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Page last updated
Thu 3 September 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Search the database pages:

Bivalvia | Unionoida | Unionoidea | Unionidae | Unioninae | Anodontini

subtribe CRISTARIINA Lopes-Lima, Bogan & Froufe in Lopes-Lima et al., 2017: cladomics
     10 genera

Cladomics Summary

number of species = 36 (39 total)
number of subtaxa (rank: genus) = 10
analyzed in 188 cladograms in 110 publications
maximum number of terminals in an analysis = 33 (mean = 4.65, median = 4)
maximum number of subtaxa (genus) = 8 (mean = 2.61, median = 2)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 135
maximum number of included terminals = 33, (mean = 6.09 median = 4)
maximum number of subtaxa (genus) = 8 (mean = 3.24, median = 3)
proportion of cladograms that support basic monophyly = 0.85

Rigorous Monophyly

number of cladograms that test rigorous monophyly = 84
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (genus) taxon and at least one rigorous outgroup (family).
maximum number of included terminals = 27 (mean = 5.2, median = 4)
maximum number of subtaxa (genus) = 7 (mean = 3.4, median = 3)
proportion of cladograms that support rigorous monophyly = 0.83


Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.


PCoA Ordination


NJ Tree


Wu et al., 2025 [Publ 4442], fig. 2

** Top Monophyly Cladogram **

iTtTbiriSoTroTroSiD
33333182000
  +----------------- Cristaria **                  
  |                                             
  |           +----- Beringiana **                 
--+  +--------+                                 
  |  |        |  +-- Acudonta **                   
  |  |        +--+                              
  |  |           +-- Sinanodonta **                
  +--+                                          
     |        +----- Pletholophus_guangzhouensis **
     |  +-----+                                 
     |  |     |  +-- Pletholophus_tenuis **        
     |  |     +--+                              
     +--+        +-- Pletholophus_reinianus **     
        |                                       
        |  +-------- Pletholophus_honglinhensis ** 
        +--+                                    
           |  +----- Anemina **                    
           +--+                                 
              |  +-- Amuranodonta **               
              +--+                              
                 +-- Buldowskia **                 

Whelan et al., 2023 [Publ 4287], fig. 2

** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
44223737229
           +-- Anemina **             
  +--------+                       
  |        +-- Sinanodonta **         
--+                                
  |  +-------- Anodonta_cygnea     
  |  |                             
  +--+     +-- Anodonta_nuttalliana
     |  +--+                       
     |  |  +-- Pseudanodonta       
     +--+                          
        |  +-- Anodonta_anatina    
        +--+                       
           +-- Alasmidontina       

Xiang et al., 2025 [Publ 4433], fig. 6

iTtTbiriSoTroTroSiD
272725810020
  +-------------- Pletholophus **
  |                           
  |        +----- Beringiana **  
--+  +-----+                  
  |  |     |  +-- Mimo **        
  |  |     +--+               
  +--+        +-- Sinanodonta ** 
     |                        
     |  +-------- Cristaria **   
     +--+                     
        |  +----- Anemina **     
        +--+                  
           |  +-- Amuranodonta **
           +--+               
              +-- Buldowskia **  

Dai et al., 2024 [Publ 4385], fig. 3-BI

iTtTbiriSoTroTroSiD
272724740021
        +----- Amuranodonta **
  +-----+                  
  |     |  +-- Anemina **     
  |     +--+               
--+        +-- Buldowskia **  
  |                        
  |  +-------- Cristaria **   
  +--+                     
     |  +----- Pletholophus **
     +--+                  
        |  +-- Beringiana **  
        +--+               
           +-- Sinanodonta ** 

Bogan et al., 2023 [Publ 4295], fig. 3

iTtTbiriSoTroTroSiD
272724741121
  +-------------- Anodontina  
  |                           
--+  +----------- Cristaria **   
  |  |                        
  |  |     +----- Amuranodonta **
  +--+  +--+                  
     |  |  |  +-- Anemina **     
     |  |  +--+               
     +--+     +-- Buldowskia **  
        |                     
        |  +----- Pletholophus **
        +--+                  
           |  +-- Beringiana **  
           +--+               
              +-- Sinanodonta ** 

Dai et al., 2024 [Publ 4385], fig. 3-ML

iTtTbiriSoTroTroSiD
272725740020
        +----- Amuranodonta **
  +-----+                  
  |     |  +-- Anemina **     
  |     +--+               
--+        +-- Buldowskia **  
  |                        
  |  +-------- Cristaria **   
  +--+                     
     |  +----- Pletholophus **
     +--+                  
        |  +-- Beringiana **  
        +--+               
           +-- Sinanodonta ** 

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iTtTbiriSoTroTroSiD
2424197989220
           +-- Pseudanodonta
           |                
  +--------+-- Anodonta     
  |        |                
  |        +-- Alasmidontina
  |                         
--+  +-------- Cristaria **    
  |  |                      
  |  |  +----- Anemina **      
  |  +--+                   
  +--+  |  +-- Amuranodonta ** 
     |  +--+                
     |     +-- Buldowskia **   
     |                      
     |  +----- Pletholophus ** 
     +--+                   
        |  +-- Beringiana **   
        +--+                
           +-- Sinanodonta **  

Sano et al., 2022 [Publ 4311], fig. 3

iTtTbiriSoTroTroSiD
181816500021
        +-- Beringiana **  
  +-----+               
  |     +-- Sinanodonta ** 
--+                     
  |  +----- Pletholophus **
  +--+                  
     |  +-- Cristaria **   
     +--+               
        +-- Buldowskia **  

Wu et al., 2025 [Publ 4442], fig. 6

iTtTbiriSoTroTroSiD
151513820024
  +----------- Cristaria **   
  |                        
  |     +----- Amuranodonta **
--+  +--+                  
  |  |  |  +-- Anemina **     
  |  |  +--+               
  |  |     +-- Buldowskia **  
  +--+                     
     |     +-- Beringiana **  
     |  +--+               
     |  |  +-- Pletholophus **
     +--+                  
        |  +-- Acudonta **    
        +--+               
           +-- Sinanodonta ** 

Douda et al., 2024 [Publ 4364], fig. 1

iTtTbiriSoTroTroSiD
131312210022
  +-- Beringiana ** 
--+              
  +-- Sinanodonta **

Konopleva et al., 2023 [Publ 4269], fig. S1_4

iTtTbiriSoTroTroSiD
1313116433124
  +----------- Anodontina  
  |                        
  |     +----- Anemina **     
--+  +--+                  
  |  |  |  +-- Amuranodonta **
  |  |  +--+               
  +--+     +-- Buldowskia **  
     |                     
     |  +----- Cristaria **   
     +--+                  
        |  +-- Beringiana **  
        +--+               
           +-- Sinanodonta ** 

Bogan et al., 2023 [Publ 4295], fig. 6

iTtTbiriSoTroTroSiD
12129310028
  +----- Pletholophus **
--+                  
  |  +-- Cristaria **   
  +--+               
     +-- Sinanodonta ** 

Sano et al., 2022 [Publ 4311], fig. 2

iTtTbiriSoTroTroSiD
121212500033
        +-- Beringiana **  
  +-----+               
  |     +-- Sinanodonta ** 
--+                     
  |  +----- Pletholophus **
  +--+                  
     |  +-- Cristaria **   
     +--+               
        +-- Buldowskia **  

Lopes-Lima et al., 2020 [Publ 4080], fig. 10

iTtTbiriSoTroTroSiD
12128220025
  +-- Beringiana ** 
--+              
  +-- Sinanodonta **

Sano et al., 2020 [Publ 4138], fig. 2

iTtTbiriSoTroTroSiD
1135366578251
                                +-- Anemina **      
  +-----------------------------+                
  |                             +-- Buldowskia **   
  |                                              
--+                          +----- Pletholophus ** 
  |  +-----------------------+                   
  |  |                       |  +-- Beringiana **   
  |  |                       +--+                
  +--+                          +-- Sinanodonta **  
     |                                           
     |  +-------------------------- Cristaria **    
     |  |                                        
     +--+                       +-- Anodontina   
        |  +--------------------+                
        |  |                    +-- Alasmidontina
        +--+                                     
           |  +-------------------- Lanceolaria  
           +--+                                  
              |  +----------------- Lepidodesma  
              +--+                               
                 |  +-------------- Aculamprotula
                 +--+                            
                    |  +----------- Unio         
                    +--+                         
                       |  +-------- Inversiunio  
                       +--+                      
                          |  +----- Nodularia    
                          +--+                   
                             |  +-- Tchangsinaia 
                             +--+                
                                +-- Schistodesmus

