MUSSELpdb | family Hyriidae Swainson, 1840 cladomics

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Page last updated
Thu 3 September 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

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Bivalvia | Unionida | Hyrioidea

family HYRIIDAE Swainson, 1840: cladomics
2 genera, 2 subfamilies

Cladomics Summary

number of species = 93 (132 total)
number of subtaxa (rank: subfamily) = 4
analyzed in 105 cladograms in 55 publications
maximum number of terminals in an analysis = 28 (mean = 4.07, median = 3)
maximum number of subtaxa (subfamily) = 2 (mean = 1.53, median = 2)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 61
maximum number of included terminals = 28, (mean = 6.28 median = 5)
maximum number of subtaxa (subfamily) = 2 (mean = 1.92, median = 2)
proportion of cladograms that support basic monophyly = 1

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination

NJ Tree

Campbell & Lydeard, 2026 [Publ 4810], fig. 6-14

** Top Monophyly Cladogram ** ** Top Sister-Group Cladogram **

tT	bi	riS	oT	roT	roS	iD
28	28	34	2	604	0	0	0

  +----------------------------------- Westralunio **           
  |                                                          
  |                             +----- Diplodon_suavidicus__2 **
  |  +--------------------------+                            
--+  |                          |  +-- Hyriini **               
  |  |                          +--+                         
  |  |                             +-- Castaliini **            
  |  |                                                       
  |  |                             +-- Hyridella_depressa **    
  +--+                       +-----+                         
     |                       |     +-- Hyridella_drapeta **     
     |  +--------------------+                               
     |  |                    |  +----- Diplodon_suavidicus__1 **
     |  |                    +--+                            
     |  |                       |  +-- Diplodon_chilensis **    
     |  |                       +--+                         
     +--+                          +-- Diplodon_gratus **       
        |                                                    
        |                          +-- Cucumerunio **           
        |  +-----------------------+                         
        |  |                       +-- Hyridella_australis **   
        |  |                                                 
        +--+  +----------------------- Echyridella **           
           |  |                                              
           |  |              +-------- Velesunio__1 **          
           +--+  +-----------+                               
              |  |           |  +----- Alathyria_jacksoni__1 ** 
              |  |           +--+                            
              |  |              |  +-- Velesunio_ambiguus **    
              +--+              +--+                         
                 |                 +-- Alathyria_profuga **     
                 |                                           
                 |  +----------------- Velesunio__3 **          
                 |  |                                        
                 +--+  +-------------- Microdontia **           
                    |  |                                     
                    +--+           +-- Lortiella **             
                       |  +--------+                         
                       |  |        +-- Velesunio_angasi **      
                       +--+                                  
                          |  +-------- Alathyria_pertexta__2 ** 
                          +--+                               
                             |  +----- Velesunio__2 **          
                             +--+                            
                                |  +-- Alathyria_jacksoni__2 ** 
                                +--+                         
                                   +-- Alathyria_pertexta__1 **

Santos-Neto et al., 2016 [Publ 3739], fig. 2a_b

tT	bi	riS	oT	roT	roS	iD
19	19	20	2	1	0	0	22.5

                 +----- Diplodon_suavidicus__1 **
  +--------------+                            
  |              |  +-- Hyriini **               
  |              +--+                         
  |                 +-- Castaliini **            
  |                                           
  |           +-------- Westralunio **           
  |           |                               
  +-----------+     +-- Hyridella_drapeta **     
  |           |  +--+                         
--+           |  |  +-- Hyridella_depressa **    
  |           +--+                            
  |              |  +-- Diplodon_gratus **       
  |              +--+                         
  |                 +-- Diplodon_suavidicus__2 **
  |                                           
  |                 +-- Hyridella_australis **   
  |  +--------------+                         
  |  |              +-- Cucumerunio **           
  +--+                                        
     |  +-------------- Echyridella **           
     +--+                                     
        |  +----------- Velesunio_ambiguus **    
        +--+                                  
           |  +-------- Microdontia **           
           +--+                               
              |  +----- Lortiella **             
              +--+                            
                 |  +-- Alathyria **             
                 +--+                         
                    +-- Velesunio_angasi **

