MUSSELpdb | subfamily Hyriinae Swainson, 1840 cladomics

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Page last updated
Thu 3 September 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

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Bivalvia | Unionoida | Hyrioidea | Hyriidae

subfamily HYRIINAE Swainson, 1840: cladomics
4 tribes, 1 genus

Cladomics Summary

number of species = 76 (86 total)
number of subtaxa (rank: tribe) = 5
analyzed in 93 cladograms in 49 publications
maximum number of terminals in an analysis = 13 (mean = 2.68, median = 2)
maximum number of subtaxa (tribe) = 5 (mean = 2.14, median = 2)

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 52
maximum number of included terminals = 13, (mean = 4 median = 3)
maximum number of subtaxa (tribe) = 5 (mean = 3.04, median = 3)
proportion of cladograms that support basic monophyly = 0.48

Rigorous Monophyly

number of cladograms that test rigorous monophyly = 47
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (tribe) taxon and at least one rigorous outgroup (family).
maximum number of included terminals = 13 (mean = 4, median = 3)
maximum number of subtaxa (tribe) = 5 (mean = 3.13, median = 3)
proportion of cladograms that support rigorous monophyly = 0.43

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination

NJ Tree

Santos-Neto et al., 2016 [Publ 3739], fig. 4

** Top Monophyly Cladogram **

tT	bi	riS	oT	roT	roS	iD
13	19	21	5	7	6	1	0

                    +----- Diplodon_suavidicus__2 **
  +-----------------+                            
  |                 |  +-- Hyriini **               
  |                 +--+                         
  |                    +-- Castaliini **            
  |                                              
  |              +-------- Westralunio           
--+              |                               
  |  +-----------+     +-- Hyridella_drapeta **     
  |  |           |  +--+                         
  |  |           |  |  +-- Hyridella_depressa **    
  |  |           +--+                            
  |  |              |  +-- Diplodon_suavidicus__1 **
  +--+              +--+                         
     |                 +-- Diplodon_gratus **       
     |                                           
     |  +----------------- Echyridella **           
     |  |                                        
     +--+              +-- Cucumerunio **           
        |  +-----------+                         
        |  |           +-- Hyridella_australis **   
        +--+                                     
           |  +----------- Velesunio_ambiguus    
           +--+                                  
              |  +-------- Alathyria             
              +--+                               
                 |  +----- Velesunio_angasi      
                 +--+                            
                    |  +-- Lortiella             
                    +--+                         
                       +-- Microdontia

Huang et al., 2019 [Publ 3843], fig. S4

** Top Sister-Group Cladogram **

tT	bi	riS	oT	roT	roS	iD
7	14	9	5	194	7	1	22.5

  +----------- Echyridella **       
  |                              
  +----------- Westralunio       
  |                              
  +----------- Hyridellini **       
  |                              
  |     +----- Rhipidodontini **    
  +-----+                        
--+     |  +-- Castaliini **        
  |     +--+                     
  |        +-- Hyriini **           
  |                              
  |     +----- Velesunio         
  |  +--+                        
  |  |  |  +-- Alathyria_jacksoni
  |  |  +--+                     
  +--+     +-- Alathyria_profuga 
     |                           
     |  +----- Alathyria_pertexta
     +--+                        
        |  +-- Lortiella         
        +--+                     
           +-- Microdontia

Santos-Neto et al., 2016 [Publ 3739], fig. 2a_b

tT	bi	riS	oT	roT	roS	iD
13	19	20	5	7	6	1	11

                 +----- Diplodon_suavidicus__1 **
  +--------------+                            
  |              |  +-- Hyriini **               
  |              +--+                         
  |                 +-- Castaliini **            
  |                                           
  |           +-------- Westralunio           
  |           |                               
  +-----------+     +-- Hyridella_drapeta **     
  |           |  +--+                         
--+           |  |  +-- Hyridella_depressa **    
  |           +--+                            
  |              |  +-- Diplodon_gratus **       
  |              +--+                         
  |                 +-- Diplodon_suavidicus__2 **
  |                                           
  |                 +-- Hyridella_australis **   
  |  +--------------+                         
  |  |              +-- Cucumerunio **           
  +--+                                        
     |  +-------------- Echyridella **           
     +--+                                     
        |  +----------- Velesunio_ambiguus    
        +--+                                  
           |  +-------- Microdontia           
           +--+                               
              |  +----- Lortiella             
              +--+                            
                 |  +-- Alathyria             
                 +--+                         
                    +-- Velesunio_angasi