Sano et al., 2020 [Publ 4138], fig. 4

iTtTbiriSoTroTroSiD
1135376578253
                                +-- Anemina **      
  +-----------------------------+                
  |                             +-- Buldowskia **   
  |                                              
--+                          +----- Sinanodonta **  
  |  +-----------------------+                   
  |  |                       |  +-- Pletholophus ** 
  |  |                       +--+                
  +--+                          +-- Beringiana **   
     |                                           
     |  +-------------------------- Cristaria **    
     |  |                                        
     +--+                       +-- Anodontina   
        |  +--------------------+                
        |  |                    +-- Alasmidontina
        +--+                                     
           |  +-------------------- Lepidodesma  
           +--+                                  
              |  +----------------- Lanceolaria  
              +--+                               
                 |  +-------------- Aculamprotula
                 +--+                            
                    |  +----------- Unio         
                    +--+                         
                       |  +-------- Nodularia    
                       +--+                      
                          |  +----- Inversiunio  
                          +--+                   
                             |  +-- Schistodesmus
                             +--+                
                                +-- Tchangsinaia 

Sano et al., 2020 [Publ 4138], fig. 3

iTtTbiriSoTroTroSiD
1135376578253
                                +-- Anemina **      
  +-----------------------------+                
  |                             +-- Buldowskia **   
  |                                              
--+                          +----- Sinanodonta **  
  |  +-----------------------+                   
  |  |                       |  +-- Pletholophus ** 
  |  |                       +--+                
  +--+                          +-- Beringiana **   
     |                                           
     |  +-------------------------- Cristaria **    
     |  |                                        
     +--+                       +-- Anodontina   
        |  +--------------------+                
        |  |                    +-- Alasmidontina
        +--+                                     
           |  +-------------------- Lanceolaria  
           +--+                                  
              |  +----------------- Lepidodesma  
              +--+                               
                 |  +-------------- Aculamprotula
                 +--+                            
                    |  +----------- Unio         
                    +--+                         
                       |  +-------- Inversiunio  
                       +--+                      
                          |  +----- Nodularia    
                          +--+                   
                             |  +-- Schistodesmus
                             +--+                
                                +-- Tchangsinaia 

Froufe et al., 2017 [Publ 3802], fig. 6

iTtTbiriSoTroTroSiD
10107422126
  +----------- Anodontina 
--+                       
  |  +-------- Anemina **    
  +--+                    
     |  +----- Cristaria **  
     +--+                 
        |  +-- Beringiana ** 
        +--+              
           +-- Sinanodonta **

Sano et al., 2020 [Publ 4138], fig. 1

iTtTbiriSoTroTroSiD
102256240047
  +----- Lanceolaria_grayii__1  
  |                             
  +----- Lanceolaria_grayii__2  
  |                             
  +----- Cristaria_plicata__1 **   
  |                             
  +----- Cristaria_plicata__2 **   
  |                             
  +----- Sinanodonta_lauta **      
  |                             
  +----- Beringiana_japonica **    
  |                             
  +----- Beringiana_fukuharai__1 **
  |                             
  +----- Beringiana_beringiana **  
  |                             
  +----- Beringiana_fukuharai__2 **
  |                             
  +----- Beringiana_fukuharai__3 **
  |                             
  +----- Buldowskia_iwakawai__1 ** 
  |                             
  +----- Buldowskia_iwakawai__2 ** 
  |                             
  +----- Sinanodonta_tumens__1 **  
--+                             
  +----- Sinanodonta_tumens__2 **  
  |                             
  +----- Pletholophus **           
  |                             
  +----- Anemina **                
  |                             
  +----- Lanceolaria_lanceolata 
  |                             
  +----- Cristaria_plicata__3 **   
  |                             
  +----- Lanceolaria_gladiola   
  |                             
  +----- Sinanodonta_angula **     
  |                             
  +----- Lepidodesma            
  |                             
  |  +-- Aculamprotula          
  +--+                          
  |  +-- Acuticosta             
  |                             
  |  +-- Cuneopsis              
  +--+                          
  |  +-- Nodularia              
  |                             
  |  +-- Schistodesmus          
  +--+                          
     +-- Inversiunio            

Lopes-Lima et al., 2020 [Publ 4080], fig. 6

iTtTbiriSoTroTroSiD
997420030
  +-------- Sinanodonta ** 
--+                     
  |  +----- Amuranodonta **
  +--+                  
     |  +-- Anemina **     
     +--+               
        +-- Buldowskia **  

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
997419211228
     +-------- Anemina **      
  +--+                      
  |  |  +----- Cristaria **    
  |  +--+                   
  |     |  +-- Pletholophus ** 
  |     +--+                
  |        +-- Sinanodonta **  
--+                         
  |     +----- Alasmidonta  
  |     |                   
  |     +----- Lasmigona    
  |     |                   
  |     +----- Pseudanodonta
  |     |                   
  +-----+----- Simpsonaias  
        |                   
        +----- Strophitus   
        |                   
        +----- Anodonta     
        |                   
        |  +-- Pyganodon    
        +--+                
           +-- Utterbackia  

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
94724419211265
  +-------- Acuticosta           
  |                              
  +-------- Alasmidonta          
  |                              
  +-------- Anodonta_anatina     
  |                              
  +-------- Anemina **              
  |                              
  +-------- Anodonta_cygnea      
  |                              
  +-------- Cristaria **            
  |                              
  +-------- Anodonta_nuttalliana 
  |                              
  +-------- Lasmigona            
  |                              
  +-------- Lepidodesma          
  |                              
  +-------- Pseudanodonta        
  |                              
  +-------- Pyganodon            
  |                              
--+-------- Simpsonaias          
  |                              
  +-------- Strophitus           
  |                              
  +-------- Utterbackia          
  |                              
  +-------- Lanceolaria          
  |                              
  +-------- Schistodesmus        
  |                              
  +-------- Unio                 
  |                              
  +-------- Aculamprotula        
  |                              
  |     +-- Pletholophus **         
  +-----+                        
  |     +-- Sinanodonta **          
  |                              
  |  +----- Cuneopsis_celtiformis
  |  |                           
  |  +----- Tchangsinaia         
  |  |                           
  +--+----- Cuneopsis_rufescens  
     |                           
     +----- Nodularia            
     |                           
     |  +-- Cuneopsis_szechenyii 
     +--+                        
        +-- Cuneopsis_heudei     

Bolotov et al., 2020 [Publ 4084], fig. 1

iTtTbiriSoTroTroSiD
9976273128
  +----------- Anodontina  
  |                        
  |     +----- Amuranodonta **
--+  +--+                  
  |  |  |  +-- Anemina **     
  |  |  +--+               
  +--+     +-- Buldowskia **  
     |                     
     |  +----- Cristaria **   
     +--+                  
        |  +-- Beringiana **  
        +--+               
           +-- Sinanodonta ** 

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
997419011228
        +-------- Anemina **      
  +-----+                      
  |     |  +----- Cristaria **    
  |     +--+                   
  |        |  +-- Pletholophus ** 
  |        +--+                
--+           +-- Sinanodonta **  
  |                            
  |  +----------- Anodonta     
  |  |                         
  |  |        +-- Alasmidonta  
  |  +--------+                
  +--+        +-- Lasmigona    
     |                         
     |        +-- Pyganodon    
     |  +-----+                
     |  |     +-- Utterbackia  
     +--+                      
        |  +----- Pseudanodonta
        +--+                   
           |  +-- Simpsonaias  
           +--+                
              +-- Strophitus   

Sano et al., 2017 [Publ 3799], fig. 2

iTtTbiriSoTroTroSiD
9975240032
        +-- Cristaria **   
  +-----+               
  |     +-- Buldowskia **  
--+                     
  |  +----- Beringiana **  
  +--+                  
     |  +-- Pletholophus **
     +--+               
        +-- Sinanodonta ** 

Huang et al., 2019 [Publ 3843], fig. 1

iTtTbiriSoTroTroSiD
831293370052
  +----------------------------------- Anemina **              
  |                                                         
--+  +-------------------------------- Acuticosta           
  |  |                                                      
  +--+  +----------------------------- Sinanodonta **          
     |  |                                                   
     +--+  +-------------------------- Aculamprotula        
        |  |                                                
        +--+                       +-- Cristaria **            
           |  +--------------------+                        
           |  |                    +-- Lanceolaria          
           +--+                                             
              |  +-------------------- Lepidodesma          
              +--+                                          
                 |  +----------------- Schistodesmus        
                 +--+                                       
                    |  +-------------- Nodularia            
                    +--+                                    
                       |  +----------- Cuneopsis_rufescens  
                       +--+                                 
                          |  +-------- Tchangsinaia         
                          +--+                              
                             |  +----- Cuneopsis_celtiformis
                             +--+                           
                                |  +-- Cuneopsis_szechenyii 
                                +--+                        
                                   +-- Cuneopsis_heudei     

Bespalaya et al., 2018 [Publ 3811], fig. 3

iTtTbiriSoTroTroSiD
887220032
  +-- Anemina **    
--+              
  +-- Sinanodonta **

Kondakov et al., 2018 [Publ 3914], fig. 5

iTtTbiriSoTroTroSiD
886200025
  +-- Anemina **    
--+              
  +-- Sinanodonta **

Sano et al., 2017 [Publ 3799], fig. 1

iTtTbiriSoTroTroSiD
8865130029
        +-- Cristaria **   
  +-----+               
  |     +-- Buldowskia **  
--+                     
  |  +----- Pletholophus **
  +--+                  
     |  +-- Sinanodonta ** 
     +--+               
        +-- Beringiana **  