Santos-Neto et al., 2016 [Publ 3739], fig. 4

tT	bi	riS	oT	roT	roS	iD
19	19	21	2	1	0	0	25.5

                    +----- Diplodon_suavidicus__2 **
  +-----------------+                            
  |                 |  +-- Hyriini **               
  |                 +--+                         
  |                    +-- Castaliini **            
  |                                              
  |              +-------- Westralunio **           
--+              |                               
  |  +-----------+     +-- Hyridella_drapeta **     
  |  |           |  +--+                         
  |  |           |  |  +-- Hyridella_depressa **    
  |  |           +--+                            
  |  |              |  +-- Diplodon_suavidicus__1 **
  +--+              +--+                         
     |                 +-- Diplodon_gratus **       
     |                                           
     |  +----------------- Echyridella **           
     |  |                                        
     +--+              +-- Cucumerunio **           
        |  +-----------+                         
        |  |           +-- Hyridella_australis **   
        +--+                                     
           |  +----------- Velesunio_ambiguus **    
           +--+                                  
              |  +-------- Alathyria **             
              +--+                               
                 |  +----- Velesunio_angasi **      
                 +--+                            
                    |  +-- Lortiella **             
                    +--+                         
                       +-- Microdontia **

Huang et al., 2019 [Publ 3843], fig. 2

tT	bi	riS	oT	roT	roS	iD
14	14	12	2	186	0	0	39

           +-- Westralunio **       
  +--------+                     
  |        +-- Hyriinae **          
  |                              
--+     +----- Alathyria_jacksoni **
  |  +--+                        
  |  |  |  +-- Alathyria_profuga ** 
  |  |  +--+                     
  +--+     +-- Velesunio **         
     |                           
     |  +----- Alathyria_pertexta **
     +--+                        
        |  +-- Lortiella **         
        +--+                     
           +-- Microdontia **

Huang et al., 2019 [Publ 3843], fig. S3

tT	bi	riS	oT	roT	roS	iD
14	14	12	2	184	0	0	41

           +-- Westralunio **       
  +--------+                     
  |        +-- Hyriinae **          
  |                              
--+     +----- Velesunio **         
  |  +--+                        
  |  |  |  +-- Alathyria_jacksoni **
  |  |  +--+                     
  +--+     +-- Alathyria_profuga ** 
     |                           
     |  +----- Alathyria_pertexta **
     +--+                        
        |  +-- Lortiella **         
        +--+                     
           +-- Microdontia **

Huang et al., 2019 [Publ 3843], fig. S4

tT	bi	riS	oT	roT	roS	iD
14	14	9	2	186	0	0	38

  +----------- Echyridella **       
  |                              
  +----------- Westralunio **       
  |                              
  +----------- Hyridellini **       
  |                              
  |     +----- Rhipidodontini **    
  +-----+                        
--+     |  +-- Castaliini **        
  |     +--+                     
  |        +-- Hyriini **           
  |                              
  |     +----- Velesunio **         
  |  +--+                        
  |  |  |  +-- Alathyria_jacksoni **
  |  |  +--+                     
  +--+     +-- Alathyria_profuga ** 
     |                           
     |  +----- Alathyria_pertexta **
     +--+                        
        |  +-- Lortiella **         
        +--+                     
           +-- Microdontia **

Graf et al., 2015 [Publ 3619], fig. 1

tT	bi	riS	oT	roT	roS	iD
14	14	12	2	6	0	0	37

  +-- Velesunioninae **
--+                 
  +-- Hyriinae **

Marshall et al., 2014 [Publ 3593], fig. 1

tT	bi	riS	oT	roT	roS	iD
10	10	8	2	4	0	0	31.5

  +-------- Echyridella **   
--+                       
  |  +----- Velesunioninae **
  +--+                    
     |  +-- Hyridellini **   
     +--+                 
        +-- Rhipidodontini **

Walker et al., 2006 [Publ 2666], fig. 6

tT	bi	riS	oT	roT	roS	iD
10	10	7	2	26	0	0	29

  +-------- Castaliini **         
  |                            
  +-------- Rhipidodontini **     
--+                            
  |  +----- Hyridella_depressa ** 
  |  |                         
  +--+----- Hyridella_australis **
     |                         
     |  +-- Echyridella **        
     +--+                      
        +-- Velesunioninae **