Huang et al., 2019 [Publ 3843], fig. 2

tT	bi	riS	oT	roT	roS	iD
7	7	5	5	194	7	1	22

     +-------------- Westralunio       
  +--+                                 
  |  |  +----------- Echyridella **       
  |  +--+                              
  |     |  +-------- Hyridellini **       
  |     +--+                           
  |        |  +----- Rhipidodontini **    
  |        +--+                        
--+           |  +-- Castaliini **        
  |           +--+                     
  |              +-- Hyriini **           
  |                                    
  |           +----- Alathyria_jacksoni
  |        +--+                        
  |        |  |  +-- Alathyria_profuga 
  |        |  +--+                     
  +--------+     +-- Velesunio         
           |                           
           |  +----- Alathyria_pertexta
           +--+                        
              |  +-- Lortiella         
              +--+                     
                 +-- Microdontia

Marshall et al., 2014 [Publ 3593], fig. 1

tT	bi	riS	oT	roT	roS	iD
7	10	8	3	7	3	1	25

  +-------- Echyridella **   
--+                       
  |  +----- Velesunioninae
  +--+                    
     |  +-- Hyridellini **   
     +--+                 
        +-- Rhipidodontini **

Santos-Neto et al., 2016 [Publ 3739], fig. 3_5

tT	bi	riS	oT	roT	roS	iD
7	7	6	3	0	0	0	18

  +----- Rhipidodontini **
--+                    
  |  +-- Hyriini **       
  +--+                 
     +-- Castaliini **

Graf et al., 2015 [Publ 3619], fig. 1

tT	bi	riS	oT	roT	roS	iD
7	7	5	4	13	7	1	20

  +----------- Velesunioninae
--+                          
  |  +-------- Hyridellini **   
  +--+                       
     |  +----- Rhipidodontini **
     +--+                    
        |  +-- Hyriini **       
        +--+                 
           +-- Castaliini **

Huang et al., 2019 [Publ 3843], fig. S3

tT	bi	riS	oT	roT	roS	iD
7	7	5	5	192	7	1	22

     +-------------- Westralunio       
  +--+                                 
  |  |  +----------- Echyridella **       
  |  +--+                              
  |     |  +-------- Hyridellini **       
  |     +--+                           
  |        |  +----- Rhipidodontini **    
  |        +--+                        
--+           |  +-- Castaliini **        
  |           +--+                     
  |              +-- Hyriini **           
  |                                    
  |           +----- Velesunio         
  |        +--+                        
  |        |  |  +-- Alathyria_jacksoni
  |        |  +--+                     
  +--------+     +-- Alathyria_profuga 
           |                           
           |  +----- Alathyria_pertexta
           +--+                        
              |  +-- Lortiella         
              +--+                     
                 +-- Microdontia

Hoeh et al., 2009 [Publ 2716], fig. 4

tT	bi	riS	oT	roT	roS	iD
6	6	4	4	40	3	1	25.5

  +----------- Velesunioninae
--+                          
  |  +-------- Echyridella **   
  +--+                       
     |  +----- Hyridellini **   
     +--+                    
        |  +-- Castaliini **    
        +--+                 
           +-- Rhipidodontini **

Hoeh et al., 2009 [Publ 2716], fig. 5

tT	bi	riS	oT	roT	roS	iD
6	6	4	4	40	3	1	25.5

  +----------- Velesunioninae
--+                          
  |  +-------- Echyridella **   
  +--+                       
     |  +----- Hyridellini **   
     +--+                    
        |  +-- Castaliini **    
        +--+                 
           +-- Rhipidodontini **

Hoeh et al., 2009 [Publ 2716], fig. 3

tT	bi	riS	oT	roT	roS	iD
6	6	4	4	41	3	1	25.5

  +----------- Velesunioninae
--+                          
  |  +-------- Echyridella **   
  +--+                       
     |  +----- Hyridellini **   
     +--+                    
        |  +-- Castaliini **    
        +--+                 
           +-- Rhipidodontini **

Graf & Cummings, 2006 [Publ 1968], fig. 2

tT	bi	riS	oT	roT	roS	iD
6	6	4	4	41	3	1	25.5

  +----------- Velesunioninae
--+                          
  |  +-------- Echyridella **   
  +--+                       
     |  +----- Hyridellini **   
     +--+                    
        |  +-- Castaliini **    
        +--+                 
           +-- Rhipidodontini **