Vikhrev et al., 2017 [Publ 3810], fig. 4

iTtTbiriSoTroTroSiD
886220025
  +-- Anemina **    
--+              
  +-- Sinanodonta **

Huang et al., 2019 [Publ 3843], fig. S1

iTtTbiriSoTroTroSiD
833383390067
  +-------------------------- Anemina **      
  |                                        
--+  +----------------------- Sinanodonta **  
  |  |                                     
  +--+  +-------------------- Acuticosta   
     |  |                                  
     +--+                 +-- Lepidodesma  
        |  +--------------+                
        |  |              +-- Cristaria **    
        +--+                               
           |  +-------------- Lanceolaria  
           +--+                            
              |  +----------- Aculamprotula
              +--+                         
                 |  +-------- Nodularia    
                 +--+                      
                    |  +----- Tchangsinaia 
                    +--+                   
                       |  +-- Cuneopsis    
                       +--+                
                          +-- Schistodesmus

Bauer et al., 2021 [Publ 4365], fig. 2

iTtTbiriSoTroTroSiD
885200026
  +-- Anemina **    
--+              
  +-- Sinanodonta **

Xiang et al., 2025 [Publ 4433], fig. 7

iTtTbiriSoTroTroSiD
775562128
  +-------------- Anodontina 
--+                          
  |  +----------- Cristaria **  
  +--+                       
     |  +-------- Anemina **    
     +--+                    
        |  +----- Beringiana ** 
        +--+                 
           |  +-- Mimo **       
           +--+              
              +-- Sinanodonta **

Wu et al., 2024 [Publ 4379], fig. 5

iTtTbiriSoTroTroSiD
66444712227
           +-------- Cristaria **           
  +--------+                             
  |        |  +----- Anemina **             
  |        +--+                          
  |           |  +-- Beringiana **          
--+           +--+                       
  |              +-- Sinanodonta **         
  |                                      
  |  +-------------- Anodonta_nuttalliana
  +--+                                   
     |  +----------- Alasmidontina       
     +--+                                
        |  +-------- Pseudanodonta       
        +--+                             
           |  +----- Anodonta_anatina    
           +--+                          
              |  +-- Anodonta_cygnea     
              +--+                       
                 +-- Anodonta_exulcerata 

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iTtTbiriSoTroTroSiD
66448612227
           +-------- Cristaria **           
  +--------+                             
  |        |  +----- Anemina **             
  |        +--+                          
  |           |  +-- Beringiana **          
--+           +--+                       
  |              +-- Sinanodonta **         
  |                                      
  |  +-------------- Anodonta_nuttalliana
  +--+                                   
     |  +----------- Alasmidontina       
     +--+                                
        |  +-------- Pseudanodonta       
        +--+                             
           |  +----- Anodonta_anatina    
           +--+                          
              |  +-- Anodonta_cygnea     
              +--+                       
                 +-- Anodonta_exulcerata 

Lopes-Lima et al., 2020 [Publ 4080], fig. 7

iTtTbiriSoTroTroSiD
664220027
  +-- Sinanodonta **
--+              
  +-- Beringiana ** 

Dai et al., 2025 [Publ 4440], fig. 2

iTtTbiriSoTroTroSiD
636306394261
                                   +-- Anemina **               
  +--------------------------------+                         
  |                                +-- Amuranodonta **          
  |                                                          
  |                             +----- Pletholophus **          
--+  +--------------------------+                            
  |  |                          |  +-- Sinanodonta **           
  |  |                          +--+                         
  |  |                             +-- Beringiana **            
  +--+                                                       
     |                             +-- Alasmidontina         
     |  +--------------------------+                         
     |  |                          +-- Anodontina            
     |  |                                                    
     +--+  +-------------------------- Lanceolaria_gladiola  
        |  |                                                 
        |  +-------------------------- Lanceolaria_oxyrhyncha
        |  |                                                 
        +--+-------------------------- Cristaria **             
           |                                                 
           |  +----------------------- Acuticosta            
           |  |                                              
           |  +----------------------- Lepidodesma           
           +--+                                              
              |  +-------------------- Globunio              
              |  |                                           
              +--+  +----------------- Aculamprotula         
                 |  |                                        
                 |  |              +-- Middendorffinaia      
                 +--+  +-----------+                         
                    |  |           +-- Nodularia             
                    |  |                                     
                    +--+  +----------- Cuneopsis             
                       |  |                                  
                       |  |        +-- Diaurora              
                       +--+--------+                         
                          |        +-- Schistodesmus         
                          |                                  
                          |  +-------- Pseudocuneopsis       
                          +--+                               
                             |  +----- Inversiunio           
                             +--+                            
                                |  +-- Tchangsinaia          
                                +--+                         
                                   +-- Unio                  

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iTtTbiriSoTroTroSiD
66449012229
        +-- Anodontina   
  +-----+                
  |     +-- Alasmidontina
  |                      
--+     +-- Anemina **      
  |  +--+                
  |  |  +-- Cristaria **    
  +--+                   
     |  +-- Beringiana **   
     +--+                
        +-- Sinanodonta **  

Wu et al., 2018 [Publ 3836], fig. 2

iTtTbiriSoTroTroSiD
6633477228
        +-- Anemina **         
        |                   
  +-----+-- Cristaria **       
  |     |                   
--+     +-- Sinanodonta **     
  |                         
  |  +----- Anodonta_cygnea 
  +--+                      
     |  +-- Anodonta_anatina
     +--+                   
        +-- Alasmidontina   

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iTtTbiriSoTroTroSiD
66449012227
     +-------- Cristaria **           
  +--+                             
  |  |  +----- Anemina **             
  |  +--+                          
  |     |  +-- Beringiana **          
  |     +--+                       
--+        +-- Sinanodonta **         
  |                                
  |  +-------- Anodonta_nuttalliana
  |  |                             
  |  +-------- Pseudanodonta       
  +--+                             
     |  +----- Anodonta_cygnea     
     |  |                          
     +--+----- Alasmidontina       
        |                          
        |  +-- Anodonta_exulcerata 
        +--+                       
           +-- Anodonta_anatina    

Chen et al., 2023 [Publ 4302], fig. 2

iTtTbiriSoTroTroSiD
6646354229
              +-- Alasmidontina
  +-----------+                
  |           +-- Anodontina   
--+                            
  |  +----------- Cristaria **    
  |  |                         
  +--+        +-- Anemina **      
     |  +-----+                
     |  |     +-- Amuranodonta ** 
     +--+                      
        |  +----- Pletholophus ** 
        +--+                   
           |  +-- Sinanodonta **  
           +--+                
              +-- Beringiana **   

Bolotov et al., 2017 [Publ 3761], fig. S1

iTtTbiriSoTroTroSiD
66431339227
           +----- Cristaria **           
  +--------+                          
  |        |  +-- Pletholophus **        
  |        +--+                       
--+           +-- Sinanodonta **         
  |                                   
  |  +----------- Anodonta_anatina    
  +--+                                
     |  +-------- Alasmidontina       
     +--+                             
        |  +----- Pseudanodonta       
        +--+                          
           |  +-- Anodonta_cygnea     
           +--+                       
              +-- Anodonta_nuttalliana

Bolotov et al., 2016 [Publ 3732], fig. 2

iTtTbiriSoTroTroSiD
665120032
1

Bogan et al., 2023 [Publ 4295], fig. 4

iTtTbiriSoTroTroSiD
664310029
  +----- Cristaria **   
--+                  
  |  +-- Sinanodonta ** 
  +--+               
     +-- Pletholophus **

Dai et al., 2025 [Publ 4440], fig. 1

iTtTbiriSoTroTroSiD
639326394264
                                         +-- Anemina **         
  +--------------------------------------+                   
  |                                      +-- Amuranodonta **    
  |                                                          
  |                                   +----- Parreysia       
  +-----------------------------------+                      
  |                                   |  +-- Coelatura       
  |                                   +--+                   
--+                                      +-- Indonaia        
  |                                                          
  |                                   +----- Pletholophus **    
  |  +--------------------------------+                      
  |  |                                |  +-- Sinanodonta **     
  |  |                                +--+                   
  |  |                                   +-- Beringiana **      
  +--+                                                       
     |  +----------------------------------- Cristaria **       
     |  |                                                    
     |  |                                +-- Alasmidontina   
     |  +--------------------------------+                   
     +--+                                +-- Anodontina      
        |                                                    
        |  +-------------------------------- Lanceolaria     
        |  |                                                 
        +--+  +----------------------------- Lepidodesma     
           |  |                                              
           +--+  +-------------------------- Acuticosta      
              |  |                                           
              +--+  +----------------------- Globunio        
                 |  |                                        
                 +--+  +-------------------- Aculamprotula   
                    |  |                                     
                    +--+  +----------------- Middendorffinaia
                       |  |                                  
                       +--+  +-------------- Nodularia       
                          |  |                               
                          +--+  +----------- Cuneopsis       
                             |  |                            
                             |  |        +-- Diaurora        
                             +--+--------+                   
                                |        +-- Schistodesmus   
                                |                            
                                |  +-------- Pseudocuneopsis 
                                +--+                         
                                   |  +----- Unio            
                                   +--+                      
                                      |  +-- Tchangsinaia    
                                      +--+                   
                                         +-- Inversiunio     

