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Page last updated
Thu 3 September 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

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Bivalvia | Unionida | Hyrioidea | Hyriidae | Hyriinae

tribe HYRIDELLINI McMichael, 1956 (1934): cladomics
     4 genera

Cladomics Summary

number of species = 9 (13 total)
number of subtaxa (rank: genus) = 4
analyzed in 46 cladograms in 28 publications
maximum number of terminals in an analysis = 4 (mean = 1.59, median = 1)
maximum number of subtaxa (genus) = 2 (mean = 1.11, median = 1)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 16
maximum number of included terminals = 4, (mean = 2.69 median = 2)
maximum number of subtaxa (genus) = 2 (mean = 1.31, median = 1)
proportion of cladograms that support basic monophyly = 0.56

Rigorous Monophyly

number of cladograms that test rigorous monophyly = 5
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (genus) taxon and at least one rigorous outgroup (family).
maximum number of included terminals = 4 (mean = 3.6, median = 4)
maximum number of subtaxa (genus) = 2 (mean = 2, median = 2)
proportion of cladograms that support rigorous monophyly = 0.4

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.


PCoA Ordination


NJ Tree

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

iTtTbiriSoTroTroSiD
4182026281440
                             +----- Diplodon_suavidicus__2
  +--------------------------+                            
  |                          |  +-- Hyriini               
  |                          +--+                         
  |                             +-- Castaliini            
  |                                                       
  |                             +-- Hyridella_depressa **    
--+                       +-----+                         
  |                       |     +-- Hyridella_drapeta **     
  |  +--------------------+                               
  |  |                    |  +----- Diplodon_suavidicus__1
  |  |                    +--+                            
  |  |                       |  +-- Diplodon_chilensis    
  |  |                       +--+                         
  +--+                          +-- Diplodon_gratus       
     |                                                    
     |                          +-- Cucumerunio **           
     |  +-----------------------+                         
     |  |                       +-- Hyridella_australis **   
     |  |                                                 
     +--+  +----------------------- Echyridella           
        |  |                                              
        |  |              +-------- Velesunio__1          
        +--+  +-----------+                               
           |  |           |  +----- Alathyria_jacksoni__1 
           |  |           +--+                            
           |  |              |  +-- Velesunio_ambiguus    
           +--+              +--+                         
              |                 +-- Alathyria_profuga     
              |                                           
              |  +----------------- Velesunio__3          
              |  |                                        
              +--+  +-------------- Microdontia           
                 |  |                                     
                 +--+           +-- Lortiella             
                    |  +--------+                         
                    |  |        +-- Velesunio_angasi      
                    +--+                                  
                       |  +-------- Alathyria_pertexta__2 
                       +--+                               
                          |  +----- Velesunio__2          
                          +--+                            
                             |  +-- Alathyria_jacksoni__2 
                             +--+                         
                                +-- Alathyria_pertexta__1

Santos-Neto et al., 2016 [Publ 3739], fig. 4

iTtTbiriSoTroTroSiD
413112169417.5
                    +----- Diplodon_suavidicus__2
  +-----------------+                            
  |                 |  +-- Hyriini               
  |                 +--+                         
  |                    +-- Castaliini            
  |                                              
  |              +-------- Westralunio           
--+              |                               
  |  +-----------+     +-- Hyridella_drapeta **     
  |  |           |  +--+                         
  |  |           |  |  +-- Hyridella_depressa **    
  |  |           +--+                            
  |  |              |  +-- Diplodon_suavidicus__1
  +--+              +--+                         
     |                 +-- Diplodon_gratus       
     |                                           
     |  +----------------- Echyridella           
     |  |                                        
     +--+              +-- Cucumerunio **           
        |  +-----------+                         
        |  |           +-- Hyridella_australis **   
        +--+                                     
           |  +----------- Velesunio_ambiguus    
           +--+                                  
              |  +-------- Alathyria             
              +--+                               
                 |  +----- Velesunio_angasi      
                 +--+                            
                    |  +-- Lortiella             
                    +--+                         
                       +-- Microdontia

Graf et al., 2015 [Publ 3619], fig. 1

iTtTbiriSoTroTroSiD
4422163318
     +----- Rhipidodontini     
  +--+                         
  |  |  +-- Hyriini            
  |  +--+                      
  |     +-- Castaliini         
--+                            
  |     +-- Hyridella_depressa ** 
  |  +--+                      
  |  |  +-- Hyridella_drapeta **  
  +--+                         
     |  +-- Hyridella_australis **
     +--+                      
        +-- Cucumerunio **