Baker et al., 2003 [Publ 2690], fig. 5

tT	bi	riS	oT	roT	roS	iD
5	7	5	4	3	2	1	24

           +-- Hyridella_australis **
  +--------+                      
  |        +-- Rhipidodontini **     
--+                               
  |  +-------- Hyridella_depressa ** 
  +--+                            
     |  +----- Velesunioninae     
     +--+                         
        |  +-- Echyridella **        
        +--+                      
           +-- Castaliini **

Santos-Neto et al., 2018 [Publ 3888], fig. 5

tT	bi	riS	oT	roT	roS	iD
5	5	3	2	0	0	0	18

  +-- Hyriini **   
--+             
  +-- Castaliini **

Walker et al., 2006 [Publ 2666], fig. 6

tT	bi	riS	oT	roT	roS	iD
5	10	7	4	31	5	1	21.5

  +-------- Castaliini **         
  |                            
  +-------- Rhipidodontini **     
--+                            
  |  +----- Hyridella_depressa ** 
  |  |                         
  +--+----- Hyridella_australis **
     |                         
     |  +-- Echyridella **        
     +--+                      
        +-- Velesunioninae

Graf & Ó Foighil, 2000 [Publ 2649], fig. 1_2

tT	bi	riS	oT	roT	roS	iD
5	5	4	4	6	1	1	26.5

  +-------------- Velesunioninae        
  |                                     
--+           +-- Rhipidodontini **        
  |  +--------+                         
  |  |        +-- Castaliini **            
  +--+                                  
     |  +-------- Hyridella_australis__2 **
     +--+                               
        |  +----- Hyridella_australis__1 **
        +--+                            
           |  +-- Echyridella **           
           +--+                         
              +-- Hyridella_depressa **

Baker et al., 2004 [Publ 2685], fig. 2-3

tT	bi	riS	oT	roT	roS	iD
5	8	9	4	4	3	1	25

     +----- Hyridella_depressa ** 
  +--+                         
  |  |  +-- Hyridella_australis **
  |  +--+                      
--+     +-- Rhipidodontini **     
  |                            
  |  +----- Velesunioninae     
  +--+                         
     |  +-- Echyridella **        
     +--+                      
        +-- Castaliini **

Guerra et al., 2019 [Publ 4168], fig. 2

tT	bi	riS	oT	roT	roS	iD
4	4	2	4	37	1	1	22

  +----------- Velesunioninae
--+                          
  |  +-------- Echyridella **   
  +--+                       
     |  +----- Rhipidodontini **
     +--+                    
        |  +-- Castaliini **    
        +--+                 
           +-- Hyriini **

Guerra et al., 2019 [Publ 4168], fig. 1

tT	bi	riS	oT	roT	roS	iD
4	5	3	4	37	1	1	23

        +-- Echyridella **   
  +-----+                 
  |     +-- Velesunioninae
--+                       
  |  +----- Rhipidodontini **
  +--+                    
     |  +-- Castaliini **    
     +--+                 
        +-- Hyriini **

Konopleva et al., 2017 [Publ 3807], fig. 6

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	28	0	0	20

  +----- Rhipidodontini **
--+                    
  |  +-- Hyriini **       
  +--+                 
     +-- Castaliini **

Bogan & Hoeh, 2000 [Publ 2650], fig. 1

tT	bi	riS	oT	roT	roS	iD
3	5	3	3	31	2	1	20.5

     +-- Castaliini **    
  +--+                 
  |  +-- Rhipidodontini **
--+                    
  |  +-- Echyridella **   
  +--+                 
     +-- Velesunioninae

Hoeh et al., 2002 [Publ 2653], fig. 2

tT	bi	riS	oT	roT	roS	iD
3	5	3	3	28	2	1	20

     +-- Castaliini **    
  +--+                 
  |  +-- Rhipidodontini **
--+                    
  |  +-- Echyridella **   
  +--+                 
     +-- Velesunioninae

Roe & Hoeh, 2003 [Publ 2655], fig. 4f3

tT	bi	riS	oT	roT	roS	iD
3	5	3	3	28	2	1	20.5

     +-- Castaliini **    
  +--+                 
  |  +-- Rhipidodontini **
--+                    
  |  +-- Echyridella **   
  +--+                 
     +-- Velesunioninae