Bolotov et al., 2017 [Publ 3761], fig. S2

iTtTbiriSoTroTroSiD
66431339227
  +----- Cristaria **   
--+                  
  |  +-- Pletholophus **
  +--+               
     +-- Sinanodonta ** 

Chen et al., 2023 [Publ 4302], fig. 1

iTtTbiriSoTroTroSiD
632246354257
                          +-- Anemina **               
  +-----------------------+                         
  |                       +-- Amuranodonta **          
  |                                                 
  |                    +----- Pletholophus **          
  |  +-----------------+                            
--+  |                 |  +-- Sinanodonta **           
  |  |                 +--+                         
  |  |                    +-- Beringiana **            
  |  |                                              
  |  |  +-------------------- Lanceolaria_gladiola  
  +--+  |                                           
     |  +-------------------- Acuticosta            
     |  |                                           
     |  +-------------------- Lepidodesma           
     |  |                                           
     |  +-------------------- Lanceolaria_oxyrhyncha
     |  |                                           
     +--+-------------------- Cristaria **             
        |                                           
        |                 +-- Alasmidontina         
        +-----------------+                         
        |                 +-- Anodontina            
        |                                           
        |  +----------------- Aculamprotula         
        |  |                                        
        |  |              +-- Middendorffinaia      
        +--+  +-----------+                         
           |  |           +-- Nodularia             
           |  |                                     
           +--+  +----------- Cuneopsis             
              |  |                                  
              |  |        +-- Diaurora              
              +--+--------+                         
                 |        +-- Schistodesmus         
                 |                                  
                 |  +-------- Pseudocuneopsis       
                 +--+                               
                    |  +----- Inversiunio           
                    +--+                            
                       |  +-- Tchangsinaia          
                       +--+                         
                          +-- Unio                  

Bolotov et al., 2017 [Publ 3764], fig. S1

iTtTbiriSoTroTroSiD
617031199242
  +-- Alasmidonta          
  |                        
  +-- Anodonta_anatina     
  |                        
  +-- Anodonta_cygnea      
  |                        
  +-- Anodonta_nuttalliana 
  |                        
  +-- Lasmigona            
  |                        
  +-- Pseudanodonta        
  |                        
  +-- Pyganodon            
  |                        
  +-- Simpsonaias          
  |                        
  +-- Strophitus           
--+                        
  +-- Sinanodonta_lauta **    
  |                        
  +-- Cristaria_plicata **    
  |                        
  +-- Cristaria_radiata **    
  |                        
  +-- Pletholophus **         
  |                        
  +-- Sinanodonta_lucida **   
  |                        
  +-- Sinanodonta_jourdyi **  
  |                        
  +-- Lanceolaria_gladiola 
  |                        
  +-- Lanceolaria_grayii__1
  |                        
  +-- Lanceolaria_grayii__2

Hou et al., 2025 [Publ 4445], fig. 5

iTtTbiriSoTroTroSiD
66444712227
           +-------- Cristaria **           
  +--------+                             
  |        |  +----- Anemina **             
  |        +--+                          
  |           |  +-- Beringiana **          
--+           +--+                       
  |              +-- Sinanodonta **         
  |                                      
  |  +-------------- Anodonta_nuttalliana
  +--+                                   
     |  +----------- Alasmidontina       
     +--+                                
        |  +-------- Pseudanodonta       
        +--+                             
           |  +----- Anodonta_anatina    
           +--+                          
              |  +-- Anodonta_cygnea     
              +--+                       
                 +-- Anodonta_exulcerata 

Hou et al., 2024 [Publ 4400], fig. 4

iTtTbiriSoTroTroSiD
66447012227
        +-------- Cristaria **           
  +-----+                             
  |     |  +----- Anemina **             
  |     +--+                          
  |        |  +-- Beringiana **          
  |        +--+                       
--+           +-- Sinanodonta **         
  |                                   
  |           +-- Anodonta_nuttalliana
  |  +--------+                       
  |  |        +-- Alasmidontina       
  +--+                                
     |  +-------- Pseudanodonta       
     +--+                             
        |  +----- Anodonta_anatina    
        +--+                          
           |  +-- Anodonta_cygnea     
           +--+                       
              +-- Anodonta_exulcerata 

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iTtTbiriSoTroTroSiD
66448612227
        +-------- Cristaria **           
  +-----+                             
  |     |  +----- Anemina **             
  |     +--+                          
  |        |  +-- Beringiana **          
  |        +--+                       
--+           +-- Sinanodonta **         
  |                                   
  |           +-- Anodonta_nuttalliana
  |  +--------+                       
  |  |        +-- Alasmidontina       
  +--+                                
     |  +-------- Pseudanodonta       
     +--+                             
        |  +----- Anodonta_anatina    
        +--+                          
           |  +-- Anodonta_cygnea     
           +--+                       
              +-- Anodonta_exulcerata 

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

iTtTbiriSoTroTroSiD
5533719228
        +-- Anodontina   
  +-----+                
  |     +-- Alasmidontina
--+                      
  |  +----- Cristaria **    
  +--+                   
     |  +-- Pletholophus ** 
     +--+                
        +-- Sinanodonta **  

Mabuchi & Nishida, 2020 [Publ 4120], fig. 1

iTtTbiriSoTroTroSiD
555370035
  +-------------- Cristaria **               
  |                                       
  |  +----------- Sinanodonta_tumens **      
  |  |                                    
  |  |        +-- Sinanodonta_lauta__5 **    
  |  |        |                           
--+  |        +-- Sinanodonta_lauta__6 **    
  |  |        |                           
  |  |        +-- Beringiana_japonica__15 ** 
  |  |        |                           
  |  |        +-- Beringiana_japonica__16 ** 
  |  |  +-----+                           
  |  |  |     +-- Beringiana_japonica__17 ** 
  +--+  |     |                           
     |  |     +-- Beringiana_japonica__18 ** 
     |  |     |                           
     |  |     +-- Beringiana_japonica__19 ** 
     |  |     |                           
     |  |     +-- Beringiana_japonica__20 ** 
     |  |                                 
     |  |  +----- Beringiana_japonica__1 **  
     |  |  |                              
     |  |  +----- Beringiana_japonica__2 **  
     |  |  |                              
     +--+  +----- Beringiana_japonica__3 **  
        |  |                              
        |  +----- Beringiana_japonica__4 **  
        |  |                              
        |  +----- Beringiana_japonica__5 **  
        |  |                              
        |  +----- Sinanodonta_lauta__1 **    
        |  |                              
        |  +----- Beringiana_japonica__6 **  
        |  |                              
        |  +----- Sinanodonta_lauta__2 **    
        |  |                              
        |  +----- Sinanodonta_calipygos__1 **
        |  |                              
        +--+----- Beringiana_japonica__7 **  
           |                              
           +----- Sinanodonta_lauta__4 **    
           |                              
           +----- Beringiana_japonica__8 **  
           |                              
           +----- Beringiana_japonica__9 **  
           |                              
           +----- Beringiana_japonica__10 ** 
           |                              
           +----- Beringiana_japonica__11 ** 
           |                              
           +----- Beringiana_japonica__12 ** 
           |                              
           +----- Beringiana_japonica__13 ** 
           |                              
           +----- Beringiana_japonica__14 ** 
           |                              
           |  +-- Sinanodonta_lauta__3 **    
           +--+                           
              +-- Sinanodonta_calipygos__2 **

Bogan et al., 2023 [Publ 4295], fig. 5

iTtTbiriSoTroTroSiD
553310028
  +----- Cristaria **   
--+                  
  |  +-- Sinanodonta ** 
  +--+               
     +-- Pletholophus **

Lopes-Lima et al., 2020 [Publ 4080], fig. 8

iTtTbiriSoTroTroSiD
553220028
  +-- Sinanodonta **
--+              
  +-- Cristaria **  

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iTtTbiriSoTroTroSiD
5534759230
           +-- Anodontina   
  +--------+                
  |        +-- Alasmidontina
--+                         
  |  +-------- Cristaria **    
  +--+                      
     |  +----- Anemina **      
     +--+                   
        |  +-- Pletholophus ** 
        +--+                
           +-- Sinanodonta **  

Soroka, 2010 [Publ 3404], fig. 9a

iTtTbiriSoTroTroSiD
553300028
  +----- Buldowskia ** 
--+                 
  |  +-- Beringiana ** 
  +--+              
     +-- Sinanodonta **

Chen et al., 2025 [Publ 4448], fig. 3

iTtTbiriSoTroTroSiD
5533816228
        +-- Anodontina   
  +-----+                
  |     +-- Alasmidontina
--+                      
  |  +----- Cristaria **    
  +--+                   
     |  +-- Anemina **      
     +--+                
        +-- Sinanodonta **  