Hoeh et al., 2009 [Publ 2716], fig. 5

tT	bi	riS	oT	roT	roS	iD
9	9	7	2	37	0	0	32.5

  +-- Velesunioninae **
--+                 
  +-- Hyriinae **

Hoeh et al., 2009 [Publ 2716], fig. 3

tT	bi	riS	oT	roT	roS	iD
9	9	7	2	38	0	0	32.5

  +-- Velesunioninae **
--+                 
  +-- Hyriinae **

Hoeh et al., 2009 [Publ 2716], fig. 4

tT	bi	riS	oT	roT	roS	iD
9	9	7	2	37	0	0	32.5

  +-- Velesunioninae **
--+                 
  +-- Hyriinae **

Bolotov et al., 2016 [Publ 3743], fig. 4

tT	bi	riS	oT	roT	roS	iD
9	9	7	2	21	0	0	38

  +-- Hyriinae **      
--+                 
  +-- Velesunioninae **

Bolotov et al., 2017 [Publ 3761], fig. S2

tT	bi	riS	oT	roT	roS	iD
9	9	7	2	129	0	0	38

  +-- Hyriinae **      
--+                 
  +-- Velesunioninae **

Graf & Cummings, 2006 [Publ 1968], fig. 2

tT	bi	riS	oT	roT	roS	iD
9	9	7	2	38	0	0	32.5

  +-- Velesunioninae **
--+                 
  +-- Hyriinae **

Bolotov et al., 2017 [Publ 3761], fig. S1

tT	bi	riS	oT	roT	roS	iD
9	9	7	2	129	0	0	38

  +-- Hyriinae **      
--+                 
  +-- Velesunioninae **

Bolotov et al., 2016 [Publ 3743], fig. 2

tT	bi	riS	oT	roT	roS	iD
9	9	7	2	21	0	0	38

  +-- Hyriinae **      
--+                 
  +-- Velesunioninae **

Baker et al., 2004 [Publ 2685], fig. 2-3

tT	bi	riS	oT	roT	roS	iD
8	8	9	2	1	0	0	35

     +----- Hyridella_depressa ** 
  +--+                         
  |  |  +-- Hyridella_australis **
  |  +--+                      
--+     +-- Rhipidodontini **     
  |                            
  |  +----- Velesunioninae **     
  +--+                         
     |  +-- Echyridella **        
     +--+                      
        +-- Castaliini **

Baker et al., 2003 [Publ 2690], fig. 5

tT	bi	riS	oT	roT	roS	iD
7	7	5	2	1	0	0	37

           +-- Hyridella_australis **
  +--------+                      
  |        +-- Rhipidodontini **     
--+                               
  |  +-------- Hyridella_depressa ** 
  +--+                            
     |  +----- Velesunioninae **     
     +--+                         
        |  +-- Echyridella **        
        +--+                      
           +-- Castaliini **

Santos-Neto et al., 2016 [Publ 3739], fig. 3_5

tT	bi	riS	oT	roT	roS	iD
7	7	6	1	0	0	0	29

Graf & Ó Foighil, 2000 [Publ 2649], fig. 1_2

tT	bi	riS	oT	roT	roS	iD
6	6	5	2	5	0	0	36.5

  +-- Velesunioninae **
--+                 
  +-- Hyriinae **

Guerra et al., 2019 [Publ 4168], fig. 2

tT	bi	riS	oT	roT	roS	iD
5	5	3	2	36	0	0	33

  +-- Velesunioninae **
--+                 
  +-- Hyriinae **

Roe & Hoeh, 2003 [Publ 2655], fig. 4f2

tT	bi	riS	oT	roT	roS	iD
5	5	2	2	26	0	0	36

     +-- Castaliini **    
  +--+                 
  |  +-- Velesunio **     
--+                    
  |  +-- Echyridella **   
  |  |                 
  +--+-- Lortiella **     
     |                 
     +-- Rhipidodontini **

Bogan & Hoeh, 2000 [Publ 2650], fig. 1

tT	bi	riS	oT	roT	roS	iD
5	5	3	2	29	0	0	33

     +-- Castaliini **    
  +--+                 
  |  +-- Rhipidodontini **
--+                    
  |  +-- Echyridella **   
  +--+                 
     +-- Velesunioninae **

Campbell & Lydeard, 2026 [Publ 4810], fig. 3_5

tT	bi	riS	oT	roT	roS	iD
5	5	3	2	136	0	0	33

  +-- Velesunioninae **
--+                 
  +-- Hyriinae **

Hoeh et al., 2001 [Publ 2652], fig. 14f4

tT	bi	riS	oT	roT	roS	iD
5	5	3	2	26	0	0	33

     +-- Castaliini **    
  +--+                 
  |  +-- Rhipidodontini **
--+                    
  |  +-- Echyridella **   
  +--+                 
     +-- Velesunioninae **