Bolotov et al., 2017 [Publ 3761], fig. S1

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	136	6	1	20

  +-------- Velesunioninae
--+                       
  |  +----- Rhipidodontini **
  +--+                    
     |  +-- Castaliini **    
     +--+                 
        +-- Hyriini **

Graf, 2002 [Publ 2651], fig. 2

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	15	1	1	23.5

  +-------- Velesunioninae
--+                       
  |  +----- Hyridellini **   
  +--+                    
     |  +-- Rhipidodontini **
     +--+                 
        +-- Castaliini **

Olivera-Hyde et al., 2020 [Publ 4249], fig. 10

tT	bi	riS	oT	roT	roS	iD
3	3	1	1	0	0	0	20

Bolotov et al., 2016 [Publ 3743], fig. 2

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	27	6	1	20

  +-------- Velesunioninae
--+                       
  |  +----- Rhipidodontini **
  +--+                    
     |  +-- Hyriini **       
     +--+                 
        +-- Castaliini **

Bolotov et al., 2017 [Publ 3761], fig. S2

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	136	6	1	20

  +-------- Velesunioninae
--+                       
  |  +----- Rhipidodontini **
  +--+                    
     |  +-- Castaliini **    
     +--+                 
        +-- Hyriini **

Roe & Hoeh, 2003 [Publ 2655], fig. 4f2

tT	bi	riS	oT	roT	roS	iD
3	5	2	3	28	2	1	23.5

     +-- Castaliini **    
  +--+                 
  |  +-- Velesunio     
--+                    
  |  +-- Echyridella **   
  |  |                 
  +--+-- Lortiella     
     |                 
     +-- Rhipidodontini **

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	37	1	1	20

  +-------- Velesunioninae
--+                       
  |  +----- Hyridellini **   
  +--+                    
     |  +-- Hyriini **       
     +--+                 
        +-- Castaliini **

Hoeh et al., 2001 [Publ 2652], fig. 14f2

tT	bi	riS	oT	roT	roS	iD
3	5	1	3	28	2	1	22.5

  +----- Rhipidodontini **
  |                    
  +----- Echyridella **   
--+                    
  +----- Lortiella     
  |                    
  |  +-- Castaliini **    
  +--+                 
     +-- Velesunio

Bolotov et al., 2016 [Publ 3743], fig. 4

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	27	6	1	20

  +-------- Velesunioninae
--+                       
  |  +----- Rhipidodontini **
  +--+                    
     |  +-- Hyriini **       
     +--+                 
        +-- Castaliini **

Graf, 2000 [Publ 2647], fig. 1

tT	bi	riS	oT	roT	roS	iD
3	4	2	3	14	1	1	23

  +-------- Castaliini **    
--+                       
  |  +----- Rhipidodontini **
  +--+                    
     |  +-- Hyridellini **   
     +--+                 
        +-- Velesunioninae

Whelan et al., 2011 [Publ 3380], fig. 1-ML

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	33	1	1	20

  +-------- Velesunioninae
--+                       
  |  +----- Hyridellini **   
  +--+                    
     |  +-- Castaliini **    
     +--+                 
        +-- Hyriini **

Whelan et al., 2011 [Publ 3380], fig. 1-MP

tT	bi	riS	oT	roT	roS	iD
3	3	1	3	33	1	1	20

  +-------- Velesunioninae
--+                       
  |  +----- Hyridellini **   
  +--+                    
     |  +-- Castaliini **    
     +--+                 
        +-- Hyriini **

Hoeh et al., 2001 [Publ 2652], fig. 14f4

tT	bi	riS	oT	roT	roS	iD
3	5	3	3	28	2	1	20.5

     +-- Castaliini **    
  +--+                 
  |  +-- Rhipidodontini **
--+                    
  |  +-- Echyridella **   
  +--+                 
     +-- Velesunioninae

Combosch et al., 2017 [Publ 3806], fig. 2

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	13	1	1	--

  +----- Velesunioninae
--+                    
  |  +-- Hyriini **       
  +--+                 
     +-- Hyridellini **

Wu et al., 2021 [Publ 4366], fig. 2_I

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	89	1	1	--

  +----- Velesunioninae
--+                    
  |  +-- Castaliini **    
  +--+                 
     +-- Rhipidodontini **