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

iTtTbiriSoTroTroSiD
5533719228
        +-- Anodontina   
  +-----+                
  |     +-- Alasmidontina
--+                      
  |  +----- Cristaria **    
  +--+                   
     |  +-- Pletholophus ** 
     +--+                
        +-- Sinanodonta **  

Soroka, 2010 [Publ 3404], fig. 9b

iTtTbiriSoTroTroSiD
442200029
  +-- Buldowskia ** 
--+              
  +-- Sinanodonta **

Wu et al., 2024 [Publ 4388], fig. 6

iTtTbiriSoTroTroSiD
4424503229
           +-- Anodontina   
  +--------+                
  |        +-- Alasmidontina
--+                         
  |  +-------- Cristaria **    
  +--+                      
     |  +----- Anemina **      
     +--+                   
        |  +-- Beringiana **   
        +--+                
           +-- Sinanodonta **  

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

iTtTbiriSoTroTroSiD
4423352229
        +-- Anodontina   
  +-----+                
  |     +-- Alasmidontina
--+                      
  |  +----- Cristaria **    
  +--+                   
     |  +-- Anemina **      
     +--+                
        +-- Sinanodonta **  

Wu et al., 2019 [Publ 3864], fig. 3

iTtTbiriSoTroTroSiD
4423425229
        +-- Anodontina   
  +-----+                
  |     +-- Alasmidontina
--+                      
  |  +----- Cristaria **    
  +--+                   
     |  +-- Anemina **      
     +--+                
        +-- Sinanodonta **  

Soroka, 2010 [Publ 3404], fig. 9c

iTtTbiriSoTroTroSiD
443200032
  +-- Buldowskia ** 
--+              
  +-- Sinanodonta **

Wu et al., 2022 [Publ 4238], fig. 3

iTtTbiriSoTroTroSiD
4423706229
        +-- Anodontina   
  +-----+                
  |     +-- Alasmidontina
--+                      
  |  +----- Cristaria **    
  +--+                   
     |  +-- Anemina **      
     +--+                
        +-- Sinanodonta **  

Sayenko et al., 2017 [Publ 3760], fig. 3

iTtTbiriSoTroTroSiD
442200029
  +-- Beringiana ** 
--+              
  +-- Sinanodonta **

Wang et al., 2018 [Publ 3846], fig. 2

iTtTbiriSoTroTroSiD
4423273229
        +-- Anodontina   
  +-----+                
  |     +-- Alasmidontina
--+                      
  |  +----- Cristaria **    
  +--+                   
     |  +-- Anemina **      
     +--+                
        +-- Sinanodonta **  

Huang et al., 2019 [Publ 3843], fig. 3

iTtTbiriSoTroTroSiD
4423526229
     +----- Cristaria **       
  +--+                      
  |  |  +-- Anemina **         
  |  +--+                   
--+     +-- Sinanodonta **     
  |                         
  |  +----- Anodonta_cygnea 
  +--+                      
     |  +-- Anodonta_anatina
     +--+                   
        +-- Alasmidontina   

Lopes-Lima et al., 2020 [Publ 4080], fig. 9

iTtTbiriSoTroTroSiD
442320029
  +----- Cristaria **   
--+                  
  |  +-- Sinanodonta ** 
  +--+               
     +-- Pletholophus **

Soroka & Burzynski, 2017 [Publ 3837], fig. 1

iTtTbiriSoTroTroSiD
4423196229
        +-- Anodontina   
  +-----+                
  |     +-- Alasmidontina
--+                      
  |  +----- Cristaria **    
  +--+                   
     |  +-- Anemina **      
     +--+                
        +-- Sinanodonta **  

Riccardi et al., 2020 [Publ 4089], fig. 2

iTtTbiriSoTroTroSiD
4423159229
           +----- Cristaria **           
  +--------+                          
  |        |  +-- Anemina **             
  |        +--+                       
--+           +-- Sinanodonta **         
  |                                   
  |  +----------- Anodonta_anatina    
  |  |                                
  +--+        +-- Anodonta_nuttalliana
     |  +-----+                       
     |  |     +-- Alasmidontina       
     +--+                             
        |  +----- Pseudanodonta       
        +--+                          
           |  +-- Anodonta_exulcerata 
           +--+                       
              +-- Anodonta_cygnea     

Fonseca et al., 2016 [Publ 3775], fig. 1

iTtTbiriSoTroTroSiD
4423275229
        +-- Anodontina   
  +-----+                
  |     +-- Alasmidontina
--+                      
  |  +----- Cristaria **    
  +--+                   
     |  +-- Anemina **      
     +--+                
        +-- Sinanodonta **  

Wu et al., 2023 [Publ 4339], fig. 5

iTtTbiriSoTroTroSiD
4423155231
        +-- Alasmidontina
  +-----+                
  |     +-- Anodontina   
--+                      
  |  +----- Anemina **      
  +--+                   
     |  +-- Cristaria **    
     +--+                
        +-- Sinanodonta **  

Wu et al., 2021 [Publ 4366], fig. 2_I

iTtTbiriSoTroTroSiD
4423876231
           +----- Anemina **      
  +--------+                   
  |        |  +-- Cristaria **    
  |        +--+                
--+           +-- Sinanodonta **  
  |                            
  |  +----------- Lasmigona    
  +--+                         
     |  +-------- Anodonta     
     +--+                      
        |  +----- Pseudanodonta
        +--+                   
           |  +-- Pyganodon    
           +--+                
              +-- Strophitus   

Wu et al., 2021 [Publ 4366], fig. 3-BI

iTtTbiriSoTroTroSiD
4423709229
     +----- Cristaria **       
  +--+                      
  |  |  +-- Anemina **         
  |  +--+                   
--+     +-- Sinanodonta **     
  |                         
  |  +----- Anodonta_cygnea 
  +--+                      
     |  +-- Anodonta_anatina
     +--+                   
        +-- Alasmidontina   

Fukata & Iigo, 2020 [Publ 4106], fig. 1

iTtTbiriSoTroTroSiD
4423506229
        +-- Anodontina   
  +-----+                
  |     +-- Alasmidontina
--+                      
  |  +----- Cristaria **    
  +--+                   
     |  +-- Anemina **      
     +--+                
        +-- Sinanodonta **  

Xue et al., 2016 [Publ 3659], fig. 1

iTtTbiriSoTroTroSiD
4533111130
     +-- Cristaria **    
  +--+                
  |  +-- Alasmidontina
--+                   
  |  +-- Anemina **      
  +--+                
     +-- Sinanodonta **  

Galal-Khallaf et al., 2024 [Publ 4459], fig. 6

iTtTbiriSoTroTroSiD
4422249231
              +-- Beringiana **             
  +-----------+                          
  |           +-- Sinanodonta **            
  |                                      
--+           +-- Anodonta_exulcerata    
  |  +--------+                          
  |  |        +-- Anodonta_cygnea        
  |  |                                   
  +--+        +-- Anodonta_californiensis
     |  +-----+                          
     |  |     +-- Anodonta_nuttalliana   
     |  |                                
     +--+     +-- Anodonta_kennerlyi     
        |  +--+                          
        |  |  +-- Anodonta_oregonensis   
        +--+                             
           |  +-- Anodonta_anatina       
           +--+                          
              +-- Alasmidontina          

Cha et al., 2018 [Publ 3916], fig. 1

iTtTbiriSoTroTroSiD
4423345229
        +-- Anodontina   
  +-----+                
  |     +-- Alasmidontina
--+                      
  |  +----- Cristaria **    
  +--+                   
     |  +-- Anemina **      
     +--+                
        +-- Sinanodonta **  

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

iTtTbiriSoTroTroSiD
4423445229
        +-- Anodontina   
  +-----+                
  |     +-- Alasmidontina
--+                      
  |  +----- Cristaria **    
  +--+                   
     |  +-- Anemina **      
     +--+                
        +-- Sinanodonta **  

Aunins et al., 2018 [Publ 3863], fig. 1

iTtTbiriSoTroTroSiD
4423367229
        +-- Anodontina   
  +-----+                
  |     +-- Alasmidontina
--+                      
  |  +----- Cristaria **    
  +--+                   
     |  +-- Anemina **      
     +--+                
        +-- Sinanodonta **  

Klishko et al., 2016 [Publ 3733], fig. 1

iTtTbiriSoTroTroSiD
442300029
  +----- Cristaria **  
--+                 
  |  +-- Sinanodonta **
  +--+              
     +-- Beringiana ** 

Wu et al., 2021 [Publ 4366], fig. 3-ML

iTtTbiriSoTroTroSiD
4423709229
     +----- Cristaria **       
  +--+                      
  |  |  +-- Anemina **         
  |  +--+                   
--+     +-- Sinanodonta **     
  |                         
  |  +----- Anodonta_cygnea 
  +--+                      
     |  +-- Anodonta_anatina
     +--+                   
        +-- Alasmidontina   

Froufe et al., 2019 [Publ 3936], fig. 6

iTtTbiriSoTroTroSiD
4423405229
        +-- Anodontina   
  +-----+                
  |     +-- Alasmidontina
--+                      
  |  +----- Cristaria **    
  +--+                   
     |  +-- Anemina **      
     +--+                
        +-- Sinanodonta **  