Santos-Neto et al., 2018 [Publ 3888], fig. 5

tT	bi	riS	oT	roT	roS	iD
5	5	3	1	0	0	0	31

Roe & Hoeh, 2003 [Publ 2655], fig. 4f3

tT	bi	riS	oT	roT	roS	iD
5	5	3	2	26	0	0	33

     +-- Castaliini **    
  +--+                 
  |  +-- Rhipidodontini **
--+                    
  |  +-- Echyridella **   
  +--+                 
     +-- Velesunioninae **

Hoeh et al., 2001 [Publ 2652], fig. 14f2

tT	bi	riS	oT	roT	roS	iD
5	5	1	2	26	0	0	35

  +----- Rhipidodontini **
  |                    
  +----- Echyridella **   
--+                    
  +----- Lortiella **     
  |                    
  |  +-- Castaliini **    
  +--+                 
     +-- Velesunio **

Hoeh et al., 2002 [Publ 2653], fig. 2

tT	bi	riS	oT	roT	roS	iD
5	5	3	2	26	0	0	33.5

     +-- Castaliini **    
  +--+                 
  |  +-- Rhipidodontini **
--+                    
  |  +-- Echyridella **   
  +--+                 
     +-- Velesunioninae **

Guerra et al., 2019 [Publ 4168], fig. 1

tT	bi	riS	oT	roT	roS	iD
5	5	3	2	36	0	0	35

        +-- Echyridella **   
  +-----+                 
  |     +-- Velesunioninae **
--+                       
  |  +----- Rhipidodontini **
  +--+                    
     |  +-- Castaliini **    
     +--+                 
        +-- Hyriini **

Klunzinger et al., 2021 [Publ 4086], fig. 2

tT	bi	riS	oT	roT	roS	iD
4	4	2	2	2	0	0	32

  +----- Westralunio **
--+                 
  |  +-- Velesunio **  
  +--+              
     +-- Hyriinae **

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

tT	bi	riS	oT	roT	roS	iD
4	4	2	2	36	0	0	34

  +-- Velesunioninae **
--+                 
  +-- Hyriinae **

Whelan et al., 2011 [Publ 3380], fig. 1-ML

tT	bi	riS	oT	roT	roS	iD
4	4	2	2	32	0	0	34

  +-- Velesunioninae **
--+                 
  +-- Hyriinae **

Graf, 2002 [Publ 2651], fig. 2

tT	bi	riS	oT	roT	roS	iD
4	4	2	2	14	0	0	31.5

  +-- Velesunioninae **
--+                 
  +-- Hyriinae **

Klunzinger et al., 2022 [Publ 4259], fig. 1

tT	bi	riS	oT	roT	roS	iD
4	4	2	2	0	0	0	34

  +-- Hyriinae **      
--+                 
  +-- Velesunioninae **

Whelan et al., 2011 [Publ 3380], fig. 1-MP

tT	bi	riS	oT	roT	roS	iD
4	4	2	2	32	0	0	34

  +-- Velesunioninae **
--+                 
  +-- Hyriinae **

Graf, 2000 [Publ 2647], fig. 1

tT	bi	riS	oT	roT	roS	iD
4	4	2	2	13	0	0	34.5

  +-------- Castaliini **    
--+                       
  |  +----- Rhipidodontini **
  +--+                    
     |  +-- Hyridellini **   
     +--+                 
        +-- Velesunioninae **

Sonowal et al., 2021 [Publ 4221], fig. 4

tT	bi	riS	oT	roT	roS	iD
3	3	1	2	13	0	0	33

  +----- Hyriini **       
--+                    
  |  +-- Velesunioninae **
  +--+                 
     +-- Hyridellini **

Sonowal et al., 2021 [Publ 4221], fig. 3a

tT	bi	riS	oT	roT	roS	iD
3	3	0	2	12	0	0	34

  +-- Hyriini **       
  |                 
--+-- Hyridellini **   
  |                 
  +-- Velesunioninae **

Sonowal et al., 2021 [Publ 4221], fig. 3b

tT	bi	riS	oT	roT	roS	iD
3	3	0	2	12	0	0	34

  +-- Hyriini **       
  |                 
--+-- Velesunioninae **
  |                 
  +-- Hyridellini **

Konopleva et al., 2017 [Publ 3807], fig. 6

tT	bi	riS	oT	roT	roS	iD
3	3	1	1	28	0	0	33

Sano et al., 2020 [Publ 4138], fig. 4

tT	bi	riS	oT	roT	roS	iD
3	3	1	2	65	0	0	33

  +----- Hyriini **       
--+                    
  |  +-- Hyridellini **   
  +--+                 
     +-- Velesunioninae **