Wu et al., 2020 [Publ 4121], fig. 2

tT	bi	riS	oT	roT	roS	iD
2	3	0	2	32	1	1	20

  +----- Hyriini **       
--+                    
  |  +-- Hyridellini **   
  +--+                 
     +-- Velesunioninae

Lopes-Lima et al., 2018 [Publ 3829], fig. 1

tT	bi	riS	oT	roT	roS	iD
2	3	0	2	26	1	1	20

  +----- Hyriini **       
--+                    
  |  +-- Velesunioninae
  +--+                 
     +-- Hyridellini **

Sano et al., 2020 [Publ 4138], fig. 2

tT	bi	riS	oT	roT	roS	iD
2	3	0	2	66	1	1	20

  +----- Hyriini **       
--+                    
  |  +-- Hyridellini **   
  +--+                 
     +-- Velesunioninae

Combosch et al., 2017 [Publ 3806], fig. 1

tT	bi	riS	oT	roT	roS	iD
2	3	0	2	13	1	1	20

  +----- Hyriini **       
--+                    
  |  +-- Velesunioninae
  +--+                 
     +-- Hyridellini **

Lopes-Lima et al., 2018 [Publ 3829], fig. 3

tT	bi	riS	oT	roT	roS	iD
2	3	0	2	26	1	1	20

  +----- Hyriini **       
--+                    
  |  +-- Velesunioninae
  +--+                 
     +-- Hyridellini **

Sonowal et al., 2021 [Publ 4221], fig. 4

tT	bi	riS	oT	roT	roS	iD
2	3	0	2	14	1	1	20

  +----- Hyriini **       
--+                    
  |  +-- Velesunioninae
  +--+                 
     +-- Hyridellini **

Huang et al., 2018 [Publ 3828], fig. 3_5

tT	bi	riS	oT	roT	roS	iD
2	3	0	2	27	1	1	20

  +----- Hyriini **       
--+                    
  |  +-- Velesunioninae
  +--+                 
     +-- Hyridellini **

Wu et al., 2020 [Publ 4121], fig. S1

tT	bi	riS	oT	roT	roS	iD
2	3	0	2	32	1	1	20

  +----- Hyriini **       
--+                    
  |  +-- Hyridellini **   
  +--+                 
     +-- Velesunioninae

Wu et al., 2021 [Publ 4366], fig. 6

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	89	1	1	--

  +----- Velesunioninae
--+                    
  |  +-- Rhipidodontini **
  +--+                 
     +-- Castaliini **

Sonowal et al., 2021 [Publ 4221], fig. 3b

tT	bi	riS	oT	roT	roS	iD
2	3	0	2	13	1	1	21

  +-- Hyriini **       
  |                 
--+-- Velesunioninae
  |                 
  +-- Hyridellini **

Chen et al., 2025 [Publ 4455], fig. 1

tT	bi	riS	oT	roT	roS	iD
2	2	0	2	24	0	0	--

  +-- Hyridellini **
--+              
  +-- Hyriini **

Sano et al., 2020 [Publ 4138], fig. 4

tT	bi	riS	oT	roT	roS	iD
2	3	0	2	66	1	1	20

  +----- Hyriini **       
--+                    
  |  +-- Hyridellini **   
  +--+                 
     +-- Velesunioninae

Sonowal et al., 2021 [Publ 4221], fig. 3a

tT	bi	riS	oT	roT	roS	iD
2	3	0	2	13	1	1	21

  +-- Hyriini **       
  |                 
--+-- Hyridellini **   
  |                 
  +-- Velesunioninae

Sano et al., 2020 [Publ 4138], fig. 3

tT	bi	riS	oT	roT	roS	iD
2	3	0	2	66	1	1	20

  +----- Hyriini **       
--+                    
  |  +-- Hyridellini **   
  +--+                 
     +-- Velesunioninae

Wu et al., 2024 [Publ 4350], fig. 8b_S2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	91	0	0	--

-- Echyridella_menziesii **

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	29	0	0	--

-- Hyridella_depressa **

Wu et al., 2019 [Publ 3864], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	45	0	0	--

-- Echyridella_menziesii **

Wu et al., 2022 [Publ 4238], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	73	0	0	--