Santos-Neto et al., 2016 [Publ 3739], fig. 2a_b

iTtTbiriSoTroTroSiD
419202169417.5
                 +----- Diplodon_suavidicus__1
  +--------------+                            
  |              |  +-- Hyriini               
  |              +--+                         
  |                 +-- Castaliini            
  |                                           
  |           +-------- Westralunio           
  |           |                               
  +-----------+     +-- Hyridella_drapeta **     
  |           |  +--+                         
--+           |  |  +-- Hyridella_depressa **    
  |           +--+                            
  |              |  +-- Diplodon_gratus       
  |              +--+                         
  |                 +-- Diplodon_suavidicus__2
  |                                           
  |                 +-- Hyridella_australis **   
  |  +--------------+                         
  |  |              +-- Cucumerunio **           
  +--+                                        
     |  +-------------- Echyridella           
     +--+                                     
        |  +----------- Velesunio_ambiguus    
        +--+                                  
           |  +-------- Microdontia           
           +--+                               
              |  +----- Lortiella             
              +--+                            
                 |  +-- Alathyria             
                 +--+                         
                    +-- Velesunio_angasi

Huang et al., 2019 [Publ 3843], fig. S4

iTtTbiriSoTroTroSiD
33111974419
1

Huang et al., 2019 [Publ 3843], fig. 2

iTtTbiriSoTroTroSiD
33111974419
  +----------- Echyridella   
--+                          
  |  +-------- Hyridella **     
  +--+                       
     |  +----- Rhipidodontini
     +--+                    
        |  +-- Castaliini    
        +--+                 
           +-- Hyriini

Huang et al., 2019 [Publ 3843], fig. S3

iTtTbiriSoTroTroSiD
33111954419
  +----------- Echyridella   
--+                          
  |  +-------- Hyridella **     
  +--+                       
     |  +----- Rhipidodontini
     +--+                    
        |  +-- Castaliini    
        +--+                 
           +-- Hyriini

Marshall et al., 2014 [Publ 3593], fig. 1

iTtTbiriSoTroTroSiD
22021252--
  +----- Rhipidodontini
--+                    
  |  +-- Cucumerunio **   
  +--+                 
     +-- Hyridella **

Graf & Ó Foighil, 2000 [Publ 2649], fig. 1_2

iTtTbiriSoTroTroSiD
230193322
           +-- Rhipidodontini        
  +--------+                         
  |        +-- Castaliini            
--+                                  
  |  +-------- Hyridella_australis__2 **
  +--+                               
     |  +----- Hyridella_australis__1 **
     +--+                            
        |  +-- Echyridella           
        +--+                         
           +-- Hyridella_depressa **

Hoeh et al., 2009 [Publ 2716], fig. 3

iTtTbiriSoTroTroSiD
22014543--
  +-------- Echyridella   
--+                       
  |  +----- Hyridella **     
  +--+                    
     |  +-- Castaliini    
     +--+                 
        +-- Rhipidodontini

Walker et al., 2006 [Publ 2666], fig. 6

iTtTbiriSoTroTroSiD
2801343314
  +----------------- Castaliini         
  |                                     
  +----------------- Rhipidodontini     
--+                                     
  |  +-------------- Hyridella_depressa ** 
  |  |                                  
  |  +-------------- Hyridella_australis **
  +--+                                  
     |  +----------- Echyridella        
     |  |                               
     |  |        +-- Velesunio_ambiguus 
     +--+  +-----+                      
        |  |     +-- Velesunio__3       
        |  |                            
        +--+  +----- Velesunio__2       
           |  |                         
           |  |  +-- Alathyria          
           +--+--+                      
              |  +-- Velesunio__1       
              |                         
              |  +-- Velesunio_angasi   
              +--+                      
                 +-- Lortiella

Baker et al., 2003 [Publ 2690], fig. 5

iTtTbiriSoTroTroSiD
270163321
           +-- Hyridella_australis **
  +--------+                      
  |        +-- Rhipidodontini     
--+                               
  |  +-------- Hyridella_depressa ** 
  +--+                            
     |  +----- Velesunio          
     +--+                         
        |  +-- Echyridella        
        +--+                      
           +-- Castaliini

Graf & Cummings, 2006 [Publ 1968], fig. 2

iTtTbiriSoTroTroSiD
22014543--
  +-------- Echyridella   
--+                       
  |  +----- Hyridella **     
  +--+                    
     |  +-- Castaliini    
     +--+                 
        +-- Rhipidodontini

Hoeh et al., 2009 [Publ 2716], fig. 5

iTtTbiriSoTroTroSiD
22014443--
  +-------- Echyridella   
--+                       
  |  +----- Hyridella **     
  +--+                    
     |  +-- Castaliini    
     +--+                 
        +-- Rhipidodontini

Baker et al., 2004 [Publ 2685], fig. 2-3

iTtTbiriSoTroTroSiD
230173321
  +----- Hyridella_depressa ** 
--+                         
  |  +-- Hyridella_australis **
  +--+                      
     +-- Rhipidodontini