Bolotov et al., 2018 [Publ 3826], fig. 2

iTtTbiriSoTroTroSiD
4423796229
        +-- Anodontina   
  +-----+                
  |     +-- Alasmidontina
--+                      
  |  +----- Cristaria **    
  +--+                   
     |  +-- Pletholophus ** 
     +--+                
        +-- Sinanodonta **  

Bulakhova et al., 2023 [Publ 4325], fig. 5

iTtTbiriSoTroTroSiD
442100031
1

Dai et al., 2025 [Publ 4440], fig. 3

iTtTbiriSoTroTroSiD
4423846229
        +-- Anodontina   
  +-----+                
  |     +-- Alasmidontina
--+                      
  |  +----- Cristaria **    
  +--+                   
     |  +-- Anemina **      
     +--+                
        +-- Sinanodonta **  

Wu et al., 2022 [Publ 4238], fig. 2d

iTtTbiriSoTroTroSiD
4423356229
        +-- Anodontina   
  +-----+                
  |     +-- Alasmidontina
--+                      
  |  +----- Cristaria **    
  +--+                   
     |  +-- Anemina **      
     +--+                
        +-- Sinanodonta **  

Wu et al., 2021 [Publ 4366], fig. 6

iTtTbiriSoTroTroSiD
4423876231
           +----- Anemina **         
  +--------+                      
  |        |  +-- Cristaria **       
  |        +--+                   
--+           +-- Sinanodonta **     
  |                               
  |  +----------- Anodonta_cygnea 
  |  |                            
  +--+        +-- Lasmigona       
     |  +-----+                   
     |  |     +-- Anodonta_anatina
     +--+                         
        |  +----- Pseudanodonta   
        +--+                      
           |  +-- Pyganodon       
           +--+                   
              +-- Strophitus      

Lopes-Lima et al., 2017 [Publ 3792], fig. 2

iTtTbiriSoTroTroSiD
4423285229
        +-- Anodontina   
  +-----+                
  |     +-- Alasmidontina
--+                      
  |  +----- Cristaria **    
  +--+                   
     |  +-- Anemina **      
     +--+                
        +-- Sinanodonta **  

Klishko et al., 2014 [Publ 3666], fig. 8

iTtTbiriSoTroTroSiD
442300029
  +----- Cristaria **  
--+                 
  |  +-- Beringiana ** 
  +--+              
     +-- Sinanodonta **

Huang et al., 2018 [Publ 3828], fig. 6

iTtTbiriSoTroTroSiD
4423385229
        +-- Anodontina   
  +-----+                
  |     +-- Alasmidontina
--+                      
  |  +----- Cristaria **    
  +--+                   
     |  +-- Anemina **      
     +--+                
        +-- Sinanodonta **  

Wu et al., 2020 [Publ 4121], fig. 3

iTtTbiriSoTroTroSiD
4423455229
        +-- Anodontina   
  +-----+                
  |     +-- Alasmidontina
--+                      
  |  +----- Cristaria **    
  +--+                   
     |  +-- Anemina **      
     +--+                
        +-- Sinanodonta **  

Froufe et al., 2019 [Publ 3936], fig. 3

iTtTbiriSoTroTroSiD
4423405229
        +-- Anodontina   
  +-----+                
  |     +-- Alasmidontina
--+                      
  |  +----- Cristaria **    
  +--+                   
     |  +-- Anemina **      
     +--+                
        +-- Sinanodonta **  

Wu et al., 2021 [Publ 4366], fig. 4

iTtTbiriSoTroTroSiD
4423658229
     +----- Cristaria **       
  +--+                      
  |  |  +-- Anemina **         
  |  +--+                   
--+     +-- Sinanodonta **     
  |                         
  |  +----- Anodonta_cygnea 
  +--+                      
     |  +-- Anodonta_anatina
     +--+                   
        +-- Alasmidontina   

Wang et al., 2018 [Publ 3865], fig. 2

iTtTbiriSoTroTroSiD
4423273229
        +-- Anodontina   
  +-----+                
  |     +-- Alasmidontina
--+                      
  |  +----- Cristaria **    
  +--+                   
     |  +-- Anemina **      
     +--+                
        +-- Sinanodonta **  

Wu et al., 2016 [Publ 3711], fig. 1

iTtTbiriSoTroTroSiD
3753184236
  +----------- Sinanodonta **  
--+                         
  |  +-------- Anemina **      
  +--+                      
     |  +----- Cristaria **    
     +--+                   
        |  +-- Anodontina   
        +--+                
           +-- Alasmidontina

Dai et al., 2024 [Publ 4358], fig. 4

iTtTbiriSoTroTroSiD
3313504230
        +-- Anodontina   
  +-----+                
  |     +-- Alasmidontina
--+                      
  |  +----- Cristaria **    
  +--+                   
     |  +-- Anemina **      
     +--+                
        +-- Sinanodonta **  

Dai et al., 2025 [Publ 4441], fig. 1

iTtTbiriSoTroTroSiD
3313450032
  +----- Sinanodonta **
--+                 
  |  +-- Anemina **    
  +--+              
     +-- Cristaria **  

Wu et al., 2020 [Publ 4147], fig. 3

iTtTbiriSoTroTroSiD
3313274230
        +-- Anodontina   
  +-----+                
  |     +-- Alasmidontina
--+                      
  |  +----- Cristaria **    
  +--+                   
     |  +-- Sinanodonta **  
     +--+                
        +-- Anemina **      

Wu et al., 2024 [Publ 4379], fig. 4_6

iTtTbiriSoTroTroSiD
3312190030
  +-- Cristaria **  
--+              
  +-- Sinanodonta **

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

iTtTbiriSoTroTroSiD
3313342230
        +-- Anodontina   
  +-----+                
  |     +-- Alasmidontina
--+                      
  |  +----- Cristaria **    
  +--+                   
     |  +-- Sinanodonta **  
     +--+                
        +-- Anemina **      

Feng et al., 2016 [Publ 3800], fig. 2

iTtTbiriSoTroTroSiD
3312113230
  +----- Alasmidontina
--+                   
  |  +-- Anemina **      
  +--+                
     +-- Sinanodonta **  

Gomes-dos-Santos et al., 2019 [Publ 4042], fig. 8

iTtTbiriSoTroTroSiD
331333130
  +-------- Anodontina 
--+                    
  |  +----- Cristaria **  
  +--+                 
     |  +-- Anemina **    
     +--+              
        +-- Sinanodonta **

Chen et al., 2020 [Publ 4170], fig. 1

iTtTbiriSoTroTroSiD
334323318263
                       +----- Prolasmidonta           
                    +--+                              
                    |  |  +-- Alasmidonta             
                    |  +--+                           
  +-----------------+     +-- Anemina **                 
  |                 |                                 
  |                 |  +----- Pseudodontina           
  |                 +--+                              
  |                    |  +-- Lamprotula_caveata      
  |                    +--+                           
  |                       +-- Chamberlainia           
  |                                                   
  |                    +----- Schistodesmus           
  |              +-----+                              
  |              |     |  +-- Lepidodesma             
--+              |     +--+                           
  |              |        +-- Aculamprotula           
  |  +-----------+                                    
  |  |           |     +----- Platynaias              
  |  |           |  +--+                              
  |  |           |  |  |  +-- Utterbackia_imbecillis  
  |  |           |  |  +--+                           
  |  |           +--+     +-- Pyganodon               
  |  |              |                                 
  |  |              |  +----- Sinanodonta **             
  |  |              +--+                              
  |  |                 |  +-- Cristaria **               
  +--+                 +--+                           
     |                    +-- Anodontina              
     |                                                
     |                    +-- Utterbackia_peninsularis
     |           +--------+                           
     |           |        +-- Potamilus_alatus        
     |  +--------+                                    
     |  |        |  +-------- Lamprotula_leaii        
     |  |        +--+                                 
     |  |           |  +----- Sinohyriopsis           
     |  |           +--+                              
     +--+              |  +-- Parvasolenaia           
        |              +--+                           
        |                 +-- Sinosolenaia            
        |                                             
        |  +----------------- Gibbosula               
        +--+                                          
           |  +-------------- Quadrulina              
           +--+                                       
              |  +----------- Toxolasma               
              +--+                                    
                 |  +-------- Lampsilis_ornata        
                 +--+                                 
                    |  +----- Potamilus_leptodon      
                    +--+                              
                       |  +-- Lampsilis_siliquoidea   
                       +--+                           
                          +-- Lampsilis_powellii      

Ouyang et al., 2011 [Publ 3408], fig. 1

iTtTbiriSoTroTroSiD
3312100030
  +-- Anemina **    
--+              
  +-- Sinanodonta **

Kano et al., 2019 [Publ 4194], fig. 6

iTtTbiriSoTroTroSiD
331370032
  +----- Sinanodonta ** 
--+                  
  |  +-- Pletholophus **
  +--+               
     +-- Beringiana **  

Wu et al., 2018 [Publ 4143], fig. 2

iTtTbiriSoTroTroSiD
3753194236
  +----------- Sinanodonta **  
--+                         
  |  +-------- Anemina **      
  +--+                      
     |  +----- Cristaria **    
     +--+                   
        |  +-- Anodontina   
        +--+                
           +-- Alasmidontina