-- Echyridella_menziesii **

Guerra et al., 2017 [Publ 3772], fig. 2a

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	24	0	0	--

-- Echyridella_menziesii **

Walker et al., 2006 [Publ 2666], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	14	0	0	--

-- Hyridella **

Huang et al., 2019 [Publ 3843], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	55	0	0	--

-- Echyridella_menziesii **

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	95	0	0	--

-- Echyridella_menziesii **

Walker et al., 2006 [Publ 2666], fig. 6-M

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	13	0	0	--

-- Echyridella_menziesii **

Pfeiffer et al., 2019 [Publ 3937], fig. 6

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	56	0	0	--

-- Hyridella_australis **

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	95	0	0	--

-- Echyridella_menziesii **

Guerra et al., 2017 [Publ 3772], fig. 2b-M

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	7	0	0	--

-- Echyridella_menziesii **

Guerra et al., 2017 [Publ 3772], fig. 2a-M

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	7	0	0	--

-- Echyridella_menziesii **

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	56	0	0	--

-- Hyridella_australis **

Chase et al., 2018 [Publ 3861], fig. 1-M

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	10	0	0	--

-- Echyridella_menziesii **

Wu et al., 2021 [Publ 4366], fig. 3-BI

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	73	0	0	--

-- Echyridella_menziesii **

Huang et al., 2019 [Publ 3843], fig. 3-M

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	22	0	0	--

-- Echyridella_menziesii **

Huang et al., 2018 [Publ 3828], fig. 6-M

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	16	0	0	--

-- Echyridella_menziesii **

Guerra et al., 2017 [Publ 3772], fig. 2b

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	24	0	0	--

-- Echyridella_menziesii **

Chase et al., 2018 [Publ 3861], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	10	0	0	--

-- Echyridella_menziesii **

Guerra et al., 2019 [Publ 4168], fig. 2-M

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	21	1	1	--

  +-- Echyridella **   
--+                 
  +-- Velesunioninae

Walker et al., 2006 [Publ 2666], fig. 4

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	14	0	0	--

-- Hyridella **

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	29	0	0	--

-- Hyridella_depressa **

Wu et al., 2021 [Publ 4366], fig. 3-ML

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	73	0	0	--

-- Echyridella_menziesii **

Chase et al., 2018 [Publ 3847], fig. 5

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	9	0	0	--

-- Echyridella_menziesii **

Dai et al., 2025 [Publ 4440], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	87	0	0	--

-- Echyridella_menziesii **

Walker et al., 2006 [Publ 2666], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	14	0	0	--

-- Hyridella **

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	79	1	1	--

  +-- Hyridellini **   
--+                 
  +-- Velesunioninae

Breton et al., 2011 [Publ 3376], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	36	0	0	--

-- Echyridella_menziesii **

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	121	1	1	--

  +-- Hyridellini **   
--+                 
  +-- Velesunioninae

Chen et al., 2025 [Publ 4448], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	85	0	0	--

-- Echyridella_menziesii **

Klunzinger et al., 2022 [Publ 4259], fig. 1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	3	3	1	--

  +-- Hyridellini **   
--+                 
  +-- Velesunioninae

Guerra et al., 2019 [Publ 4168], fig. 1-M

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	21	1	1	--

  +-- Echyridella **   
--+                 
  +-- Velesunioninae

Klunzinger et al., 2021 [Publ 4086], fig. 2

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	5	3	1	--

  +----- Westralunio
--+                 
  |  +-- Velesunio  
  +--+              
     +-- Hyridellini **

Huang et al., 2018 [Publ 3828], fig. 6

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	41	0	0	--

-- Echyridella_menziesii **

Chase et al., 2018 [Publ 3847], fig. 4

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	37	0	0	--

-- Echyridella_menziesii **

Hoeh et al., 2002 [Publ 2673], fig. 1-M

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	8	0	0	--

-- Echyridella_menziesii **

Wu et al., 2020 [Publ 4121], fig. 3

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	48	0	0	--

-- Echyridella_menziesii **

Wu et al., 2024 [Publ 4350], fig. 8a_S1

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	91	0	0	--

-- Echyridella_menziesii **

Wu et al., 2021 [Publ 4366], fig. 4

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	68	0	0	--

-- Echyridella_menziesii **

Hoeh et al., 2002 [Publ 2673], fig. 1-F

tT	bi	riS	oT	roT	roS	iD
1	1	0	1	8	0	0	--

-- Echyridella_menziesii **

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