Olivera-Hyde et al., 2020 [Publ 4249], fig. 10

tT	bi	riS	oT	roT	roS	iD
3	3	1	1	0	0	0	33

Lopes-Lima et al., 2018 [Publ 3829], fig. 1

tT	bi	riS	oT	roT	roS	iD
3	3	1	2	25	0	0	33

  +----- Hyriini **       
--+                    
  |  +-- Velesunioninae **
  +--+                 
     +-- Hyridellini **

Combosch et al., 2017 [Publ 3806], fig. 1

tT	bi	riS	oT	roT	roS	iD
3	3	1	2	12	0	0	33

  +----- Hyriini **       
--+                    
  |  +-- Velesunioninae **
  +--+                 
     +-- Hyridellini **

Lopes-Lima et al., 2018 [Publ 3829], fig. 3

tT	bi	riS	oT	roT	roS	iD
3	3	1	2	25	0	0	33

  +----- Hyriini **       
--+                    
  |  +-- Velesunioninae **
  +--+                 
     +-- Hyridellini **

Huang et al., 2018 [Publ 3828], fig. 3_5

tT	bi	riS	oT	roT	roS	iD
3	3	1	2	26	0	0	33

  +----- Hyriini **       
--+                    
  |  +-- Velesunioninae **
  +--+                 
     +-- Hyridellini **

Combosch et al., 2017 [Publ 3806], fig. 2

tT	bi	riS	oT	roT	roS	iD
3	3	1	2	12	0	0	35

  +-- Velesunioninae **
--+                 
  +-- Hyriinae **

Wu et al., 2020 [Publ 4121], fig. S1

tT	bi	riS	oT	roT	roS	iD
3	3	1	2	31	0	0	33

  +----- Hyriini **       
--+                    
  |  +-- Hyridellini **   
  +--+                 
     +-- Velesunioninae **

Wu et al., 2021 [Publ 4366], fig. 2_I

tT	bi	riS	oT	roT	roS	iD
3	3	1	2	88	0	0	35

  +-- Velesunioninae **
--+                 
  +-- Hyriinae **

Wu et al., 2020 [Publ 4121], fig. 2

tT	bi	riS	oT	roT	roS	iD
3	3	1	2	31	0	0	33

  +----- Hyriini **       
--+                    
  |  +-- Hyridellini **   
  +--+                 
     +-- Velesunioninae **

Sano et al., 2020 [Publ 4138], fig. 2

tT	bi	riS	oT	roT	roS	iD
3	3	1	2	65	0	0	33

  +----- Hyriini **       
--+                    
  |  +-- Hyridellini **   
  +--+                 
     +-- Velesunioninae **

Wu et al., 2021 [Publ 4366], fig. 6

tT	bi	riS	oT	roT	roS	iD
3	3	1	2	88	0	0	35

  +-- Velesunioninae **
--+                 
  +-- Hyriinae **

Sano et al., 2020 [Publ 4138], fig. 3

tT	bi	riS	oT	roT	roS	iD
3	3	1	2	65	0	0	33

  +----- Hyriini **       
--+                    
  |  +-- Hyridellini **   
  +--+                 
     +-- Velesunioninae **

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	78	0	0	--

  +-- Hyriinae **      
--+                 
  +-- Velesunioninae **

Guerra et al., 2019 [Publ 4168], fig. 2-M

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	20	0	0	--

  +-- Hyriinae **      
--+                 
  +-- Velesunioninae **

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	120	0	0	--

  +-- Hyriinae **      
--+                 
  +-- Velesunioninae **

Guerra et al., 2019 [Publ 4168], fig. 1-M

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	20	0	0	--

  +-- Hyriinae **      
--+                 
  +-- Velesunioninae **

Chen et al., 2025 [Publ 4455], fig. 1

tT	bi	riS	oT	roT	roS	iD
2	2	0	1	24	0	0	--

Campbell & Lydeard, 2026 [Publ 4810], fig. 4

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	45	0	0	--

  +-- Hyriinae **      
--+                 
  +-- Velesunioninae **

Wu et al., 2019 [Publ 3864], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	45	0	0	--

-- Echyridella_menziesii **

Guerra et al., 2017 [Publ 3772], fig. 2a-M

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	7	0	0	--

-- Echyridella_menziesii **

Huang et al., 2018 [Publ 3828], fig. 6

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	41	0	0	--

-- Echyridella_menziesii **

Wu et al., 2022 [Publ 4238], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	73	0	0	--