Hoeh et al., 2009 [Publ 2716], fig. 4

iTtTbiriSoTroTroSiD
22014443--
  +-------- Echyridella   
--+                       
  |  +----- Hyridella **     
  +--+                    
     |  +-- Castaliini    
     +--+                 
        +-- Rhipidodontini

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

iTtTbiriSoTroTroSiD
11013922--
  +----- Hyridella ** 
--+                
  |  +-- Hyriini   
  +--+             
     +-- Castaliini

Sonowal et al., 2021 [Publ 4221], fig. 4

iTtTbiriSoTroTroSiD
11011511--
-- Hyridella_australis **

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iTtTbiriSoTroTroSiD
11017900--
-- Hyridella_depressa **

Combosch et al., 2017 [Publ 3806], fig. 1

iTtTbiriSoTroTroSiD
11011411--
-- Hyridella_australis **

Walker et al., 2006 [Publ 2666], fig. 3

iTtTbiriSoTroTroSiD
11011400--
-- Hyridella **

Klunzinger et al., 2021 [Publ 4086], fig. 2

iTtTbiriSoTroTroSiD
1101500--
-- Cucumerunio_novaehollandiae **

Graf, 2002 [Publ 2651], fig. 2

iTtTbiriSoTroTroSiD
11011722--
  +----- Hyridella **     
--+                    
  |  +-- Rhipidodontini
  +--+                 
     +-- Castaliini

Whelan et al., 2011 [Publ 3380], fig. 1-ML

iTtTbiriSoTroTroSiD
11013522--
  +----- Hyridella ** 
--+                
  |  +-- Castaliini
  +--+             
     +-- Hyriini

Sonowal et al., 2021 [Publ 4221], fig. 3a

iTtTbiriSoTroTroSiD
11011411--
-- Hyridella_australis **

Sano et al., 2020 [Publ 4138], fig. 3

iTtTbiriSoTroTroSiD
11016711--
-- Hyridella_australis **

Klunzinger et al., 2022 [Publ 4259], fig. 1

iTtTbiriSoTroTroSiD
1101300--
-- Cucumerunio_novaehollandiae **

Chen et al., 2025 [Publ 4455], fig. 1

iTtTbiriSoTroTroSiD
11012511--
  +-- Hyridella **
--+            
  +-- Hyriini

Wu et al., 2020 [Publ 4121], fig. 2

iTtTbiriSoTroTroSiD
11013311--
-- Hyridella_australis **

Whelan et al., 2011 [Publ 3380], fig. 1-MP

iTtTbiriSoTroTroSiD
11013522--
  +----- Hyridella ** 
--+                
  |  +-- Castaliini
  +--+             
     +-- Hyriini

Sonowal et al., 2021 [Publ 4221], fig. 3b

iTtTbiriSoTroTroSiD
11011411--
-- Hyridella_australis **

Combosch et al., 2017 [Publ 3806], fig. 2

iTtTbiriSoTroTroSiD
11011411--
  +-- Hyriini  
--+            
  +-- Hyridella **

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B

iTtTbiriSoTroTroSiD
11012900--
-- Hyridella_depressa **

Walker et al., 2006 [Publ 2666], fig. 2

iTtTbiriSoTroTroSiD
11011400--
-- Hyridella **

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iTtTbiriSoTroTroSiD
11015600--
-- Hyridella_australis **

Walker et al., 2006 [Publ 2666], fig. 4

iTtTbiriSoTroTroSiD
11011400--
-- Hyridella **

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iTtTbiriSoTroTroSiD
110112100--
-- Hyridella_depressa **

Sano et al., 2020 [Publ 4138], fig. 2

iTtTbiriSoTroTroSiD
11016711--
-- Hyridella_australis **

Huang et al., 2018 [Publ 3828], fig. 3_5

iTtTbiriSoTroTroSiD
11012811--
-- Hyridella_australis **

Sano et al., 2020 [Publ 4138], fig. 4

iTtTbiriSoTroTroSiD
11016711--
-- Hyridella_australis **

Wu et al., 2020 [Publ 4121], fig. S1

iTtTbiriSoTroTroSiD
11013311--
-- Hyridella_australis **

Lopes-Lima et al., 2018 [Publ 3829], fig. 1

iTtTbiriSoTroTroSiD
11012711--
-- Hyridella_australis **

Lopes-Lima et al., 2018 [Publ 3829], fig. 3

iTtTbiriSoTroTroSiD
11012711--
-- Hyridella_australis **

Graf, 2000 [Publ 2647], fig. 1

iTtTbiriSoTroTroSiD
11011622--
-- Hyridella **

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A

iTtTbiriSoTroTroSiD
11012900--
-- Hyridella_depressa **

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iTtTbiriSoTroTroSiD
11015600--
-- Hyridella_australis **
 
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