Dai et al., 2023 [Publ 4285], fig. 5

iTtTbiriSoTroTroSiD
3313504230
        +-- Anodontina   
  +-----+                
  |     +-- Alasmidontina
--+                      
  |  +----- Cristaria **    
  +--+                   
     |  +-- Anemina **      
     +--+                
        +-- Sinanodonta **  

Dai et al., 2024 [Publ 4389], fig. 3

iTtTbiriSoTroTroSiD
3313440032
  +----- Sinanodonta **
--+                 
  |  +-- Anemina **    
  +--+              
     +-- Cristaria **  

Guerra et al., 2017 [Publ 3772], fig. 2a

iTtTbiriSoTroTroSiD
2802236237
  +-------- Sinanodonta **  
--+                      
  |  +----- Cristaria **    
  +--+                   
     |  +-- Anodontina   
     +--+                
        +-- Alasmidontina

Zhang et al., 2004 [Publ 2735], fig. 1

iTtTbiriSoTroTroSiD
22021200--
  +-- Sinanodonta **
--+              
  +-- Cristaria **  

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iTtTbiriSoTroTroSiD
22025522--
     +-- Cristaria **    
  +--+                
  |  +-- Sinanodonta **  
--+                   
  |  +-- Anodontina   
  +--+                
     +-- Alasmidontina

Huang et al., 2002 [Publ 2657], fig. 1

iTtTbiriSoTroTroSiD
22023251--
  +----- Alasmidontina
--+                   
  |  +-- Sinanodonta **  
  +--+                
     +-- Cristaria **    

Soroka, 2010 [Publ 3404], fig. 3a-M

iTtTbiriSoTroTroSiD
2201411--
1

Zhou et al., 2016 [Publ 3710], fig. 1

iTtTbiriSoTroTroSiD
2602184135
  +----- Sinanodonta **  
--+                   
  |  +-- Cristaria **    
  +--+                
     +-- Alasmidontina

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iTtTbiriSoTroTroSiD
22025522--
     +-- Cristaria **    
  +--+                
  |  +-- Sinanodonta **  
--+                   
  |  +-- Anodontina   
  +--+                
     +-- Alasmidontina

Guerra et al., 2017 [Publ 3772], fig. 2b

iTtTbiriSoTroTroSiD
22022362--
     +-- Cristaria **    
  +--+                
  |  +-- Sinanodonta **  
--+                   
  |  +-- Anodontina   
  +--+                
     +-- Alasmidontina

Zhang et al., 2013 [Publ 4214], fig. 5

iTtTbiriSoTroTroSiD
240220033
  +-------- Cristaria **    
--+                      
  |  +----- Sinanodonta **  
  +--+                   
     |  +-- Margaritifera
     +--+                
        +-- Lamprotula   

Soroka, 2010 [Publ 3404], fig. 5a

iTtTbiriSoTroTroSiD
2201631--
  +-------- Anodonta_anatina
--+                         
  |  +----- Sinanodonta **     
  +--+                      
     |  +-- Anodonta_cygnea 
     +--+                   
        +-- Pseudanodonta   

Zhou et al., 2007 [Publ 2658], fig. 2

iTtTbiriSoTroTroSiD
21802160047
  +----------------- Sinanodonta **  
  |                               
--+  +-------------- Cristaria **    
  |  |                            
  +--+           +-- Lamprotula   
     |  +--------+                
     |  |        +-- Quadrulina   
     +--+                         
        |  +-------- Aculamprotula
        +--+                      
           |  +----- Acuticosta   
           +--+                   
              |  +-- Nodularia    
              +--+                
                 +-- Cuneopsis    

Soroka, 2010 [Publ 3404], fig. 5b

iTtTbiriSoTroTroSiD
2201631--
1

Riccardi et al., 2020 [Publ 4089], fig. 1

iTtTbiriSoTroTroSiD
2202441--
  +----- Anodontina 
--+                 
  |  +-- Sinanodonta **
  +--+              
     +-- Anemina **    

Soroka, 2010 [Publ 3404], fig. 3b-M

iTtTbiriSoTroTroSiD
2201411--
1

Tomilova et al., 2020 [Publ 4097], fig. 3

iTtTbiriSoTroTroSiD
2201641--
1

Wen et al., 2017 [Publ 3795], fig. 3

iTtTbiriSoTroTroSiD
22021411--
  +----- Anodontina 
--+                 
  |  +-- Sinanodonta **
  +--+              
     +-- Cristaria **  

Xia et al., 2016 [Publ 3712], fig. 7B

iTtTbiriSoTroTroSiD
230210032
  +----- Sinanodonta **  
--+                   
  |  +-- Cristaria **    
  +--+                
     +-- Sinohyriopsis

Huang et al., 2013 [Publ 3629], fig. 3

iTtTbiriSoTroTroSiD
22021441--
  +----- Alasmidontina
--+                   
  |  +-- Sinanodonta **  
  +--+                
     +-- Cristaria **    

Soroka, 2010 [Publ 3404], fig. 3b

iTtTbiriSoTroTroSiD
2201731--
1

Zhou et al., 2007 [Publ 2658], fig. 1

iTtTbiriSoTroTroSiD
22021900--
  +-- Sinanodonta **
--+              
  +-- Cristaria **  

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

iTtTbiriSoTroTroSiD
22022931--
  +----- Alasmidontina
--+                   
  |  +-- Cristaria **    
  +--+                
     +-- Sinanodonta **  

Huang et al., 2002 [Publ 2657], fig. 2

iTtTbiriSoTroTroSiD
22023251--
  +-- Sinanodonta **
--+              
  +-- Cristaria **  

Chong et al., 2008 [Publ 2668], fig. 3

iTtTbiriSoTroTroSiD
220216152--
  +-- Beringiana ** 
--+              
  +-- Sinanodonta **

Pfarrer et al., 2022 [Publ 4305], fig. 2-5

iTtTbiriSoTroTroSiD
22021031--
  +----- Anodontina 
--+                 
  |  +-- Cristaria **  
  +--+              
     +-- Sinanodonta **

Soroka, 2010 [Publ 3404], fig. 3a

iTtTbiriSoTroTroSiD
2201731--
1

Dai et al., 2024 [Publ 4386], fig. 5-BI

iTtTbiriSoTroTroSiD
11012011--
-- Anemina_arcaeformis **

Mezhzherin et al., 2014 [Publ 3705], fig. 4

iTtTbiriSoTroTroSiD
1101331--
  +-- Sinanodonta **
--+              
  +-- Anodontina 

Mock et al., 2004 [Publ 2686], fig. 4

iTtTbiriSoTroTroSiD
1101111--
  +-- Anodontina 
--+              
  +-- Sinanodonta **

Wu et al., 2022 [Publ 4238], fig. 2e

iTtTbiriSoTroTroSiD
11011732--
  +----- Sinanodonta **  
--+                   
  |  +-- Anodontina   
  +--+                
     +-- Alasmidontina

Smith et al., 2018 [Publ 3790], fig. 1

iTtTbiriSoTroTroSiD
110117171--
  +-- Cristaria **    
--+                
  +-- Alasmidontina

Froufe et al., 2017 [Publ 3802], fig. 7

iTtTbiriSoTroTroSiD
1101931--
  +-- Sinanodonta **
--+              
  +-- Anodontina 

Burzynski & Soroka, 2018 [Publ 3862], fig. 3

iTtTbiriSoTroTroSiD
1101332--
  +----- Sinanodonta **  
--+                   
  |  +-- Anodontina   
  +--+                
     +-- Alasmidontina

Soroka, 2010 [Publ 3404], fig. 4-M

iTtTbiriSoTroTroSiD
1101111--
  +-- Anodontina 
--+              
  +-- Sinanodonta **

Soroka, 2010 [Publ 3404], fig. 8b

iTtTbiriSoTroTroSiD
1101521--
-- Sinanodonta_woodiana **

Amaro et al., 2017 [Publ 3791], fig. 2-M

iTtTbiriSoTroTroSiD
1101931--
  +----- Pyganodon  
  |                 
--+  +-- Lasmigona  
  |  |              
  +--+-- Sinanodonta **
     |              
     +-- Utterbackia

Mock et al., 2004 [Publ 2686], fig. 3

iTtTbiriSoTroTroSiD
1101111--
  +-- Anodontina 
--+              
  +-- Sinanodonta **

Soroka, 2010 [Publ 3404], fig. 8a-M

iTtTbiriSoTroTroSiD
1101200--
-- Sinanodonta_woodiana **

Froufe et al., 2019 [Publ 3936], fig. 3-M

iTtTbiriSoTroTroSiD
11011832--
  +----- Sinanodonta **  
--+                   
  |  +-- Anodontina   
  +--+                
     +-- Alasmidontina

Soroka, 2010 [Publ 3404], fig. 5a-M

iTtTbiriSoTroTroSiD
1101111--
  +-- Anodontina 
--+              
  +-- Sinanodonta **