-- Echyridella_menziesii **

González & Giribet, 2015 [Publ 3632], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	5	0	0	--

-- Velesunio_ambiguus **

González & Giribet, 2015 [Publ 3632], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	5	0	0	--

-- Velesunio_ambiguus **

González & Giribet, 2015 [Publ 3632], fig. 4

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	5	0	0	--

-- Velesunio_ambiguus **

Bieler et al., 2014 [Publ 3622], fig. 26_27_29_30

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	4	0	0	--

-- Velesunio_ambiguus **

Huang et al., 2019 [Publ 3843], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	55	0	0	--

-- Echyridella_menziesii **

Pfeiffer et al., 2019 [Publ 3937], fig. 6

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	56	0	0	--

-- Hyridella_australis **

Chase et al., 2018 [Publ 3861], fig. 1-M

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	10	0	0	--

-- Echyridella_menziesii **

Wu et al., 2021 [Publ 4366], fig. 4

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	68	0	0	--

-- Echyridella_menziesii **

Wu et al., 2024 [Publ 4350], fig. 8b_S2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	91	0	0	--

-- Echyridella_menziesii **

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	30	0	0	--

-- Velesunio_ambiguus **

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	95	0	0	--

-- Echyridella_menziesii **

Elderkin et al., 2016 [Publ 3693], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	2	0	0	--

-- Velesunio_ambiguus **

Walker et al., 2006 [Publ 2666], fig. 6-M

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	13	0	0	--

-- Echyridella_menziesii **

Plazzi et al., 2011 [Publ 3676], fig. 5

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	17	0	0	--

-- Alathyria_jacksoni **

Wu et al., 2021 [Publ 4366], fig. 3-BI

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	73	0	0	--

-- Echyridella_menziesii **

Chase et al., 2018 [Publ 3861], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	10	0	0	--

-- Echyridella_menziesii **

Walker et al., 2006 [Publ 2666], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	14	0	0	--

-- Hyridella **

Wu et al., 2024 [Publ 4350], fig. 8a_S1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	91	0	0	--

-- Echyridella_menziesii **

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	29	0	0	--

-- Hyridella_depressa **

Dai et al., 2025 [Publ 4440], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	87	0	0	--

-- Echyridella_menziesii **

Walker et al., 2006 [Publ 2666], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	14	0	0	--

-- Hyridella **

Hoeh et al., 2002 [Publ 2673], fig. 1-F

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	8	0	0	--

-- Echyridella_menziesii **

Guerra et al., 2017 [Publ 3772], fig. 2b-M

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	7	0	0	--

-- Echyridella_menziesii **

Breton et al., 2011 [Publ 3376], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	37	0	0	--

-- Echyridella_menziesii **

Chen et al., 2025 [Publ 4448], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	85	0	0	--

-- Echyridella_menziesii **

Hoeh et al., 2002 [Publ 2673], fig. 1-M

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	8	0	0	--

-- Echyridella_menziesii **

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	29	0	0	--

-- Hyridella_depressa **

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	95	0	0	--

-- Echyridella_menziesii **

González & Giribet, 2015 [Publ 3632], fig. 6

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	5	0	0	--

-- Velesunio_ambiguus **

Chase et al., 2018 [Publ 3847], fig. 5

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	9	0	0	--

-- Echyridella_menziesii **

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	56	0	0	--

-- Hyridella_australis **

Guerra et al., 2017 [Publ 3772], fig. 2a

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	24	0	0	--

-- Echyridella_menziesii **

Guerra et al., 2017 [Publ 3772], fig. 2b

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	24	0	0	--

-- Echyridella_menziesii **

Wu et al., 2020 [Publ 4121], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	48	0	0	--

-- Echyridella_menziesii **

Walker et al., 2006 [Publ 2666], fig. 4

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	14	0	0	--

-- Hyridella **

Wu et al., 2021 [Publ 4366], fig. 3-ML

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	73	0	0	--

-- Echyridella_menziesii **

Huang et al., 2018 [Publ 3828], fig. 6-M

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	16	0	0	--

-- Echyridella_menziesii **

Huang et al., 2019 [Publ 3843], fig. 3-M

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	22	0	0	--

-- Echyridella_menziesii **

Chase et al., 2018 [Publ 3847], fig. 4

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	38	0	0	--

-- Echyridella_menziesii **

Bieler et al., 2014 [Publ 3622], fig. 28

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	4	0	0	--

-- Velesunio_ambiguus **

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