404], fig. 2-M

iTtTbiriSoTroTroSiD
1101411--
-- Sinanodonta_woodiana **
 
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