Soroka, 2010 [Publ 3404], fig. 5b-M

iTtTbiriSoTroTroSiD
1101111--
  +-- Anodontina 
--+              
  +-- Sinanodonta **

Dai et al., 2024 [Publ 4386], fig. 5-ML

iTtTbiriSoTroTroSiD
11012011--
-- Anemina_arcaeformis **

Sheng et al., 2014 [Publ 3704], fig. 2

iTtTbiriSoTroTroSiD
1101711--
-- Cristaria_plicata **

Plazzi & Passamonti, 2010 [Publ 3354], fig. 2-3

iTtTbiriSoTroTroSiD
1101300--
-- Sinanodonta_woodiana **

Soroka, 2010 [Publ 3404], fig. 2

iTtTbiriSoTroTroSiD
1101631--
-- Sinanodonta_woodiana **

Plazzi et al., 2011 [Publ 3676], fig. 5

iTtTbiriSoTroTroSiD
11011742--
-- Cristaria_plicata **

Froufe et al., 2017 [Publ 3802], fig. 2

iTtTbiriSoTroTroSiD
1101331--
  +-- Cristaria ** 
--+             
  +-- Anodontina

Gustafson & Iwamoto, 2005 [Publ 2741], fig. 2

iTtTbiriSoTroTroSiD
1101421--
  +-- Beringiana **
--+             
  +-- Anodontina

Inoue et al., 2018 [Publ 3858], fig. 1_3a-c

iTtTbiriSoTroTroSiD
1101103192--
  +-- Alasmidonta_marginata      
  |                              
  +-- Prolasmidonta              
  |                              
  +-- Alasmidonta_undulata       
  |                              
  +-- Beringiana **                 
  |                              
  +-- Anodonta_kennerlyi         
  |                              
  +-- Anodonta_nuttalliana       
  |                              
  +-- Anodonta_oregonensis       
  |                              
  +-- Arcidens_confragosus       
  |                              
  +-- Arcidens_wheeleri          
  |                              
  +-- Pterosyna                  
  |                              
--+-- Platynaias_compressa       
  |                              
  +-- Lasmigona                  
  |                              
  +-- Platynaias_decorata        
  |                              
  +-- Platynaias_subviridis      
  |                              
  +-- Pyganodon_grandis          
  |                              
  +-- Pyganodon_lacustris        
  |                              
  +-- Pseudodontoideus           
  |                              
  +-- Strophitus                 
  |                              
  +-- Utterbackia_imbecillis     
  |                              
  +-- Utterbackia_peninsularis__1
  |                              
  +-- Utterbackia_peninsularis__2

Nagel et al., 1996 [Publ 3400], fig. 351

iTtTbiriSoTroTroSiD
1101441--
  +-------------- Pseudanodonta   
--+                               
  |  +----------- Anodonta_cygnea 
  +--+                            
     |  +-------- Sinanodonta **     
     +--+                         
        |  +----- Anodonta__2     
        +--+                      
           |  +-- Anodonta_anatina
           +--+                   
              +-- Anodonta__1     

Huang et al., 2019 [Publ 3843], fig. 3-M

iTtTbiriSoTroTroSiD
11012232--
  +----- Sinanodonta **  
--+                   
  |  +-- Anodontina   
  +--+                
     +-- Alasmidontina

Wei et al., 2016 [Publ 3665], fig. 0

iTtTbiriSoTroTroSiD
1101511--
-- Cristaria_plicata **

Huang et al., 2018 [Publ 3828], fig. 6-M

iTtTbiriSoTroTroSiD
11011632--
  +----- Sinanodonta **  
--+                   
  |  +-- Anodontina   
  +--+                
     +-- Alasmidontina

Guerra et al., 2019 [Publ 4168], fig. 2-M

iTtTbiriSoTroTroSiD
11012132--
  +----- Alasmidontina
--+                   
  |  +-- Anodontina   
  +--+                
     +-- Sinanodonta **  

Soroka, 2008 [Publ 3695], fig. 1-M

iTtTbiriSoTroTroSiD
1101322--
  +----- Sinanodonta **  
--+                   
  |  +-- Anodontina   
  +--+                
     +-- Alasmidontina

Froufe et al., 2019 [Publ 3936], fig. 4

iTtTbiriSoTroTroSiD
11011832--
  +----- Sinanodonta **  
--+                   
  |  +-- Anodontina   
  +--+                
     +-- Alasmidontina

Liu et al., 2017 [Publ 3939], fig. 2

iTtTbiriSoTroTroSiD
11011400--
-- Anemina_arcaeformis **

Soroka, 2010 [Publ 3404], fig. 2-M

iTtTbiriSoTroTroSiD
1101411--
-- Sinanodonta_woodiana **

Burzynski & Soroka, 2018 [Publ 3862], fig. 5b

iTtTbiriSoTroTroSiD
1101932--
  +----- Sinanodonta **  
--+                   
  |  +-- Anodontina   
  +--+                
     +-- Alasmidontina

Guerra et al., 2019 [Publ 4168], fig. 2

iTtTbiriSoTroTroSiD
11014062--
  +----- Sinanodonta **  
--+                   
  |  +-- Anodontina   
  +--+                
     +-- Alasmidontina

Jiang et al., 2010 [Publ 3438], fig. 5

iTtTbiriSoTroTroSiD
1101300--
-- Cristaria_plicata **

Mezhzherin et al., 2014 [Publ 3705], fig. 2

iTtTbiriSoTroTroSiD
1101331--
  +-- Sinanodonta **
--+              
  +-- Anodontina 

Soroka, 2008 [Publ 3695], fig. 1

iTtTbiriSoTroTroSiD
1101322--
  +----- Sinanodonta **  
--+                   
  |  +-- Anodontina   
  +--+                
     +-- Alasmidontina

Soroka, 2010 [Publ 3404], fig. 8b-M

iTtTbiriSoTroTroSiD
1101200--
-- Sinanodonta_woodiana **

Smith, 2021 [Publ 4331], fig. 1

iTtTbiriSoTroTroSiD
11011921--
  +-- Cristaria **    
--+                
  +-- Alasmidontina

Nagel et al., 1998 [Publ 1946], fig. 7

iTtTbiriSoTroTroSiD
1101441--
  +-------------- Pseudanodonta   
--+                               
  |  +----------- Anodonta_cygnea 
  +--+                            
     |  +-------- Sinanodonta **     
     +--+                         
        |  +----- Anodonta__2     
        +--+                      
           |  +-- Anodonta_anatina
           +--+                   
              +-- Anodonta__1     

Soroka, 2010 [Publ 3404], fig. 8a

iTtTbiriSoTroTroSiD
1101521--
  +-- Sinanodonta **
--+              
  +-- Anodontina 

Zieritz et al., 2021 [Publ 4231], fig. 3-M_4

iTtTbiriSoTroTroSiD
11012232--
  +----- Sinanodonta **  
--+                   
  |  +-- Anodontina   
  +--+                
     +-- Alasmidontina

Soroka, 2010 [Publ 3404], fig. 4

iTtTbiriSoTroTroSiD
1101631--
  +-- Sinanodonta **
--+              
  +-- Anodontina 

Breton et al., 2011 [Publ 3376], fig. 3

iTtTbiriSoTroTroSiD
110136111--
  +-- Cristaria **    
--+                
  +-- Alasmidontina

Burzynski & Soroka, 2018 [Publ 3862], fig. 5a

iTtTbiriSoTroTroSiD
11011932--
  +-- Sinanodonta **  
  |                
--+-- Anodontina   
  |                
  +-- Alasmidontina

Peng et al., 2012 [Publ 3689], fig. 1

iTtTbiriSoTroTroSiD
1101200--
-- Cristaria_plicata **

Guerra et al., 2019 [Publ 4168], fig. 1-M

iTtTbiriSoTroTroSiD
11012132--
  +----- Sinanodonta **  
--+                   
  |  +-- Anodontina   
  +--+                
     +-- Alasmidontina

Chase et al., 2018 [Publ 3847], fig. 4

iTtTbiriSoTroTroSiD
110137132--
  +----- Sinanodonta **  
--+                   
  |  +-- Anodontina   
  +--+                
     +-- Alasmidontina

Zieritz et al., 2012 [Publ 3673], fig. 4

iTtTbiriSoTroTroSiD
1101731--
-- Sinanodonta_woodiana **

Guerra et al., 2019 [Publ 4168], fig. 1

iTtTbiriSoTroTroSiD
11014062--
  +----- Sinanodonta **  
--+                   
  |  +-- Anodontina   
  +--+                
     +-- Alasmidontina

Zhou et al., 2017 [Publ 3788], fig. 2

iTtTbiriSoTroTroSiD
11011500--
-- Anemina_arcaeformis **

Chen et al., 2012 [Publ 3624], fig. 4

iTtTbiriSoTroTroSiD
1101611--
  +-- Alasmidontina
--+                
  +-- Cristaria **    

Dai et al., 2024 [Publ 4386], fig. 6

iTtTbiriSoTroTroSiD
11014021--
  +-- Sinanodonta **  
--+                
  +-- Alasmidontina
 
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